The human ear is a complex and wonderful instrument to say though that all it does is to enable us to hear is to really understate the things that is capable of. It is first of all, a collector of sound waves, it catches the sound waves and directs them down into the external canal so that they can find their way on further into the inner ear where they can be received and interpreted. It is also an energy transducer that is, it changes the acoustic energy of the airborne signal to mechanical energy to hydraulic energy and finally to electrical energy. It is also a sound amplifier that is, it actually increases the loudness of the sound after it reaches the ear. It's also a sound analyzer that is, it allows us to be able to interpret the complex noise that we call speech. And finally, it contains the mechanism that enables us to be able to maintain our balance and to stay in an upright position. Now, I'd like to tell you some things about the ear. But before we do that, I'd like to review with you some of the basic notions of sound and sound. Waves, how they're generated where they come from and how they travel through the air sound sources can be of varying kinds. Some examples are the reed of the clarinet, the bell, the human voice and a tuning fork. All of them produce sound in the same basic way. Let's review then the four essentials that we need for a sound to be created and to be heard. First of all, we need a vibrator such as a tuning fork. Secondly, we need some force to set the vibrator in motion. Thirdly, we need a medium such as the air to convey the wave motion created by the vibrator. And finally, we need the hearing mechanism itself which can receive and interpret the sound. I think the best example to explain how a sound wave is generated is the tuning fork. If you were to squeeze the tides of the tuning fork together and then let them snap apart, you would of course generate a pure tone. The reason why the tone is generated is because as the time springs back past its original resting position, it pushes the molecules of the air together and causes them to bunch up, it condenses them. Then as it springs back again, in the opposite direction, it creates a vacuum or a rare faction where there are fewer molecules than there should be. This constant repetition of condensation and rare faction increase in pressure, decrease in pressure is the basis for the sound wave. Now, it's important to remember that it's not the molecules in the air themselves that travel. But rather the pressure changes that travel through the air to the ear. There are two basic characteristics of a sound wave that I think you should be familiar with. The first is frequency, frequency is analogous to pitch. And it's defined as being the number of times the repetition of condensation and rare faction occurs in some unit time. For example, in 1 2nd, the fewer number of times in a second that the conversation and rare faction cycle goes on. Then the lower the pitch, the greater the number of times that this condensation rare faction cycle goes on. Then the higher the perceived pitch, if you will look at the screen, you will be able to see a representation of a pure tone that's been created by an oscilloscope. The uppermost portion of the sound wave is that represents the condensation portion. The lower most portion represents the rare faction portion. You can see that these waves are some distance apart and probably represent a tone with a frequency of about 250 cycles per second as we increase the tone so that the frequency is close to 1000 cycles per second, the waves come closer together. And we see more a greater number of repetitions of condensation and rare faction. In a second. The other aspect of a sound wave that I would want you to be familiar with is amplitude. Amplitude simply relates to the fact that the strength of the particle or molecular energy is greater. And so we have a louder sound. This is represented on the screen by the sound wave increasing in amplitude, you can see that the envelope becomes wider as the sound becomes louder. Then as the sound becomes softer, the envelope of the sound wave decreases and becomes smaller. Now, this example, from the tuning fork is a fairly simple representation of a sound wave and it's normally not one that we run into every day. I better a different example which I would like to show you is what the human voice would look like if you were to put it on a screen as I'm talking. Now, you can see the jumble of lines and angles on the screen representing my voice as it comes to you. It's, it's really a wonderful thing to consider the human ear that it can take such a jumble of noise and make sense out of it. Now, let's take a look at the year just how it's made and just how it works. The year is first of all located in the petrus portion of the temporal bone and this is the hardest bone in the body and nature designed it that way to be sure and protect the inner ear from damage. As a matter of fact, the word petrus derives from the Latin word petrus, which means rock, the outer ear is connected to the skull by a series of muscles ligaments and other tissue. Now, most of the muscles that attach the outer ear to the skull are vestigial. That is at one time, we used to use them the same way an animal uses them now to turn his ears to localize sound. But uh they are no longer of any real use to us at this time. The outer ear is the first thing that receives the sound. It's ovoid in shape and contains a number of depressions and humps. Its function is to collect the sound waves as they hit it and direct them down and into the external canal. It provides us with about 10 decib of amplification. Now, this amount of amplification is really not significant. As a matter of fact, if we were to lose the outer ear, it would not cause any real loss of hearing for us. So even the outer ear itself is not that useful to us today and serves primarily a cosmetic function. Now, as we move from the outer ear into the external canal, remember now that the sound waves coming through the air are collected by the outer ear and directed down into the canal where they finally will hit the ear drum. The external canal is about an inch long and it's about a quarter to a half inch in diameter. The tissue that uh is made that makes up the wall of the external canal is the same tissue that is connected with the tissue that makes that up that tissue of the ear, the hand, the back and so forth. There are two things about the external canal that I think you should be familiar with. One is that it contains a number of hairs along this portion here. And the function of those hairs is to protect it from insects intruding into the external canal. In addition to that, there are also in here, a number of glands which are called saruman. It's glands and they will produce saruman or what's more popularly known as earwax. Now, this earwax or saruman is very bitter tasting and repugnant material. And it also serves the function of protecting the ear from the intrusion of insects. Now, the sound wave comes down from the article into the external canal and finally, it comes up to the middle ear, the middle ear is made up of the ear drum, The articular chain which is made up of three tiny bones called the, asse. the Incas and the stay peas. The function of the middle ear is to transform the acoustic energy into mechanical energy and also to increase or amplify the amount of sound that that is reaching the ear. And it does this by means of what is called the a real ratio. That is if we take the ratio of the area of the Eardrum to the ratio of the area of this foot plate. Here, we find that the increase in sound is about 18 times. Now, it's necessary for this to occur because the sound after it leaves, the middle ear will go into the inner ear, which is filled with fluid. And it's necessary to overcome the impedance which is offered by the fluid, the fluid in the inner ear. There are two muscles in the middle ear, which we should be familiar with. And those are called the tensor tympani. And the step PDS muscle, the tensor tympani originates here and comes across the middle ear space to the handle of the malas. The step pds muscle originates on the posterior wall behind these bones and it inserts right here in the head of the stay peas. These two muscles serve a protective function that is whenever a sound which is loud enough to cause damage to the ear occurs, they contract very quickly and they pull the articular chain off its normal course of action. And so it protects the inner ear from uh from damage. Now, when the sound wave it comes down into the external canal and it hits the eardrum, it causes the eardrum to vibrate at the same frequency and amplitude as the sound as the original sound. The articular chain is connected to the eardrum. And so that also vibrates at the same frequency and amplitude. And so the sound wave is now conducted across this middle ear space up to the inner ear as as of yet though no, no hearing has taken place. The only thing that's really happened is that the sound wave has been conducted up to the inner ear. The other part of the inner ear that you should be familiar with is something called the U station tube. And this is located in the anterior inferior portion of the middle ear space. The station tube runs from the middle airspace down to the nasal firing. So to the back of your throat And it serves two functions. First of all, it provides a constant source of oxygen to the middle ear space. The second thing it does is to is to provide an equalization of pressure whenever there are changes in altitude. If you were to experience a severe change in altitude, for example, the pressure on the outside of your ear would be less than the pressure on your, on the inside of your ear. And it would force the ear drum outward. If you yawn, then you open the station tube and it allows the air to come up into the middle ear. And so it equalizes this pressure. I'm sure if you've flown in an airplane, you have probably experienced this and have had to equalize that pressure in order to get rid of the uncomfortable or stuffy feeling that you find in your ear. Now, once the sound wave again, has crossed the middle ear, it enters the inner ear it is in the inner ear that the essential organ of hearing is contained. The inner ear contains also the balance mechanism. And this mechanism is made up of the three semicircular canals, the sack you'll and the you trickle. And this mechanism, all of these parts working together provide us with a sense of balance to keep ourselves upright. This yellow line that you see here is the vestibular portion of the auditory nerve. The other portion of the inner ear or the labyrinth is called the cochlea. And this is a spiral or snail shaped bony labyrinth within which the essential organ of hearing is located. This portion of the inner ear is called the vestibule. And you can see that there are three channels here. This is the scale of vestibular or the vestibular channel. The scallop media or the middle channel and the skeleton pony or the tim panic channel. This area in here is filled with a fluid which is called parallel. And this middle channel floats in that parallel and it protects it from bumping up against the walls of the bony uh capsule. The fluid which is inside the scallop media is called indolent. And this is much different than the parallel and arises from somewhere near the cortex. Now, when, when we hear the acoustic signal, which has been changed to uh mechanical energy now has to come into the inner ear and be somehow changed into hydraulic energy and then into electrical energy. Now, what I would like to do is to show you a more detailed version of what the cochlea looks like and just what it's made up of. Now, this picture over here is a much more detailed version of the cochlea. Right here. We have the auditory branch of the 8th cranial nerve coming on up into the cochlea and branching off at every level. You can see the snail shaped appearance of the cochlea. And this picture shows it as though it had been cut straight down the middle. The arrows that you see indicate the direction uh the force, the direction of the force of the energy which is created by the sound wave, it moves on up and then at the top turns around comes back down. Again. Now, here's an even more detailed picture of of the cochlea. This is just one section of it. And again, you can see a fiber from the eighth cranial nerve coming on through the spiral lamanna to connect with the essential organ of hearing. This is the scalar vestibular, the scalar tympani and the scalar media within the scalar media contains, as I said, the essential organ of hearing, the scalar media is bounded by prisoners membrane here, the base layer membrane here and the spiral ligament here within the scalar media, we find the sectorial membrane and the organ of corti, the pink area in the organ of corti. You'll see hair cells located along its whole length. Each of these hair cells is connected to the nerve of hearing when they say no comes from the middle ear into the inner ear. A number of things happen. First of all, the stay peas moves inward as it does, it compresses the fluid in the inner ear. Now that pressure has to be released somewhere. And right down here in the skeleton pony is something called a round window. The staples moves in through the over window, compresses. The fluid causes it to move. That pressure is released at the round window. So the signal again comes by way of the article to the external canal. It has changed to mechanical energy and amplified across the time panic membrane of secular change structure. And now it is causing the fluid of the inner ear to move. And so now it has been changed into hydraulic energy. You can see the dotted lines here and they indicate that as that fluid is moving through the inner ear, it causes this membrane to bulge inward. That membrane causes the fluid in here to move. That in turn causes the tech tutorial membrane to bend downward. And now you see the hair cells here which are connected between the organ of corti and the tech tutorial membrane. When the tech tutorial membrane bends downward, it distorts or bends these hair cells out of shape as soon as it does that, that triggers off an electrical signal, Which is then picked up by the fibers of the 8th cranial nerve and carried on up through the central nervous system to the cortex. The movement of the fluid, of course, being pressure continues on. And you can see the basilar membrane has bent down also that causes this fluid to continue moving. And if we can go back over here again for a minute, it would be released here at the oval window. Let's take a look at schema ties version of the inner ear. Remember the inner ear is coiled snail shaped of 2.5 turns. And so we've schema ties it a little bit here in order to show you in some detail, just how it works. The sound comes down through the the external canal hits the eardrum that is connected to the articular chain. Now, the foot plate of the stay peas moves into the the vestibule and it compresses this fluid in here. Now, you can see that at this point, this continuation or movement of fluid through the inner ears taking place, risers membrane is bending downward. The fluid continues to compress it causes the basal membrane to bend and comes right on down through and finds its release here at the round window. These arrows here indicate the pressure which is not necessary here or not, uh which is not necessary at this point going on up around through the whole system and eventually coming out through the round window. The basil er end of the inner ear contains those uh hair cells and fibers which are responsible for low pitched sounds. I beg your pardon for high pitched sounds. The a pickle end up here is responsible for those uh hair cells and fibers which are responsible for low pitched sounds. Once the the hair cells have been excited and trigger off this electrical response, they will excite the fibers of the eighth cranial nerve And they all come together in a bunch of, of nerve fibers, which is called the spiral ganglion. And then they find their way on up through the central nervous system to the cortex where the sound is interpreted. Now, what we've been discussing so far is the way we hear by what is called air conduction. That is the signal travels from the air directly into the ear canal into the inner ear. This is the way that we normally hear in most everyday situations. There is another way though that we can hear and it's called hearing by bone conduction. Now, if we can take a look at this chart, again, recall that the inner ear is encased in a capsule of bone and that bone is part of the skull. When a sound wave is strong enough to cause the bones of the skull to vibrate, those vibrations can cause the fluid in the inner ear to move and to carry out the same essential process as happens by air conduction. Now, this is normally not important for us as long as the outer ear and the middle ear are intact. But for those persons who have some pathology of the outer and middle ear, who may perhaps need a hearing aid, we can provide a hearing aid that would put a vibrator right here on the skull and that would cause the bones of the skull to vibrate at an in an amplified way, excite the inner ear. And so allow him to be able to hear normally. Now, let's just take a look again at the whole process of hearing by air conduction. Remember that we have a signal which is generated, for example, by a tuning fork, it generates a number of compensations and rare factions changes in pressure. Those pressure changes find their way through the air are collected by the external ear and forced down into the ear canal. They reached the eardrum secular change structure where they are changed to mechanical energy and amplified. They then find their way into the inner ear when the foot plate of the staples moves in and out in time to the signal that came into the year originally. And that causes the fluid to move, causes the membrane structure to move. And so excites the nerves of hearing from there. They then find their way on up through the central nervous system to the cortex where they're received analyzed and interpreted. Now, that's essentially how the ear works. Uh it's come a long way since the time when it was a simple system for that in a fish millions of years ago. So that the fish could detect the pressure changes in the water and maintain an upright and balanced position. We've learned a lot about it since the early days of, since 1950, with, with the invention of the electron microscope, but we still have a lot more to learn and I'm sure that in the next few years we'll, we'll see a lot more being added to our store of knowledge about the year and how we hear.