S~£:^>^;^*:^: HUMAN PHYSIOLOGY: '**' DESIGNED FOR COLLEGES AND THE HIGHER CLASSES IN SCHOOLS, FOR GENERAL READING. 1/ BY WORTH1NGTON HOOKER, M. D., PROFESSOR OF THI THEORY AND PRATTIC*. "K >1LDICINE IN TALE COLLEGE; AUTHOR OF " PHYSICIAN AND PATIENT." 3 ^EON Illustrated by nearly 200 Engravings. /-n~->>___ NEW YORK PUBLISHED BY FARMER, BRACE, & CO. NO. 4, COURTLANDT-STREET. 1854. 2 5- Entered, according to Act of Congress, in the year of our Lord, One thousand eight hundred and fifty-four, BY WORTHINGTON HOOKER, H. D., In the Clerk's OfRce of the District Court of Connecticut. R. H. HOBBS, pt j^ At-VORn Stereotypy, Hartford, Ct. Printer,'New York. - - 123 CONTENTS. PART I. CHAPTER I. Pack Organized and Unorganized Substances, ... 13 CHAPTER II. The Distinction between Animals and Plants, . . 21 CHAPTER III. Man in his relations to the Three Kingdoms of Na- ture, ..............27 PART II. CHAPTER IV. General views of Physiology, with a Brief Account of some of the Structures of the Body, . . 35 CHAPTER V. Digestion,..............42 CHAPTER VI. Circulation of the Blood,.........64 CHAPTER VII. Respiration,.............86 CHAPTER VIII. Formation and Repair,.........109 CHAPTER IX. Cell-Life, .............123 IV CONTENTS. PART III. CHAPTER X. The Nervous System,.........139 CHAPTER XL The Bones,.............1™ CHAPTER XII. The Muscles,............196 CHAPTER XIII. The Language of the Muscles,......222 CHAPTER XIV. The Voice, ......'.......243 CHAPTER XV. The Ear,..............271 CHAPTER XVI. The Eye,..............287 CHAPTER XVII. Connection of the Mind with the Body, . . . 318 CHAPTER XVIII. Differences between Man and the Inferior Animals, 347 CHAPTER XIX. Varieties of the Human Race,......367 CHAPTER XX. Life and Death,...........381 PREFACE. I have aimed so to write this book, that it shall be fitted both for gen- eral reading, and for instruction. It is designed for the family as well as for the school. It seemed desirable that these two objects should be ac- complished at the same time, and I have not found them to be at all in- compatible. The instruction needed by the family on this subject, differs not from that which is required in the school-room, either in regard to the facts to be communicated, or the manner in which it should be done. No one will question the truth of this, so far as the facts are concerned. But it is true even as to the mode of communicating them. In both cases there needio be clearness in statement, arid fullness of illustration. Actual instruction is to be given in both cases, and to minds that are very nearly in the same attitude. I could not, therefore, see the necessity of writing a book on this subject for the people which should differ from one written for the school. Besides, it has seemed to me desirable that there should be a greater community of interest between the school and the family than as yet exists; and this object books equally interesting to both will tend to promote. It may be proper for me to say a word in relation to the style of the work. I have adopted the style of the lecture-room, because, that while it is not inconsistent with conciseness, it is the more natural mode of in- struction, especially when so much reference is made to illustrative figures. It has enabled me also to keep in view more effectually the attitude of the minds I address. I have had my readers before me continually as an imaginary audience. I have avoided technical terms as far as possible. Whenever they are used they are sufficiently explained at the time, so that no glossary is needed. Some points commonly considered hard to be understood are treated of, but I have endeavored to simplify them, by vi PREFACE. full illustration, and by a presentation of the truth uncomplicated with speculations and hypotheses. And these points are so introduced, that the mind is prepared by the previous investigation to understand them. I have aimed so to arrange the topics, as to have a preparation constantly going on in the mind of the student, fitting him for the proper under- standing of what is to come after. By this natural gradation in the de- velopment of the whole subject some of the deep things in Physiology can be made clear, which it would otherwise be impossible for the student to understand. It is proper to state here, that I intend to prepare a work for younger scholars, in which some of the simple points in Physiology will be illustrated. This, by familiarizing their minds with the subject, will fit them for a more thorough understanding of the present work. Although Physiology is becoming a prominent study in the schools and colleges in some parts of our country, its importance is no where as yet appreciated as it should be. It should be made a regular branch in our Educational System. This has been already done in France. " A com- petent knowledge," says Carpenter, " of Animal Physiology and Zoology is there required from every candidate for University honors ; and men of the highest scientific reputation do not think it beneath them to write elementary books, for the instruction of the beginner." The importance of Physiology as a study, will appear from various con- siderations. Many of the subjects comprised in Physiology have, in the case of most students, been already studied in a different phase, or mode, in other branches. Thus, if the student has attended to the Mechanical Powers in his Natural Philosophy, he finds in the human body the principles of the pulley and the lever illustrated in great variety and perfection. The principles in relation to strength in the form and arrangement of struc- ture he sees exemplified in the frame-work of the body in the most ad- mirable manner. If he has studied Hydraulics, he sees in the body the most perfect, and at the same time the most complicated hydraulic ma- chinery, working incessantly throughout life in the circulation of the blood. The principles of Pneumatics he finds applied in the respiration— those of Optics in the eye—those of Acoustics in the ear—and those of Musical Sounds in the apparatus of the voice. And then, his chemical PREFACE. Vll knowledge meets with new applications in his observation of the changes and the processes going on in the body. The relations, then, of Physiology to some of the common branches taught in the higher classes in schools, are of the most intimate charac- ter. Physiology, in part, merely extends these branches into a new and interesting field 5 and the student who has once entered this field recurs to these same branches with a renewed interest. Hydraulics, Pneuma- tics, Optics, &c, have now a new attraction for him, from this, to him novel, application of their principles. The interest thus awakened in his mind is worth much in itself, aside from the mere addition- made to his knowledge. And the interest is enhanced by the consideration, that in the human body he sees the applications of these principles to mechan- ism that exhibits the skill of perfect wisdom and almighty power. But there are relations of Physiology to still other studies which should be noticed. The analogies that exist between the human body and all other living things, in relation to structure and growth, are numerous and striking. Though life is so diverse in its processes and in the forms which we see it evolve in the whole range of animated nature, it in some important re- spects displays a great similarity, which it is interesting to trace through- out its diversified manifestations. Growth, or nutrition, as you will see | in the following pages, is essentially the same in the Plant as it is in the Animal. Botany, therefore, taught as it should be, has quite an intimate relation to Animal Physiology. The Science of Life is, in many respects, one Science; and if, in studying any of its subdivisions, we fail to take this broad view of it, and to trace out the analogies referred, to, we lose a large part of the interest of the study. Human Physiology, the subject of study in this book, is but a part of a science which offers to the student wide fields of observation exceedingly diversified and full of interest. This being so, I could not avoid in the following pages making occasional reference to the analogies existing between the phenomena of life as ex- hibited in the human system, and those which we see in the living world around us. So that as the student proceeds with the study, he will find himself interested in the phenomena of life in whatever form they are presented. ■•,/■' '' '['■ VU1 PREFACE. This leads me to say that this study of nature, in its broad common re- lations and its beautiful and extensive analogies, should be made very prominent in our systems of education. It is the application of the prin- ciples of abstract science to the forms, and especially the living forms of nature all about us, that gives Interest to these principles, and makes us to understand and appreciate them. It is here that we find a very seri- ous defect in the prevalent mode of education, even at the present time, notwithstanding all our improvements. Let us look at it a moment. We live in the midst of a material world, animate and inanimate, and have daily converse, so to speak, with material forms of every variety, present- ing phenomena of the highest interest and of endless diversity. And yet, through almost all the period of childhood, and perhaps we may say youth also, this book of nature is in the school-room very nearly a sealed book. The very process of education shuts in the pupil from this broad contemplation of the world in which he lives. He is drilled through spelling, reading, grammar, &c, but he is left in total ignorance of the beautiful flowers, and the majestic trees outside of the school-room. How very few even of thoroughly educated adults, know the processes by which a plant or a tree grows! And the same can be said of other phenomena of nature. The defect which I have pointed out runs through the whole of educa- tion. We can see it even in the prevalent mode of teaching the natural sciences themselves. One would suppose that here the facts, the phe- nomena, would command the chief attention of the teacher and the stu- dent. But it is very commonly not so. The mere technicalities and the classification are made much too prominent. Botany, really one of the most interesting of all branches of natural science, is thus ordinarily made one of the driest of studies. To teach this aright, the phenomena of vegetation, so varied and so beautiful, should constitute the chief ma- terial of instruction, and the mere classification should be considered al- though necessary, as wholly a secondary thing. The great facts of the world, both of mind and matter, should furnish really the material for education, and those branches that are ordinarily pursued with such assiduity should be considered as merely subsidiary to the teaching of these facts. The whole order of education must be re- PREFACE. IX versed. Instead of beginning the child's education with learning to spell and read, the object should be to make -him an observer of nature, and the spelling and reading should be done in connection with this, and as subsidiary to it. Things and not words, or mere signs, should from the first, constitute the substantial part of instruction. The child should be made, at home, in the school, and everywhere, a naturalist in the largest sense of that word. We should aim to impart to him a spirit in con- sonance with the following precept of Hugh Miller, the famous self- taught geologist. " Learn to make a right use of your eyes; the com- monest things are worth looking at—even stones and weeds, and the most familiar animals." As it is now, no one becomes a naturalist early in life, except in spite \ of the tendencies of his education. The study of nature is not only not i encouraged, but is absolutely discouraged in our educational system. If any one, like Hugh Miller, by the force of a taste that can not be repress- ed by the training of the school-room, undertakes to make a " right use of his eyes," and curiously examines '^ stones and weeds," he is regard- ed by the world of spellers and readers and grammarians and cipherers, as a strange genius. But he is pursuing from an irresistible internal force, the very course that I would have every student, even from his childhood, encouraged to pursue, in a measure at least, by the external circumstances of his education. The tendencies of his training should be decidedly in this direction. If the general mode of education were changed in the manner indicated, education would have much less of the character of mere drudgery than it now has. Not that there would be any the less labor; but the labor would be made lighter by the interest imparted to it—the interest, which always results from the study of facts and phenomena, and never from the learning of mere words and technicalities and classifications. I would gladly dwell on this subject, and show by varied illustrations how the mode of instruction referred to, should be pursued, and especially with younger scholars; but the limits of a preface will not allow me to enter so large a field. The change which I have pointed out can not be effected at once. It will require time. Confirmed traditional customs are to be done away, X PREFACE. the habits of teachers are to be altered, and-the proper books are to a great extent to be yet written, especially such as are fitted for the first years of education. If the study of nature should be thus made prominent in education, human physiology would be considered altogether its most interesting and important branch, and for several reasons. First: there is no where to be found so curious a collection of mechanisms, or so interesting and wonderful a series of processes, as in the human body. In nothing else in the wide world are the principles of so many departments of science so extensively and perfectly exemplified. Life works here its most com- plicated set of machinery. Secondly: the singular and mysterious con- nection of the immaterial and immortal soul with the material and perish- able body, gives intense interest to this study. In Physiology we do not study matter alone, or spirit alone, but both matter and spirit united, and often acting together. This circumstance distinguishes this from all other studies. Thirdly: it is our own frames, moved by the spirit within us, that we study. The subject has a personal interest for us, that is not presented by most studies, and by none in so large a degree as in this. And Fourthly: the study is of great importance, because a judicious and efficient Hygiene must be based upon a knowledge of the laws of physi- ology. We cannot know how to keep our functions in the condition of health, without understanding the laws that regulate them. I have said but little in this book in regard to hygiene, and that only incidentally, be- cause that subject would require of itself a whole volume to elucidate it properly. I have not thought it proper to indulge to any great extent in those re- flections, which the contemplation of so perfect and diversified a congeries of mechanisms as are presented in man would naturally suggest, in regard to the skill of the great builder of the universe. Such reflections would extend the book to too great length. Besides, they are so readily sug- gested to the mind of both teacher and scholar, that it is entirely un- necessary for the author to dwell on them. I have treated of some subjects, on which, from the difficulty of un- derstanding them, there has been a disposition in many minds to go be- yond what we know, and indulge in unwarranted speculation. On these PREFACE. XI points I have taken pains to draw the line very distinctly between what is known, and what is supposed. I deem it to be important to prevent the minds of the young from being led away from the simple truths of science by ingenious speculations and plausible reasonings. Let me not be understood to decry all hypothesis. I only object to the mingling of # facts and suppositions together in one indiscriminate mass, as is often done. The disposition to do this, which is more common than is generally supposed, exerts so injurious an influence upon the habits of the mind, and so confuses its views of truth, that we ought to look upon it as one of the most serious evils to be guarded against in education. It is really one of the most prominent obstacles to the progress of truth on all sub- jects, both in individual minds, and in the minds of the community at large. This disposition, so apt to be fostered in the enthusiastic mind* of youth, by ingenious but dreamy speculations, should be corrected at the outset, and the mind should in its forming stage, be habituated to the dis- crimination between the proved, the true, and that which rests on pre- sumptive, perhaps merely plausible evidence. This discrimination should therefore be exemplified in books designed for instruction, and this I have attempted in the present volume. I have divided the book into Three Parts. The First, which I have made as short as possible, is merely preliminary to the consideration of the particular subject of the book. In the Second Part, I present the human structure, simply as a structure, and show how it is constructed and kept in repair. In the Third Part, I treat of all those subjects which relate to the uses for which the structure is designed. This natural division of the whole subject, not only presents it to the mind of the student in an interesting point of view, but secures that natural grada- tion in its development, which I have spoken of as being necessary to a clear understanding of its deeper and more intricate portions. PHYSIOLOGY. PART FIRST, CONTAINING, CHAPTER I.—Organized and Unorganized Substances. CHAPTER II.—The Distinction between Animals and Plants. CHAPTER III.—Man in his Relations to the Three Kingdoms of Nature. CHAPTER I. ORGANIZED AND UNORGANIZED SUBSTANCES. 1. The crystal and the plant are both wonderful growths. As you look at them, you think of the crystal as having been formed, and of the plant as having grown. But in one sense they have both grown to be what they are. The crystal was once a minute nucleus, and the plant was once a little germ. 2. -In one respect they are alike in their growth—both have increased from particles taken from things around them. But the processes by which this is done are different in the two cases. The crystal has increased or grown by layer after layer of particles. There are no spaces or passages by which parti- cles of matter can be introduced inside of it. Any part of it, when once formed, is not altered. It can receive additions upon the outside alone. But it is not so with the plant. This enlarges by particles which are introduced into passages and interstices. It grows, as it is expressed, by absorption or by in- tussusception. 3. How, now, is this absorption effected ? It is done by cer- tain vessels or organs, constructed in the root of the plant for this purpose. These take up or absorb fluid matter from the earth. There are other organs which circulate this fluid through all the plant; and others still which use it for the purpose of growth or formation. There are no such organs in the crystal, for it has no inner growth. The plant is therefore said to be an organized substance or being, and the crystal is an unor- ganized substance. And so we speak of the organic structure, or the organization of plants. 2 14 HUMAN PHYSIOLOGY. Organized beings. Mechanical, chemical, and vital principles.___________ 4. These organs, which thus absorb, and circulate, and con- struct, do not act simply on mechanical principles. Ihe plant is not merely soaked with fluid, which the heat of the sun may expel, as it does water from a porous mineral substance. Inese organs are active agents, and they perform their duty with a force, and after a manner, for which no mechanical principles can account. No mechanical powers could alone supply the leaves of the mighty tree of the forest with sap from its deep roots; much less could they form these leaves. 5. Neither do these organs act simply on chemical princi- ples. While man, through the agency of chemistry, can form some of the crystals which are found in nature, he can not by any arrangement of constituents make a plant, a flower, or a leaf. And the plant, left alone to the action of chemical prin- ciples, wilts; and at length ceases to be a plant, and becomes common unorganized matter. 6. Mechanical and chemical principles, it is true, are both employed to some extent in the growth of plants ; but they are under the control of other principles, which we term vital. And so we speak of the plant not only as an organized substance, but as a living being. 7. What I have said of plants, in distinction from minerals, may also be said of animals. They are also organized living beings, and they have generally a more complex organization than plants, as you will see as I proceed. 8. The whole material world, then, that we see around us, we divide into two parts—the unorganized and lifeless, and the organized and living. The distinctions thus pointed out be- tween organized and unorganized matter are essential and fundamental. But let us look at some other distinctions, which either arise from these or accompany them. 9. One distinction is this. All the parts of the mineral are independent of each other, while it is otherwise with the plant or the animal. Accordingly, we examine the properties of min- erals in a different way from those of plants and animals. The chemist can ascertain all the properties of a crystal or a rock, if you give him but a small piece of it. But the botanist can not ascertain all the properties of a plant by looking at some one part of it. If he examine the flower, this gives him no knowledge of the root. In order to know all about the plant, he must examine every part by itself, and then look at it in its relations to the other parts. The same can be said of the physiologist, in his investigation of the properties of animals. ORGANIZED AND UNORGANIZED SUBSTANCES. 15 Assimilation in organized substances. 10. As the crystal is forming by layer after layer of particles, no change is effected in these particles as they are becoming arranged in the layers. But in the case of the living organ- ized being, a change is produced in the particles which are taken up by the absorbents. And the change, ordinarily, is both a gradual and a complex one. In the plant, a change is produced in the particles in the very act of absorption; but this change is only the beginning of a process which is after- wards perfected. The sap is not thoroughly fitted for nutrition when it first begins to circulate. It is carried up through the vessels of the trunk or stalk to the leaves. There the last step of the process is taken, and the sap is now ready to be used in the growth of the plant or tree. So, also, in the animal, the nutritious part of the food, taken up by the absorbents in the digestive organs, is first acted upon by certain little glands, through which it passes, is then poured into the circulation, to be mingled with the blood, and is carried with the Wood to the lungs, to be exposed to the air; and thus it is fitted for the nu- trition or growth of the body. This process, which is thus car- ried on in the plant and in the animal, is very properly called assimilation. For the particles that are taken up by the ab- sorbents in the root of the plant are, by this process, made like to the plant; and the particles taken up by the absorbents in the stomach * are made like to the "animal. So obvious is this, in the case of the animal, that some French physiologist speaks of the blood as chair coulante, or running flesh. 11. Another prominent distinction between organized and unorganized substances is in relation to permanency. Constant change appears in all organized bodies; while permanency is written upon all substances which are unorganized. In organ- ized beings, continual change is going on at every point. It is a condition of their being. This is true, not only of the de- cline of a plant or animal, but even of its growth. For, in its growth, as the parts enlarge internally as well as externally, they change not only the arrangement of the particles, but, to a great extent, they change the particles themselves. It is * The word stomach requires some little explanation, as it is used in physiology in two senses—in a limited sense, and also in an extended one. It is used in its limited sense, as referrin" to the cavity at the beginning of the alimentary canal, as it is termed; this lat- ter term bein" applied to the series of cavities, the stomach and the small and large intes- tines which are found in the digestive apparatus in the higher orders of animals. In comparisons, however, between these animals and those whicli have a more simple digest- ive apparatus, the word stomach is used in a more extended sense, as being synonymous with the term alimentary canal. It is used in this sense, also, when, as in the present case it is referred to in a comparison between animals and vegetables. 16 HUMAN PHYSIOLOGY. Organized substances changing. Unorganized permanent.____________^ true, as well of the towering tree as of the tiny plant, that these changes have been going on during all its growth; so that, at its maturity, it is, both in relation to the arrangement of its particles, and in relation to the particles themselves, a very different thing from what it was when it pushed its germ up through the ground, or even when it was but a small tree. Not only has it received into its interstices and passages new particles, but it has thrown off from the pores of its leaves, those outlets for the refuse of plants, vast quantities of parti- cles which are no longer of use in its structure. So, in all animals, the same internal changes are going on, and to a much greater extent; because, from the activity of their na- ture, there is more of wear and tear, and, therefore, more of refuse matter to be disposed of. As you will see in another part of this bools, the human body, that most complicated of organized beings, undergoes these changes very largely. 12. It is not thus with unorganized substances. The crys- tal, so fast as it is formed, becomes permanent. No changes occur within it. In itself, it is unchangeable. It can not change its own particles, as the plant or the animal does. It can be changed only by external addition, or by external dimi- nution, through the influence of agents acting upon its surface. 13. With the constant changes going on in organic nature, there is constant succession. Plants and animals produce other plants and animals, and themselves die, making room for their successors. But the crystal does not form other crystals, and then crumble into dust. In itself, it is both unchangeable and unproductive. 14. This distinction between organized and unorganized sub- stances, in relation to change and succession, meets the eye everywhere. The mountains, the rocks, and even the stones under our feet, remain the same year after year, while all vege- table and animal life is ever changing its forms and manifesta- tions. There are the changes of growth, and the changes of decay and death, all around and within us; and they are strangely mingled together. There is death even in the changes of life, as the waste particles are taken away, and are replaced by the new; and life springs out of the very bosom of death, as from decayed nature new forms of vigor and beauty arise. The mountains stand as they have stood, as the passing generations have looked upon them, while the continual changes of vegetation have been going on upon and around them. The seasons crown their battlements with the emblems ORGANIZED AND UNORGANIZED SUBSTANCES. 17 Different forms of organized and unorganized substances. of their ever-returning mutations of life, decay, and death; and even the mighty trees, that have shed their leaves from year to year, in obedience to the great law of change, but have themselves stood, at length bow their heads to the same law, and give place to other lords of the forest. From the " ever- lasting hills," which thus remain the same, though change is ever about and upon them, man gets the unchangeable and imperishable rock to construct his habitation, while he himself is changeable and perishable—the creature of a day, whose life is as a vapor. He wears the precious stones, and traffics in the golden ores, which have existed from the creation of the world, through all the changing and dying generations, and passes away, leaving them to others as changeable and perish- able as himself. 15. Another distinction between organized and unorganized substances relates to the forms which they assume. There is regularity in both, but it is different in each. Unorganized matter is disposed to arrange its particles in straight lines, and with angles mathematically exact. You see this in the beautiful crystal; and you also see it, less definitely, but magnificently, displayed in the regular battlements and columns of rocks and mountains. The tendency is to regularity; and irregularity is the result of interfering circumstances. A similar disposition to regularity is manifest in organized substances, but in a different manner. It is disposed to curved, rather than straight lines, and seldom makes lines or angles with mathematical ex- actness. We see this law of regularity exemplified both in animal and vegetable life. The leaf, for example, has the same general shape, that is, the same general arrangement of par- ticles, when it attains its full size, that it had when it was small; and the same can be said of the arm of the man, com- pared with his arm when a child. Illustrations might be cited to any extent, but these are sufficient. 16. While the law of regularity is not commonly as exact in organized substances as it is in the unorganized, it is quite as authoritative. While it does not ordinarily observe the per- fectly straight lines and the unvarying angles which we always find in the crystal, the general plan and contour are very strictly preserved amid all the changes of animal and vegetable life. And, in some cases, the same mathematical exactness that we find in the mineral world is found in organized beings. I know not that this is ever true of straight lines and angles; but it is often true of curved lines. There are many very 2* 18 HUMAN PHYSIOLOGY. Regularity in form—in some cases wonderful. beautiful examples in the vegetable world. I will give but a single one. If you look at the common white daisy, before the hundreds of little buds in its bosom have opened into tiny flowers, you will see them arranged with great exactness in crossing curved lines, such as you often see on the back of a watch case. A similar arrangement you will find in many flowers. 11. This regularity is more wonderful in organized sub- stances than in the unorganized, because it rules in them in the midst of constant change. In the case of the crystal, as there are no internal changes in it, and as each layer of it, when formed, is permanent, regularity is comparatively, so to speak, easily secured. But in the case of the leaf, as it is growing to its full size, and of the arm, as it grows from infancy to be the stalwart arm of manhood, continual change is going on at every point; and regularity here is obviously a more difficult achievement. 18. This regularity appears still more wonderful, when we look at the infinite variety of forms in organized matter, in both the vegetable and the animal world. In all these forms, each part of every animal and of every plant maintains its own peculiar plan and contour. Take, for example, the leaf in its endless varieties. How definitely does each variety preserve its individual character, and how easily is it distinguished from every other variety! The same can be said of every part of every organized being. 19. Another circumstance still must be mentioned, as adding to the wonderfulness of this regularity. It has been scrupu- lously maintained, through all the changes of the world from its creation, when God pronounced the works of his hands to be " very good." The leaf of every tree, for example, is like the leaf of its ancestral trees back to that time; and so it will be in all its successors to the end of the world. " The trees of the garden," which delighted the eyes of our first parents, and refreshed them with their shade in their innocence, and amid which they hid themselves after their sin from the presence of their Maker, undoubtedly had the same characteristic shapes, and the same leaves and flowers which their successors present to our eyes. 20. Again, it is interesting to notice that, in the midst of this regularity, so strictly maintained in each specific form from age to age, there is a measure of irregularity allowed. While each kind of tree, for example, has specific characteristics in ORGANIZED AND UNORGANIZED SUBSTANCES. 19 Variety of form; yet regularity preserved. Size. the arrangements of branches and other parts, and in the shapes of its leaves, no two trees of the same kind are exactly alike, and there is always much variety in the leaves of the same kind. The wonder is, that so much latitude is allowed in this respect, and yet the specific characteristics of each kind are so thoroughly preserved. We can readily see that if a pattern, definite in all its details, were to be copied exactly in each kind of vegetable and animal form, the distinctions between them could be more easily preserved. But Omnipotence is able to combine a wide latitude and variety of form in each kind, with a strict and uniform preservation of its characteristic contour and arrangement. We have a striking exemplification of the above remarks in the variety of the human countenance. While the face of man is so entirely different from the face of every other animal, at the same time, among the hundreds of millions of the human family, how uncommon it is to find two faces that are very nearly alike. 21. In the animal world, we see remarkable examples of the preservation of regularity of form in the exact correspondence which exists so commonly between the two halves of the body. For example, the brain has two halves, which are precisely alike, and the same is true of the nerves which are distributed from it. And so of other parts. But, mingled with this symmetri- cal arrangement of parts, there are other parts which are irreg- ular in their shape. This is the case with the stomach, the heart, the liver, etc. There are some animals which are alto- gether destitute of this arrrangement of two similar halves^ of the body. The oyster is a familiar example. The shell of this animal is strikingly in contrast, in this respect, with the shells of some other of the bivalve tribe, as, for instance, the common clam. 22. There is a distinction between organized and unorganized substances, in regard to size, which must not pass unnoticed in this connection. The size of unorganized bodies has no fixed limit. A crystal or a rock may grow to any imaginable size,- if the particles forming it are sufficiently abundant. But or- ganized bodies have limits fixed to their growth. There is, it is true, more or less latitude to these limits; but they are so well defined in the case of most vegetables and animals, that when growth reaches much beyond or below the limit, it is recognized as a remarkable fact. Gigantic and dwarfish vari- ties are rare exceptions to the general rule. 23. The last distinction, between organized and unorganized 20 HUMAN PHYSIOLOGY. Difference between organized and unorganized in structure and elements. substances, which I shall mention relates to their structure. While unorganized substances are made of one form of matter, either solid or liquid, or gaseous, organized bodies are made of a mixture of fluids and solids. They are therefore more or less soft and flexible; while the solid, unorganized substances are hard and brittle. There is a still further difference in struc- ture. Organized substances are much more compound than the unorganized. Most of the unorganized substances are composed of only two or three elements. Thus, air is com- posed of oxygen and nitrogen, water of oxygen and hydro- gen; and most of the mineral salts are composed of three elements—as, for example, carbonate of lime, or chalk, which is composed of oxygen, carbon, and calcium, the mineral base of lime. But organized substances are composed of at least three or four elements, and sometimes more. The four princi- pal elements in the composition of organized bodies are, oxygen, nitrogen, hydrogen, and carbon. But there are other elements introduced for special purposes. Thus, carbonate of lime (a combination of calcium with two of the common elements, carbon and oxygen,) is diffused very generally throughout the textures of plants, giving them firmness and strength. In the grass tribe, silex is deposited under the surface, producing the necessary combination of strength and lightness, a very small quantity of the silex answering the purpose. In animals of the higher orders, phosphate and carbonate of lime compose in part the framework of the body. We find iron, too, in the blood. Of the fifty-four elementary substances discovered in mineral bodies, only eighteen or nineteen have been found in plants and animals, and some of these in very small amounts. The essen tial components of living substances are the four non- metallic elements mentioned above—oxygen, hydrogen, nitro- gen, and carbon; while the bulk of the inorganic world is composed of the metals and their compounds, viz., the alkalies and the earths. And it is interesting to observe that, of the four elements which compose the bulk of the animal and vege- table world, both the fluids and the solids, three are gaseous, while but one, carbon, is a solid substance. DISTINCTIONS BETWEEN ANIMALS AND PLANTS. 21 Locomotion. Stomach and other central organs. CHAPTER II. THE DISTINCTIONS BETWEEN ANIMALS AND PLANTS. 24. Having pointed out in the first chapter the distinctions between organized and unorganized substances, I now proceed to consider the distinctions between the two classes of organ- ized beings—animals and vegetables. I shall first notice those differences which are obvious when we look at the great major- ity of animals and vegetables; and shall then point out those which are essential, in order that we may have a clearer view of those exceptional cases, in regard to which it is somewhat difficult to decide to which of the two kingdoms they belong. 25. One of the most obvious distinctions is in relation to locomotion. The plant remains in one place ; while the animal moves about, in the air, or in the water, or upon the surface of the earth. And the structures of the animal and the plant of course differ, so as to accommodate these two very different modes of existence. I will particularize. As the animal moves from place to place, it must, for this reason, if for no other, have an apparatus of nourishment and growth different from that of the plant. The plant, by means of its absorbents in the roots, takes up from the earth, in the form of sap, its nutri- tion, or food, as it may very properly be called. The moving about of the animal would in itself forbid its deriving its food directly from the earth, even if the earth contained the proper materials for its nourishment. Another contrivance must therefore be resorted to, in order to effect nutrition in its case. So a cavity is provided in its body, called a stomach, into which nutritious substances can be introduced. And this cav- ity is lined with absorbents, which there do for the animal just what the absorbents in the roots of the plant do for the plant. 26. Besides the stomach, there are other great central or- gans which are peculiar to most animals, in distinction from vegetables—as the heart, the liver, the lungs, &c. In the plant, there are no such central organs upon which the whole plant depends. Branches and roots may be cut off extensively, and even a large portion of the stem or trunk may be des- troyed ; and yet what remains of the plant may still live. And 22 HUMAN PHYSIOLOGY.____________ Feeling. Motion. Sensitive plant and catch-fly.________________ even more than this. A small portion of it may be made to take root and live by itself. It is not so with most animals. Mutilation can not be carried far without injuring some large organ which is essential to the life of the whole; and no part taken from its extremities can be made in any way to live by itself. 27. Another obvious distinction is this. Animals are sen- tient and spontaneously-moving beings, while vegetables are not. The animal feels the action of agents upon it, and this it can not do without consciousness and thought. The evidences of the existence of consciousness and thought, and the consequent spon- taneous motion, are very slight in some animals. Still, there is no doubt of their existence in these cases. We see these evidences plainly in the great majority of animals; and we infer, very properly, the existence of sensation and thought in those excep- tional cases, where the evidences are doubtful or absent, as we find in them other marks of animal in distinction from vege- table life. 28. The distinctions which I have mentioned are those which we see generally existing. Let us see how far they are essential and universal. 29. The distinction in regard to locomotion, if we look at the animal as a whole, has its exceptions. There are some animals that are entirely confined to one spot during all their existence, as the coral animal and the sponge. But, while some animals are thus confined, they have the power of spon- taneous motion in some of their parts, which is exercised for the purpose of obtaining food, and, in some cases, for the avoidance of danger. This power is not possessed by any plant. Some few plants, as the sensitive plant and the venus catch-fly, (dionaea muscipula,) exhibit a property which resembles it, but it is essentially a different thing. In these cases, the influence of the stimulus that excites the motion is communicated from particle to particle, from the point where the stimulus is ap- plied ; and the motion is only in one direction, and not in various directions, as is the case with spontaneous animal mo- tions. This can be very readily seen, if we compare the motion of the sensitive plant or the catch-fly with those of the little fresh- water polype, called the Hydra. This animal, of which I give you here an enlarged representation, and also a representation of its natural size, is found in ponds. It attaches itself to any floating object—a stick or straw, as seen in the Figure—by a kind of sucker. Thus supporting itself, it stretches out its long DISTINCTIONS BETWEEN ANIMALS AND PLANTS. ~ 23 Digestive cavity. Nervous system. None in plants. arms, to take for its food any minute worm or insect which may float within their reach. When it catches one, it directs it to the mouth, a, which opens into the stomach or general cav- ity. Now, in doing all this, there is a variety, a compound character in the motion, which is in plain contrast with the simple motion of the leaves of the catch-fly and the sensitive plant. 30. The distinction, in rela- tion to a digestive cavity, can not be made out in the case of some of the lower animals. And, if it could be, it is not an essen- tial distinction. For it only relates to a mere difference of arrangement in the absorbents that take up the nutritious substance in the two cases. The absorbents in the stomach of the animal, as before remarked, perform the same office that the absorbents do in the root of the plant. They only do it in a different place, and after a different manner. The same remarks, substantially, can be made in regard to the other large central organs which are found in most animals. 31. The last of the distinctions, which I mentioned as being commonly observed, is really the essential distinction between plants and animals. I mean the capacity for sensation and spontaneous motion, which exists only in the animal. There is nothing truly analogous to this in the plant. And we, ac- cordingly, find a peculiar structure in animals, devoted to these functions, and others connected with them. This structure is the nervous system. No trace of such a structure has ever been discovered in any plant. If there were any true analogy between animal motion and the motions of the sensitive plant and the catch-fly, we should be able to find in them traces of ner- vous structure; for the structure of these plants is so plainly developed, that its constituent parts are easily distinguished. 32. The nervous system is evidently not essential to nutri- tion, for this is as well effected in the plant as in the animal. 24 HUMAN PHYSIOLOGY. Thought and will. Instinctive and automatic motions. __________ This is accomplished in both in substantially the same way. The means by which it is done, and its arrangements are modi- fied, as you have seen, in the two cases, to suit the differing circumstances. The nervous system, observe, then, is, for par- ticular purposes, superadded in the animal to what is common both to the animal and the plant, and so constitutes the essen- tial difference between them. And so, all the functions relating to nutrition, which are of course common to plants and animals, are called functions of organic life. But the functions which are performed by the system superadded in the animal, the chief of which are sensation and spontaneous motion, are termed functions of animal life. These are sometimes also called functions of relation, from the especial connection which they form between the animal and all that is around him. 33. These animal functions, sensation and spontaneous mo- tion, imply thought and will. The order of action is this: sensation—thought in regard to it—action of the will in con- sequence of thought—then, from this action, an impression carried through nerves to organs termed muscles—motion in them from their contraction. This order, • however, is not always observed. The first link, sensation, may be absent. Thought, without any preceding sensation, may prompt the will, and spontaneous motion results. The action of the will, too, may be left out, or may be in opposition. Thus, emotions may produce action of the muscles, the will not concurring, and perhaps opposing; as when we laugh at what is ridiculous, or weep at what is sad, in spite of restraining efforts dictated by the will. 34. There are also instinctive motions, and motions which are termed automatic, with which the will has no direct con- nection. And the connection of sensation with them is, in some cases at least, doubtful. The action of the muscles, in swallowing, breathing, &c, and the action of that compound muscle, the heart, are examples of motions more or less dis- connected from the will, and also from sensation. The action of the heart is wholly removed from the direct influence of the will, and it is at least not obvious that it is influenced directly by sensation. It is influenced indirectly by both, through the agency of emotions awakened by them. The muscles of breath- ing, on the other hand, though ordinarily involuntary, may be directly influenced both by the will and by sensation. You can at will breathe faster and more deeply, and sensations of uneasiness in the chest modify the breathing. DISTINCTIONS BETWEEN ANIMALS AND PLANTS. 25 Central organs of nervous system. Most developed in man. 35. For all these different actions, thus produced in different ways, there are central parts of the nervous system upon which the causes of these actions produce the impressions or impulses from which the actions result. Thus, when a sensation is fol- lowed by a spontaneous action of muscles, an impression is conveyed by nerves to the central organ; the will there acts, and the impulse there given by this action of the will is car- ried by other nerves to the muscles, which execute the intended movement. 36. These central parts or organs, which are the media, the instruments of impressions, are in different parts of the body of the animal; but the most important of them is what we call the brain. This part is developed most in those animals that give the greatest evidences of intelligence; and, therefore, it is more prominent in man than in any other animal. 37. It may be remarked, as a general truth, that the nervous system, and its associate or subordinate system, the muscular, are developed in different degrees or forms, to suit the different characters and wants of animals. In man, they are more com- plex and perfect than in any other animal. The brain, in him, is a large organ, occupying the skull. The spinal marrow, and other central parts, and the nerves, are largely developed. And the muscles which are moved by this nervous system form a large portion of the bulk of the body. The organs of nutrition, analogous to those which make up nearly the whole of the plant, occupy the two cavities of the trunk of the body, the thoracic and the abdominal. But, as we descend in the animal kingdom, the nervous system becomes continually less promi- nent, and the system of mere nutrition more so. We at length come to animals, in which the nervous system is a mere small appendage to the system of nutrition, and only serves to direct the muscles in securing the food of the animal. In some of these, we not only do not find a brain, but we fail to discover any traces of a nervous system. This is true of the Hydra, noticed in § 29. 38. The nervous system, which so clearly distinguishes most animals from all plants, is fairly presumed to exist, though in an exceedingly slight degree, in those beings in which it can not be found, but in which we find other characteristics of the animal kingdom. And it is presumed, also, that the exercise of thought and the action of the will, which most animals so plainly exhibit, while they become less and less obvious as we descend in the scale, are not wholly obliterated in the very 3 26 HUMAN PHYSIOLOGY. No nervous system in some animals; yet feeling and thought.___________ lowest animals. It may, perhaps, be said, that as muscular ■action, as mentioned in § 34, is sometimes produced even in man without the intervention of thought or the will, it may be produced in animals of the lowest order altogether in this way. But we may more rationally infer that, as the chief object of motion in them is the securing of food, it is guided by the action of a will in obedience to their sensations. In other words, it is truly a spontaneous, and not a mere automatic mo- tion. And it is probable that there is in the very lowest of animals some degree, though it may indeed be slight, of enjoy- ment in the sensations received from the moving water about it, and from the satisfying of its wants in the process of nutri- tion. We will take the Hydra, a representation of which is given in Fig. 1, page 23, as an illustration of the above remarks. It is a minute gelatinous animal, in which no nervous or mus- cular fibres can be found. And yet it has an extraordinary power of extending and contracting itself. When it is alarmed, it draws in its arms, and shrinks into the form of a little glob- ule ; and if you should see it in this condition, you would not suspect that it had any arms or tentacula. But when it is searching for food, it often extends its body and its arms to a great length; and when it grasps its prey, it puts it into its stomach, which constitutes, so far as we can see, its whole body. We can not conceive of all these motions, thus exe- cuted to effect certain definite objects, without the agency of a will, and without sensations to prompt the will and guide the motions. The animal must have a power of choice, or it would put a bit of stick or straw into its stomach as readily as a worm or an insect. But the tentacula never grasp, among the various bits of things which float against them, any thing be- side the appropriate food of the animal. And it undoubtedly enjoys its food as really, though perhaps not as vividly, as any human epicure; and has in some measure the same pleasur- able sensations which locomotion produces in us, as it floats along so quietly, with its arms hanging down from its body. Though there be no nervous fibres to be seen in the loose gela- tinous structure of this little creature, yet, as the phenomena which it exhibits are known to be produced by the nervous system in those animals whose structure is more plainly and thoroughly developed, we justly infer that there must be nervous matter, in some form, in this and other similar ani- mals. 39. One more important distinction between animals and MAN IN HIS RELATIONS TO NATURE. 27 Peculiar endowments of man. Abstract reasoning. Conscience. plants remains to be noticed. It relates to their chemical com- position. I stated, in § 23, that organized substances are com- posed mostly of four elements—oxygen, hydrogen, nitrogen, and carbon. Plants differ from animals, in having but little nitro- gen in their composition. It was formerly supposed that they contained none of this element. It is found only in particular parts of plants, as the seeds. We may regard carbon as the most characteristic constituent of vegetables, and nitrogen of animals. And in this connection it is interesting to observe that, while carbon is largely thrown off from the lungs of ani- mals, in the shape of carbonic-acid gas, it is as largely absorbed by the leaves of plants. Of this fact I shall take more par- ticular notice when I come to the subject of respiration. CHAPTER III. MAN IN HIS RELATIONS TO THE THREE KINGDOMS OF NATURE. 40. Man is commonly spoken of as being at the head of the animal kingdom, and in the book of the naturalist is made an order of the class termed Mammalia. As the basis of the whole classification is mere material organization, and has no reference at all to mental or spiritual endowments, the classifica- tion, in regard to man, is in its principle correct. At the same time, it must be admitted, that it fails to recognize altogether the essential distinctions between man and other animals. These distinctions, making, as they do, a wide gap—" an im- passable chasm," as Professor Guyot expresses it—between man and the inferior animals, are to be found in certain peculiar spiritual endowments which man possesses. These I will no- tice now in the briefest manner, leaving it for another part of this book to treat more fully of this and other kindred subjects. One of these endowments is the power of abstract reasoning. Other animals in a certain sense reason, that is, they make in- ferences; but they never arrive at any general or abstract truths. Another endowment is a moral one, linking man in his spiritual nature to the Deity. It is conscience, or the knowledge and sense of what is right, in distinction from what is wrong. Other animals, in obedience to the passions of fear and love, 28 HUMAN PHYSIOLOGY. Immortality. Real relation to the animal kingdom. sometimes appear to the superficial observer to have an idea of what is right, as such; but there is not the slightest evidencer that they really have any such knowledge. 41. In view of these endowments of man, it is wrong to consider him merely as being at the head of the animal king- dom. He is something more than this. He is so much and so distinctly more, that the accepted classification of him, on the ground of mere difference of organization, gives a most inadequate idea of his true position in the scale of being. It leaves entirely out of view the essential distinctions; and it separates man from other animals, as you will see, by a distinc- tion of organization which is of rather a trivial, perhaps ques- tionable, character. 42. The force of this view of the subject is enhanced, if we take into consideration that great fact, revealed to us by God in his Word, that man is destined to immortality. It may be objected that, as this fact is learned only by revelation, and not by observation, it is not to be regarded as a scientific fact. But, granting that there is truth in the objection, it certainly is allowable to allude to the revelations of Scripture, as con- firming or enforcing views developed by scientific observation. This is all that I have done in this case. The view which I have presented is based upon endowments that are recognized by the scientific observer, without the aid of revelation; and I appeal to the revealed fact of man's immortality, as adding force to this view, and not as being at all necessary to the establishment of its truth. 43. Let us look at this subject in another point of view. The grand essential distinction between animals and plants lies, as you have seen in the last chapter, in the fact that animals have a nervous system. Now, with this system, as you have also seen, appear certain mental manifestations. These differ widely in different animals, and are most prominent in those in which this system is most prominent and complicated. As we trace upward these complications, when we come to man, we find certain mental manifestations, which separate him by "an impassable chasm" from all other animals. Till we arrive at him, the difference is one of degree, for the most part. But in his case it is a difference of kind, and a very wide one. Of such a difference the naturalist should certainly take very dis- tinct cognizance; and, if it be not consistent for him to do so in his classification, great force and prominence should be given to these views in his instructions on this subject. As the super- MAN IN HIS RELATIONS TO NATURE. 29 The hand of man. No other animal really has such a member. ^adding of the nervous system separates the animal from the plant, so, also, as Professor Guyot very justly maintains, the superadding of such endowments as we find in man separates him, by a chasm quite as " impassable," from other animals. 44. The distinction commonly received as the ground of classification for man, I have said, is a trivial, perhaps a ques- tionable one. He is said to have two hands, and so makes the order Bimana; while apes and monkeys are said to have four hands, and are, therefore, considered as making the order Quadrumana. Now, if we observe carefully and fully the won- derful endowments of the human hand, we shall hardly be willing to allow that monkeys and apes have four such mem- bers. With a full view of the capabilities of the human hand, those members can not be considered as hands, but as members possessing some of the properties of both hands and feet. They are given to these animals to enable them to climb with facility, and to grasp their food; and they have none of that infinite variety of motion, which is so striking a peculiarity of the hand of intelligent man. The ground upon which they are said to have four hands is that which is thus stated by Cuvier. "That which constitutes the hand, properly so called, is the faculty of opposing the thumb to the other fingers, so as to seize upon the most minute objects." No animal besides man has this arrangement, except the Quadrumana. It is claimed, therefore, that they have hands, although they are very imperfect when compared with the hand of man. The imperfection is indeed so great, as to make us at least reluc- tant to admit the claim set up by the naturalist. " While," says Carpenter, " the thumb in the human hand can be brought into exact opposition to the extremities of all the fingers, whe- ther singly or in combination, in those Quadrumana which most nearly approach man, the thumb is so short, and the fingers so much elongated, that their tips can scarcely be brought into opposition, and the thumb and fingers are so weak, that they can never be opposed to each other with any degree of force. Hence, although admirably adapted for clinging round bodies of a certain size, such as the small branches of trees, &c, the ex- tremities of the Quadrumana can never seize any minute object with such precision, nor support large ones with such firmness, as are essential to the dexterous performance of operations for which the hand is admirably adapted." Indeed, what is called the thumb of the Quadrumana is so short and slender, that Eus- tachius, the anatomist, very properly said that, regarded as an 30 HUMAN PHYSIOLOGY. Other peculiarities. Chin. Erect posture. Weeping and laughing. imitation of the thumb of man, it is a ridiculous affair. If, then, we take into view the extensive and varied capabilities ot the human hand, we must agree with Sir Charles Bell, when he says that "we ought to define the hand as belonging exclu- sively to man." This view of the subject has always impressed itself upon the minds of acute observers in all ages. Aristotle said, that man alone possesses hands deserving of the name. Anaxagoras said, that " man is the wisest of animals, because he possesses hands." And the opinion, thus uttered by these philosophers some centuries before the Christian era, is fully echoed at the present time. 45. It would seem, then, that, if mere organization be ad- hered to, as the basis of classification, it is desirable that some ground of distinction in relation to man be fixed upon, which is more definite than the commonly received one. It is to be remembered, however, that, in classification, some one very ob- vious peculiarity that presents itself to the eye is ordinarily made use of as a mark of distinction, while accurate and full discriminations are followed out entirely separately from the mere classification. This is done in the case of man. His structure differs in many respects from that of the inferior ani- mals. It would make this chapter too long to point out all the differences. Some of them are important, while others are not. As an example of the latter, I will mention the fact, that no animal but man has a chin. Every other animal has its lower jaw retreating from the teeth, instead of projecting for- ward below, as in man. One of the most important and strik- ing peculiarities of man's structure is that general arrangement which enables him to be in the erect posture. No other animal naturally assumes this posture, or is able to maintain it for any length of time; and most animals assume one which is en- tirely the opposite of this. Even the monkey, when taught by man to stand and walk, is by no means erect; but his lower limbs are crooked, and the moment that he escapes the neces- sity of being an imitator, he is on all fours. There is a distinc- tion of an interesting character, which concerns both the ner- vous and muscular systems. I refer to the fact, that no animal but man can shed tears, or perform those muscular motions which are necessary to the acts of weeping and laughing. In view of this marked distinction, man has sometimes been desig- nated as " a laughing and crying animal." 46. But the great essential distinctions, to which all the rest are really tributary, are, as I have before stated, of a mental or MAN IN HIS RELATIONS TO NATURE. 31 Tendency to skepticism. Robinet's doctrines. spiritual character. And these should always be made peculi- arly prominent, whenever the distinctions between man and the inferior animals are treated of by the naturalist. This should be done, not only because they are essential, but also because, as I have just hinted, all other distinctions are subor- dinate and tributary to them. It is the mental peculiarities of man, for the most part at least, that render necessary those peculiarities which distinguish his organization from that of other animals. I will not dwell on this point, as I shall speak of it in another part of this book. 47. In view of this whole subject, it may be said, that the classification upon which I have commented is not of itself of very great importance, provided that the definite distinctions, which have been pointed out as existing between man and other animals, be clearly recognized by the naturalist. The tendency, however, evidently is, to lose sight of these distinc- tions in the exclusive regard which is paid to mere material organization. This tendency, it is true, is effectually counter- acted in the case of the great majority of scientific men, by the comprehensive and Christian views which they take of the whole subject; but, still, it manifestly exists, and gives rise to many sceptical notions, especially in superficial and theorizing observers. Great care, then, should be taken to oppose this tendency in all public teaching on this subject, whether it be done by books or lectures. 48. There is a disposition, on the part of some writers, to obliterate the grand distinction between man and the inferior animals, and other distinctions which are stamped by the Cre- ator upon his works. Some go so far as to maintain, that there is not only no line to be drawn between the animal and the vegetable kingdoms, but none even between organized and un- organized substances. Robinet, and many other European authors, teach that all matter has living properties, and _ that every object that we see, whether mineral, vegetable, or animal, is the result of repeated and progressive efforts of nature. The ultimate aim of these efforts is considered to be the formation of man, who is looked upon as the perfection of organization evolved by these efforts. In advocating this theory, they make great use of resemblances and analogies, and even represent the fantastic shapes .which minerals sometimes assume, from their slight resemblance to parts of the human body, " as so many proofs," in the language of Carpenter, " of this long and bungling apprenticeship of nature to man-making." Although 32 HUMAN PHYSIOLOGY. Gradations in nature. Wrong ideas of perfection._______________ such ridiculous doctrines are seldom formally advanced, there is a disposition in many scientific men to indulge in speculations which have more or less resemblance to them. They seem dis- posed to confuse with the veil of mystic scepticism the clear characters which God has imprinted upon the manifestations of his power. It is well, therefore, to fix these characters definitely in the mind, in order to guard against the fascinating and bewildering speculations of a false science. A true science, forsaking the mazes of speculation, and inquiring only for the facts, reads with admiration and reverence the clear lines of God's handiwork, and attributes to no imaginary agency, termed Nature, what bears the marks of exquisite design and Almighty power. 49. An idea, somewhat akin to that of Robinet, is sometimes entertained, viz., that the varieties in the mineral, vegetable, and animal kingdoms are mere gradations in nature. There would be some plausibility in this notion if it were difficult to distinguish the minerals of the most perfect kind from the lowest plant, and then the plants of the highest order from the lowest animal. But the difficulty lies in other quarters. The most per- fect in the three kingdoms are distinguished from each other in the most marked manner; and it is only when the character- istic qualities are the least developed-that there is any difficulty. One kingdom is a no more perfect formation than another. The crystal, with its exact lines and angles—the plant, with its curvilinear and less definite shapes—and the symmetrical animal, are equally perfect in their kind. Each is made for a definite purpose, and is perfectly adapted to that purpose. In none is there any imperfection which could be remedied by endowments taken from another kingdom of nature. In the vast variety of forms which nature presents, there is to be seen no vain struggling after a higher and better state. There is no progressiveness, aiming at an ideal perfection. Neither are there gradations leading to it. All the works of the Creator are per- fectly adapted to the spheres which they fill. They were all from " man, made in His image," down to the humblest ani- mal or plant, pronounced to be "very good," as they came from His hand. 50. Let me not be understood to say that there are no (ni- dations in nature. There are some of a very interesting char- acter ; but they do not obey any such laws as those which are indicated by Robinet and other fanciful theorizers. There are gradations in both the animal and vegetable world. You ob- MAN IN HIS RELATIONS TO NATURE. 33 Man inferior to other animals in some respects. serve them as you go from the simplest plant up to the most complicated. And so of animals. But these two kingdoms of nature are separate in their gradations, and are not in one series together, as is represented by Robinet. And the grada- tion in each kingdom is by no means an unbroken and regular one, going up, step by step, from the lowest to the highest. For example, in the animal kingdom, there are not constant and regular additions made, as you trace the gradations up- ward. And though man stands at the top of the series, it is not as a compound, made up of all the excellencies found below him, with additional excellencies peculiar to himself. Superior as he is, as a whole, to all other animals, yet in some respects he is inferior to many of them. He is inferior to them in the wonderful capabilities of instinct. Some animals can do some things better than he can. The monkey is a better climber. Some animals can do what he can not. Birds and winged insects fly, but he can not. These points could be illustrated to any extent, but this will suffice. 51. Man is often spoken of as being the most perfect of ani- mals. This, as you will see from what was said in a previous paragraph, is not true in the strict sense of the word. His organization is more complicated, and he has more and higher endowments than any other animal; but the perfection of struc- ture, and of adaptation in contrivance to the purposes aimed at, is as manifest in all the varieties of animals as it is in man. 52. In one respect, there is a gradation existing through the three kingdoms of nature. It is in regard to formation or nu- trition. All the elements which are found in the composition of animals exist in the mineral world. But these elements, with very few exceptions, can not be transmitted directly to animals, but they are transmitted indirectly through vegeta- bles. No animal, therefore, can live on mineral substances, although these substances contain all the elements found in its composition. But vegetables draw their nutriment from the mineral world, and then furnish nutriment to animals. There is, therefore, in relation to formation, a gradation running through the three kingdoms, from the mineral up to the ani- mal. 53. At the summit of the last step in this gradation stands man. To him, not only is the animal kingdom tributary, but so, also, are the mineral and vegetable kingdoms. They are all made for him, to beautify and gladden this his temporary home, 34 HUMAN PHYSIOLOGY. All nature tributary to man. Imperfectly so. and to sustain him in it. The subjec tion $*™£^? undoubtedly perfect in his primeval condit<® *J™^™ the garden of Eden. But now, we see this tribute 7 subjecbon manifested only as a general fact, with many exceptions These mark this life as thl imperfect state, and this world as the Smporary home of manfin which he can prepare himself for the perfect and everduring state and home of another life beyond. PART SECOND. CONTAINING, CHAPTER IV.—General Vlyws or Physiology, with a Brief Account of some of the Structures in the Body. CHAPTER V.-Digestion. CHAPTER VI.—Circulation. CHAPTER VII.—Respiration. CHAPTER VIII.—Formation and Repair. CHAPTER IX.—Cell Life. CHAPTER IV. GENERAL VIEWS OP PHYSIOLOGY, WITH A BRIEF ACCOUNT OP SOME OF THE STRUCTURES IN THE BODY. 54. The contents of the previous chapters are preliminary to the consideration of the real subject of this work, the Physi- ology of Man. But they were necessary, in order to accomplish a very prominent object which I have in view. It is my wish that the student, as he examines the functions of the human system, should at the same time observe the analogies existing between man and other living beings, in the processes of life. He will, in this way, get an enlarged view of man in his rela- tions to the world around him, and will be prompted to observe the phenomena of life, in whatever department of nature they may be presented to his view. And, in order to promote this object effectually, I shall, as I proceed with the development of the physiology of man, refer occasionally to the analogous phe- nomena in other animals, and also in plants. This will serve to fix more definitely in the mind of the student the main facts that are to be communicated, at the same time that the bound- aries of his knowledge will be extended over fields full of in- terest. 55. This is a work on Physiology, and not on Anatomy. Physiology treats of the offices or functions of the different parts of the structure, while Anatomy has regard to the struc- ture itself. In the following pages, I shall introduce the anat- omy of the system only so far as it is necessary to elucidate its physiology. Before proceeding to an examination of the individual sub- jects which will claim our attention, it will be proper to present some general views of them in their relations as a whole. 36 HUMAN PHYSIOLOGY.________ Natural division of the subject. Interest of the study. _____________ 56. You have seen, in the preliminary chapters, that organ- ized living beings have much that is common to them all. This is true so far as nutrition is concerned. You have seen that the animal grows very much as the plant does, and that the arrangements for its growth vary from those of the plant only so far as the difference of the source of its nourishment, and of the circumstances under which it is obtained, require. The grand distinction, as you have seen, between animals and vegetables is to be found in the functions belonging to the nervous system. These functions are wholly separate from the nutritive 'functions, which animals perform in common with plants. 57. This view of the subject suggests a natural division of the physiology of man into two parts, viz., the nutritive func- tions, and the animal functions, or those connected with the nervous system. In other words, the first division will com- prise all those subjects which relate to the building and repair- ing of the human structure ; and the second will comprise those which relate to the uses for which the structure is made. The first class of subjects includes digestion, circulation, respiration, formation, and excretion. The second class includes locomotion, sensation, the five senses, the voice, instinct, thought, &c. 58. The student will see at one glance, that a wide range of exceedingly interesting subjects opens before him. Contem- plated as a mere mechanism, the human system is full of won- ders. The principles of common Mechanics, of Hydraulics, of Pneumatics, of Optics, of Acoustics, are abundantly illustrated in the human body, by contrivances of the most exact and ex- quisite adaptation. But this congeries of beautiful mechanisms is all regulated by a nervous system, making it, by its minute fibrils, to be alive with feeling in every part. Sensation and sympathy govern, through the nerves, in a wonderful manner, the ever-varying adjustments of all the parts of the complicated system. It is not only mechanism, but living mechanism, that develops to us its wonders, so numerous and diversified. And then, when we look at the soul—"that side of our nature which is in relation with the Infinite"—connected as it is by the nerves with every part of this mechanism, the interest of the study before us appears exceedingly great, and its variety never ending. The study is a peculiar one. It is not the body merely that you are to study in these pages; but it is the body and spirit united. The study differs from all others in this respect. In other studies, you look at either matter alone, or GENEEAL VIEWS. 37 ______________Bones. Two parts, animal and mineral. Cartilages. spirit alone; but here you look at them both, as brought to- gether in mysterious union. 59. It will be proper, here, to say something in general of the structure of the human frame, before proceeding to a par- ticular view of individual subjects. I do this in order to avoid a frequent turning aside for explanation, which would not only be inconvenient, but would mar the interest of the study. It will not be necessary to go into a full description of the nume- rous and diverse textures, or tissues, (as they are commonly called,) of the body. I will notice only some of the principal of them. J * F 60. From the osseous or bony tissue, the solid part of the framework of the body is made. Bone is composed in part of animal matter, and in part of mineral. The mineral part is mostly phosphate of lime. These two parts of bone are in different proportions to each other in the different periods of life. The mineral part in the child is about one- half of the bone; in the adult, four-fifths; and in the old, seven-eighths. The bones are therefore very brittle in old age, while they are somewhat yielding in childhood. The mineral and the animal portions of bone can be separated from each other. If a bone be put into diluted muriatic acid, the mineral part will after a time become united with the acid, and the animal part will be left, having the perfect shape of the bone. Thus separated from the mineral part, it is so flexible, that it can be tied into a knot without affecting its shape. On the other hand, by subjecting a bone for some time to the action of heat, the animal part can be removed, and the mineral part be left by itself. 61. The animal part of bone is cartilage, or gristle. This part is formed first, constituting a sort of mould, in which the bone is to be formed. The mineral matter is gradually depos- ited in the cells of the cartilage. In the very young child, you can see that this process is not completed, especially if you ob- serve the bones of the head. The bones are not united to- gether, as they are in the adult; and there is so little of mineral matter near their edges, that they can be bent with a very slight pressure. The proportion of mineral matter which is deposited in the cartilaginous bones varies much in different animals. In many fishes, there is almost none of this deposit, the skeleton retaining its cartilaginous character throughout life. 62. Besides the cartilaginous portion of bones, there are car- 4 38 HUMAN PHYSIOLOGY. Ligaments. Muscles. Tendons. Cellular tissue.________________ tilages which are destined to remain so, instead of having mineral deposits made in their cells. The ends of the bones are tipped with them. They are placed between all the twenty- four bones of the spinal column. They form the connecting links between the breastbone and the ribs. Cartilage consti- tutes the body of the outer ear, of the eyelids, and of the lower part of the nose. The transparent part of the eye is formed of cartilage. This substance is placed wherever firm- ness and tenacity are required without hardness. 63. The bones are united together by ligaments of various degrees of strength, according to the necessity of the case. They are moved by muscles, which, in man, are bundles of reddish fibres. Muscular substance is what is commonly called the meat in animals. It is of various colors in different ani- mals, or in the same animal at different periods of life. All motion in animals is produced by muscles. I will not go into an explanation of their action now, any further than to say, that they act by contracting or shortening their fibres. Com- monly there are tendons united with the muscles. These ten- dons are composed of strong white fibres, and have no power of contraction themselves. They serve merely as the cords by which the contracting muscles move the bones and other parts. 64. The most common tissue in the body is what is called by the names, cellular membrane, cellular tissue, and areolar tissue. This last name is most commonly used by physiologists at the present time. The word areolar comes from the Latin word areola, meaning a small open space. The term is appro- priate, because this tissue is filled with minute spaces or cells. The word cellular is quite as appropriate; and, as this will be more familiar to you, I shall make use of it whenever I shall have occasion to speak of this tissue. That you may under- stand what this tissue is, I will refer you to its appearance as you see it in different meats. It is the delicate white substance that you see between the different layers of muscle in a piece of meat. If you notice particularly, you will see that it is also between all the different fibres of the muscles. As the spaces or cells communicate together, butchers sometimes " blow up" this tissue in veal, in order to make the meat look more plump. This tissue is the universal packing material of the body. It is to be found almost everywhere. It surrounds every thing,—vessels, nerves, muscles, organs, &c. It enters into their composition, uniting together different tissues, and GENERAL VIEWS. 39 FIG. 2. Communication between cells of cellular tissue. Shown in dropsy. also the fibres of the same tissue. It varies much in its com- pactness in different parts. It is very fine and compact where it is necessary that it should be so ; while in other cases it is loose and delicate, allowing a free motion of the parts which it envelops and connects together. It is abundant and loose among the muscles, and between them and the skin. Fig. 2 represents a portion of cellu- lar tissue, inflated and dried, exhibiting the arrangement of its larger meshes. This is magnified twenty diame- ters. The free communica- tion which exists between the interstices or cells in this tissue is exemplified in drop- sy. These cells are bathed, in the healthy state, with a small amount of a watery fluid; and when this in- creases largely, forming the disease termed dropsy, it obeys the force of gravity in the cells, and accumulates most in the lowest parts of the lower extremities. This tissue is very elastic in health, so that if you press on the skin, there is no indentation left when the pressure is taken away, for the elastic cellular tissue at once rises from its state of compression, pushing the skin before it. But in dropsy it loses its elasticity by over distension; therefore there is pitting, as it is termed, after removing the pressure. We sometimes have an opportunity of seeing the communication between the cells manifested by the introduction of air into them. This has occurred in some cases in which an opening has been made, by disease or accident, from the air tubes in the lungs into the cel- lular tissue in the walls of the chest. The whole body has been seen largely swollen, from the air which has from this cause accumulated in this tissue directly under the skin. Among the many tricks of impostors, the inflation of the cel- lular tissue of the head has been practised ; and as it produces a frightful appearance, and therefore excites pity, the trick is a very successful one. CELLULAR TISSUE. 40 HUMAN PHYSIOLOGY. Deposits of fat. Its uses. How kept in its cells. Mucous tissue.__________ 65. There are here and there in the cellular tissue deposits of fat. Various purposes are answered by these deposits. They are sometimes of use in promoting a free motion of the adja- cent parts. The eye has, intervening between it and the bony socket, a cushion of fat, on which it rests, and against which it is pressed when any violence is offered to it. Fat, as a non- conductor, is a protection against the cold, and it is therefore deposited largely in the cellular tissue under the skin, in ani- mals that inhabit cold climates. Fat, also, sometimes serves as nourishment to the system when its necessities require it. In diseases in which food cannot be taken in any amount, the fat is absorbed into the system. The heat of the body is maintained also, in part, by this process. This occurs in the torpid condition of hybernating animals. They commonly be- • come very fat in autumn, as a preparation for the winter, and in the spring they come forth very lean, their store of fat having been used up for the purposes of nutrition and heat during their confinement. The fat thus deposited in the cellular mem- brane or tissue is not diffused merely in the interstices, but it is confined in cells of its own, which lie in these interstices. Mi- nute blood-vessels, pass from the fibres of the cellular tissue to these fat cells. The fat, which is an oily fluid, is kept from oozing through the pores of the cells that hold it by the blood, which is very nearly four-fifths water, and by the watery fluid which I have spoken of as bathing all the interstices of the cellular tissue; for oil, you know, will not pass through any porous substance that is wet with any watery fluid. If a por- tion of cellular membrane containing fat be dried, the fat, which in the moist state is wholly confined to its cells, now oozes through their pores. This is the reason that a lump of fat, as it is called, feels so oily after it has been exposed for a while to the air. 66. The mucous tissue is that which lines all the cavities of the body that have outlets. It lines the mouth and the cavities of the nose, and descends into the lungs, the stomach, &c. It takes its name from the fluid called mucus, which is constantly secreted by innumerable minute glands, that are situated in the substance of this membrane. The chief object of this viscid fluid is to protect the membrane from the substances which come in contact with it, which would otherwise produce some irritation. This membrane is continuous with the skin, shading off into it insensibly, as you may observe on the lips. 67. The serous tissue or membrane forms the outer coat or GENERAL VIEWS. 41 __________ Serous membranes. Compound character of the organs. lining of nearly all those organs the inner coat of which is mu- cous membrane. This is the case with the lungs, the stomach, and the intestines. The serous membranes are white, smooth, and shining, and are lubricated with a watery fluid, called serum. Every serous membrane forms a cavity or sac without an outlet, differing in this respect entirely from the mucous membranes. Thus, in the case of the lungs, the serous mem- brane lining the outside of each of these organs passes from the lungs to the walls of the chest, lining the inside of them, and thus makes a sac without an outlet for each lung. This sac could be dissected off, and taken out whole. When the fluid which lubricates the inside of this sac increases to any extent, the disease called dropsy in the chest is produced. The membrane which thus lines the outside of the lungs and the inside of the walls of the chest is called the pleura, and it is the seat of the disease termed pleurisy. 68. I have thus described some of the principal of the tis- sues which make up the human structure. The other tissues will be spoken of in the proper connection as we proceed. Be- fore dismissing this subject, I will call your attention to the fact, that the organs of the body are generally composed of several tissues united together. Thus, the stomach has three coats, as they are termed,—the mucous as the inner coat, the serous as the outer, and the muscular between them. And then we have the cellular tissue uniting these together. Besides these, there are arteries, and veins, and nerves, so that the stomach, which looks like a simple pouch, is really quite a composite thing. And the same can be said of the other organs. 69. Before entering upon the particular consideration of the functions by which the system is built up and kept in repair, it will be well to take a general view of them, that you may see them in their connection and mutual dependence. Each of these functions has its special and appropriate part to play, in effecting the formation and repair of the structure. The material from which all parts of the body are formed and repaired is the blood. There are organs whose special duty it is to make this material; organs which distribute it; and or- gans which use it after it is distributed. There are also organs whose duty it is to receive the blood after it has been used, and fit it to be used again. This common building material of the body is made out of the food; and the succession of processes by which it is done I will describe in the next chapter. After it is made, it is distributed by the complicated apparatus of the 42 HUMAN PHYSIOLOGY. Summary of nutritive functions. Processes of digestion._____________ circulation. This apparatus is therefore the common carrier of the building material of the system. It is the numberless little form- ative vessels, so small as to be invisible to the naked eye, that use the blood thus brought to them, and make and keep in repair all the various structures that we see in the body. When the blood has been used by these formative vessels, it is not fit to be used again until it is submitted to a purifying process by exposure to air; and to this particular object the lungs are de- voted. And besides all this, as there are continually some par- ticles which, in the wear and tear of the machinery, become useless, and even injurious, they must be got rid of in some way; and so there are organs for this purpose—organs of waste, as they are termed. It is also to be remembered, that the processes to which I have now alluded are so carried on, that the heat of the body, as will be fully explained hereafter, is steadily maintained. In the following chapters of this part, I proceed to a particular examination of the functions of which I have now given a brief summary. CHAPTER V. DIGESTION 70. I shall include, under the term digestion, all those pro- cesses which are necessary to effect the separation from the food of its nutritious portion, and the introduction of it into the circulation. A summary of these processes may be thus given. The food is broken up and ground in the mouth, and it is at the same time mixed with the saliva. It is then taken into the stomach, where it is kept in constant motion, and is under the solvent action of a fluid of a peculiar character. When it is brought into the right condition, it is passed from the stomach into the intestines. Here it is acted upon by two fluids, the bile, the secretion of the liver, and the secretion of another gland, the pancreas or sweet-bread. These secretions have some agency in separating from the mass its nutritious portion, and this is taken up or absorbed, in the form of a milky fluid, by little vessels lying on the surface of the inner membrane of the intestine. These vessels unite together to form a large tube, and through this the milky fluid is poured into the circulation, to replenish the blood. DIGESTION. 43 Mastication. Teeth various, according to different kinds of food. Having given this summary of the processes which make up digestion, I proceed to speak of them more particularly in the order of their succession. In doing so, I shall notice some of the points in which other animals differ from man, in regard to these processes and the arrangements of the apparatus of diges- tion. 71. Mastication is an important part in the process of diges- tion. The teeth, which perform this act, are very hard bodies. The body of a tooth is composed of two substances. The in- ner part is called the ivory, and the outer is called the enamel, which is exceedingly hard. The teeth are of different shapes for different modes of action. There are long and pointed teeth, for tearing; others, for cutting, which have a sharp edge; and others, for grinding, having for this purpose a broad and irregular surface. The teeth are differently shaped in animals, according to the kinds of food which they eat. Thus, the her- bivorous, or vegetable-eating animals, have grinding teeth to bruise their food; while the carnivorous, or flesh-eating animals, have sharp-edged teeth and long-pointed teeth, by which their food is torn and cut in pieces. And it is to be observed, that the movement of the jaws always corresponds with the char- acter of the teeth. In the carnivorous animals, the motion of the lower jaw upon the upper is a mere up-and-down, or hinge- like motion. As they have no grinding teeth, there is no need of any lateral or grinding motion. But in the animals that have grinding teeth, there is a lateral motion, to enable them to grind. You see this difference very plainly, if you ob- serve the dog and the horse while they are eating. In Fig. 3, you see represented the teeth of a carnivorous animal. The front teeth are long and pointed, for rending, while the back FtG. 3. TEETH OF CARNIVOROUS ANIMAL. 44 HUMAN PHYSIOLOGY. Man an omnivorous animal._________________ teeth have a sharp edge for cutting. In Fig 4, you see repre- sented the broad and irregular grinding surfaces of the teeth ot herbivorous animals. In animals that live on insects, the teeth present conical points, which press into corresponding depres- sions in the opposite jaw, as represented in Fig. 5. In those that live on soft fruits, the teeth are rounded, as in Fig. 6. These are quite in contrast with the tearing teeth of the carniv- orous, and the grinding teeth of the herbivorous. fig 5. FIG- 6- TEETH OF FRUGIVOROUS ANIMAL. TEETH OF INSECTIVOROUS ANIMAL. 72. There is an arrangement of the enamel and the ivory in the teeth of the herbivorous which ought not to pass unnoticed. Instead of having the enamel cover the ivory, as in the teeth of the carnivorous, the two substances are arranged in upright layers, as seen in Fig. 4. The object of this is plain. The ivory wears away faster than the harder enamel, and, therefore, the surface of the tooth always presents projecting hard ridges, fitting them for grinding thoroughly. A miller would say, that these are stones that never need picking. 73. So perfect is the correspondence of the teeth with the kind of food on which the animal lives, that the skillful natural- ist can infer very correctly, from the examination of the teeth of an animal alone, the character of the food on which it lives, and even the general arrangement of its structure. As man has the three kinds of teeth which I have noticed, he is said to be omnivorous, or an eater of all kinds of food. In him, the front teeth are the cutting ones; what are called the stomach and eye teeth are the tearing ones; and the large back teeth are shaped for grinding. It will be observed that the tearing teeth, as they have not a very sharp point, and are no longer than the other teeth, have but little power when compared with the long and sharp tearing teeth of a carnivorous animal, as seen in Fig. 3. As man can make use of instruments to cut DIGESTION. 45 Whales have no teeth. Substitute for teeth in birds. his food in pieces, he does not need such power in his teeth as carnivorous animals have. Allowance should be made for this in estimating the amount of carnivorous adaptation in man. 7-4. There are a few of the Mammalia that have no teeth. This is the case with the common whale. In his case, instead of teeth, there hang down from the upper jaw, as represented in Fig. 7, plates of a fibrous substance, called whalebone, which have their fibres separated at their free extremities, so as to make a sort of sieve. This is intended to catch the little gela- tinous animals, which the whale devours in great numbers. FIG. 8. FIG. 7. WHALEBONE. SKULL OF WHALE. For this purpose, he draws in the water, making it to pass through this sieve, and then expels it from the nostrils or blow- holes. Birds, too, have no teeth. Their place is supplied by a contrivance in the stomach itself, for the breaking up of the food. This will be described in another part of this chapter. 75. While the food is cut and ground by the teeth, it is at the same time thoroughly moistened by the saliva, which is poured forth from certain glands in the neighborhood. There are three pairs of these glands. Fig. 9 shows the glands on one side. The parotid gland, 1, is the largest. This is situated in front of the lower part of the ear. It is the seat of the swelling in the disease called mumps. Its duct, 2, passes over one large muscle and between the fibres of another, and pours its contents into the mouth opposite the second small grinder of the upper jaw. If you press on this part of the cheek, you can 46 HUMAN PHYSIOLOGY. Formation of the saliva. Three pairs of salivary glands. FIG. 9. SALIVARY GLANDS. feel in the mouth an increased flow of the saliva. The sub- maxillary gland, 3, is situated inside of the lower jaw at its lower part; and its duct, 4, opens into the mouth at the side of the frsenum of the tongue. The sublingual gland, 5, lies under the tongue, and discharges its secretion by a duct at the side of that organ. These saliva factories, as we may term them, are in much more active operation at some times than at others. They are especially active when we are eating ; and it is commonly estimated that, during an ordinary meal, about eight ounces of saliva are poured into the mouth. This large amount is wanted to moisten the food thoroughly before it is swallowed; and it is supposed, also, that it has some chemical influence in preparing the food for the action of the gastric juice in the stomach. More saliva than usual is needed, also, when we are speaking, in order to keep the parts properly lubri- cated, for the passage of the air in and out during speaking dries up the saliva by evaporation. And, accordingly, the mo- tion of the parts at such times stimulates a larger flow, just as pressure on the cheek will do it, as before remarked. This result is favored by the arrangement of the duct of the parotid gland, which, as you have seen, passes over one large muscle, and then through the body of another. Chewing any thing excites DIGESTION. 47 Flow of saliva affected by sympathy. Swallowing. an increased flow of the saliva; and the tobacco chewer does a real injury to the salivary glands, by keeping them constantly in excessive operation, in addition to the ruinous effects of this drug on the system at large. When he eats, these over-worked factories can not turn out as good an article as they should, nor will it be in sufficient quantity. 76. It is supposed that, besides the mere mechanical stimu- lus of the motion of the parts, the stimulus of sympathy is also concerned in exciting these glands to increased action. The glands are supposed to be affected in this way by the stimula- tion of the food on the surface of the mouth, about the orifices of their ducts. That sympathy does have an influence on their secretion is evident from the very familiar fact, that the thought of food will often, as it is expressed, cause the mouth to water. 77. The fact, that these glands do not all secrete the same kind of fluid, has led to an interesting discovery in relation to them. The submaxillary glands secrete rather a viscid fluid, while that which is poured forth by the parotid and sublingual glands is perfectly limpid. Now, it has been found, by various observations and experiments on animals, that while the teeth are cutting and grinding the food, and the parotid and sub- lingual glands are pouring out the saliva to moisten it, no secre- tion comes from the submaxillary glands. But these glands pour out their viscid fluid the moment that the tongue thrusts the food back towards the throat, in the beginning of the act of swallowing. The special object of this viscid fluid is then to cover the food, so that it may, to use a common expression, slip down easily into the stomach ; and it has nothing to do with the moistening of the food, this being the particular office of the other two pairs of glands. 78. When the food has been ground by the teeth, and moistened by the saliva, it is carried by the act of swallowing into the stomach. This act, simple as it appears to you, is a very complicated one, and is performed by the conjoined and agreeing action of many different muscles. The food is first thrust back over the surface of the tongue into the large cavity in the back of the throat, called the pharynx. In the pharynx are the openings of two tubes—the oesophagus or gullet, which is the passage into the stomach, and the trachea* or windpipe, the passage to the lungs. As the oesophagus lies behind the * This term is sometimes used, as here, to mean the whole of the tube conducting to the lungs, including the larynx, which is at the top of this tube, and sometimes it is restricted to that part of the tube which is below the larynx. 48 HUMAN PHYSIOLOGY. Parts employed in swallowing. Office of the epiglottis. trachea, the food, in passing to it, must go directly over the opening into the trachea. To prevent the food from entering the trachea, therefore, there is a little tongue-shaped body, called the epiglottis, extending back from the root of the tongue, and acting as a lid to the glottis, the opening into the trachea. When we are swallowing, this lid is shut down; but it is always raised up when we are breathing or speaking. When we swal- low, not only does the lid shut down, but the larynx rises to meet the lid, as you may readily perceive, if you place your fingers upon what is called Adam's apple while you are swal- lowing. With the aid of Figures 10 and 11, all this will be very clear to you. In Fig. 10, you have a side view of the parts engaged in swallowing, as if the head were divided into two halves by a vertical section. At i, is the cavity of the nos- tril ; at h, are the lips; a is the divided bone of the chin; b is the tongue, between which and the spinal column, /, is the large cavity of the pharynx. In front of this cavity hangs the FIG. 10. VERTICAL SECTION OF THE THROAT. palate, g, as a door or valve, to direct the air coming from the trachea, d, either through the mouth or throuo-h the nostrils according to its position. The oesophagus, e, is behind the tra- chea, and the epiglottis, c, shuts down when we swallow to let DIGESTION. 49 Mode of action of the oesophagus in swallowing. the food pass over it into the oesophagus. In Fig. 11, you have a view of the same parts from the rear. At 1, is a sec- tion of the bones at the base of the skull; 3, 3, are the cavities of the nostrils; 2, 2, the walls of the pharynx spread apart; 5, the pendulous palate; 6, 6, the arch of the palate; 8, the root of the tongue; 9, the epiglottis, and 10, the glottis, or opening into the larynx; 13, the oeso- phagus; 14, the trachea. The pharynx, you see, is a funnel- shaped cavity, tapering down to the oesophagus, the opening of which is considerably below the opening of the trachea. 79. When the food enters the oesophagus, it is carried through that tube into the stomach by the ac- tion of muscular fibres. These fibres are represented in Fig. 12. The cir- cular fibres are seen at a and b. These are removed at c, so as to show the longitudinal fibres. It is by the con- sent of action between these different sets of fibres that the food is propelled through the oesophagus. As the food descends, a dilatation of the circular fibres must everywhere take place where the food is, and a contraction of them immediately behind it—the dilatation making the way for it, and the contraction forcing it along. And in animals that chew the cud, these actions must be reversed when the ball of food is forced up through the oesoph- agus into the mouth. 80. The food being introduced into the stomach, is here subjected to the action of the gastric juice. This is a 5 FIG. 11. VIEW OF THE THROAT FROM BEHIND. FIG. 12. CESOPHAGUS LAID OPEN. 50 HUMAN PHYSIOLOGY.________ Gastric juice. Chemical in its action. peculiar fluid, somewhat acid in its character, which is secreted by very minute follicles, or bag-like cavities, situated in the sub- stance of the mucous membrane. Ordinarily there is none of this fluid in the stomach when there is no food there. _ Dr. Beaumont made some very interesting observations on this, as well as many other points, in the remarkable case which fell under his care. The individual, Alexis St. Martin, received a wound in his left side by the bursting of a gun. The wound, which opened into the stomach, never entirely closed, but an orifice remained, after the healing process had done all that it could. Through this orifice, Dr. Beaumont could look into the stomach, and observe Avhat was going on there. He describes the mucous membrane, in its healthy state, as having a velvet- like appearance, with a pale pink color, and as being covered with a very thin, transparent, viscid mucus. On introducing some food, or irritating the mucous membrane mechanically, he saw, by the aid of a magnifying glass, "innumerable lucid points " projecting on the surface, and from these there exuded a pure, limpid, colorless fluid. These points were the follicles which secrete the gastric juice, now rendered turgescent by ► being stimulated to action. 81. The amount of gastric juice secreted is generally about in proportion to the amount of food which the necessities of the system require. When the quantity of food taken is very much more than is required, there can not be a sufficient amount of gastric juice secreted to digest all of the food ; and some of the food must therefore remain undigested, and will prove a source of irritation to the stomach. If the amount of food taken from day to day is not very excessive, but is only a little above the proper quantity, there will be enough of the gastric juice made to digest it; but the daily overtaxing of the pow- ers of the secreting follicles will, after a while, produce derange- ment in the stomach, and perhaps permanent disease. 82. The action of the gastric juice upon the food is of a chemical nature. In order that it may act effectually on all portions of the contents of the stomach, this organ is" kept in constant motion by the fibres of its muscular coat. These fibres are so arranged that, as they contract and relax, they keep up a sort of churning of the contents, and thus effect a thorough mixture of them with the gastric juice. In Fig. 13, you see these fibres represented. At 1, is the opening of the oesopha- gus into the stomach ; and at 4, is the part which opens into the intestine. The fibres are in different layers, running in DIGESTION. 51 Churning of the food in the stomach. The chyme. FIG. 13. MUSCULAR FIBRES OF THE STOMACH. different directions. The outer peritoneal coat, 5, 5, is dissected off and turned back, showing some of the fibres that run length- wise of the organ, 6 ; and some of them transverse, 7 ; and others, 8, that run obliquely. You can readily see what effect the contraction of these different fibres will have on the shape of the stomach. The contraction of the longitudinal fibres, 6, brings the large, bulging end of the stomach, 2, and the small end, 3, nearer together. The transverse fibres, when they con- tract, diminish the capacity of the stomach transversely. And the oblique fibres modify these two motions by their oblique action. 83. By the combined chemical and mechanical action of the stomach, its contents are, after a little time—in three or four hours—reduced to an uniform, greyish, semi-fluid mass, called chyme. While this process has been going on, the communi- cation between the stomach and the intestines has been entirely closed by a valve, called the pylorus. This is represented at 5, in Fig. 14, which presents a view of the inside of the stomach. This valve is made of a fold of both the mucous and muscular coats of the stomach. It is a very faithful sentinel, as is indi- cated by its name, which is derived from two Greek words, sig- nifying to guard the gate. It will not ordinarily permit any undigested food to pass it. While the process of digestion is 52 HUMAN PHYSIOLOGY. The pylorus. A sentinel. On duty only during digestion. FIG. 14. INTERIOR OF THE STOMACH AND SMALL INTESTINE. going on, the motions produced by the muscular fibres cause the contents to move about, and of course they are thrown against the pylorus, as well as any other part of the stomach. But the valve remains closed, until some portion comes against it that is thoroughly changed to chyme, and is therefore fit to pass on into the intestine. It then opens to let this pass, and it does so for any other portions that have become chyme. Toward the conclusion of the digestion of a meal, small quan- tities pass at first, and after a while, the contents pass quite rapidly through the valve. 84. Although this sentinel-valve thus performs its duty so faithfully in relation to nutritive substances, it seems to let other substances pass very readily. Solid substances, swallowed by mistake, as buttons, pieces of money, the pits and skins of vari- ous fruits, often pass the valve without any trouble. The valve seems to be on duty as a sentinel only during the process of digestion; and, if the attempt to go through with this process prove unavailing, the pylorus, though it let such hard sub- stances as I have mentioned pass without difficulty, resists the passage of the undigested food, sometimes causing much un- easiness, and even perhaps pain, by so doing. In such a case, either the valve after a time gives over its resistance, or, hold- DIGESTION. 53 Theories of digestion. Eating between meals. Eating fast. ing out, the action of the stomach is reversed, and the offending matter is thrown off by vomiting. 85. It is not a little amusing to read the different theories which were formerly broached to explain the process of diges- tion. Some supposed it to be a concoction, heat being, in their view, the chief agent; some, a kind of putrefaction ; some, a chemical solution; some, a trituration; and some, a process dependent upon the action of the nerves. Of these various theories, the celebrated Hunter playfully remarked: " To ac- count for digestion, some have made the stomach a mill; some would have it to be a stewing-pot, and some, a brewing-trough ; yet, all the while, one would have thought that it must have been very evident that the stomach was neither a mill, nor a stewing-pot, nor a brewing-trough, nor any thing but a sto- mach.^ All these theories are now done with; and it is pretty well ascertained, that digestion is a chemical process—in part a solution, and in part a fermentation—and that mechanical agency is employed only for the purpose of thoroughly exposing the food to the action of the gastric juice. 86. The process of digestion, as it has been described, is a regular process, requiring a certain average period of time for its completion. If, during the progress of it, fresh food be in- troduced, its regularity is broken in upon, and the process fails to be well done. Then, too, if, immediately after the completion of the process, a new supply of food be taken, harm is done, because the organ has not its needed interval of rest. For these reasons, the practice of eating between meals is a very injurious one. Eating fast does harm, because,—1st, the food is not sufficiently ground ; 2d, it is not mixed thoroughly with the saliva; and, 3d, more food is taken than would be sufficient to satisfy the hunger if the individual ate slowly, and, therefore, more than can be easily digested. Great variety in food stimu- lates the appetite unduly, and too much is consequently eaten. Exercise' facilitates digestion, if it be not violent. An experi- ment was once tried upon two dogs, which was thought to prove that exercise hindered digestion. Two dogs were fed freely, and while one was left to lie still, the other was made to run about violently. Both dogs were killed after an hour or two, and it was found that, while digestion had gone on thor- oughly in the dog that was allowed to remain quiet, in the other the food was undigested. This only proved that violent exercise, taken immediately after eating, impedes digestion. It has been found, on the other hand, that light exercise pro- 54 HUMAN PHYSIOLOGY. Cause of hunger, state of the system. Its seat in the stomach.___________ motes the process; and daily experience, among laborers, shows, that very strong exercise does not interfere with it, if a little interval of rest be allowed, so that the process may be fairly begun. 87. The sensation of hunger has been attributed to various causes,—as the empty state of the stomach, the presence of the gastric juice irritating the mucous membrane, &c. It cannot arise from emptiness; for, if it were so, it should occur sooner than it does after eating, and it should not be absent in dis- ease, as it often is for a long time, when the stomach is, almost entirely empty. It can not arise from the irritation of the gas- tric juice ; for it was found by Dr. Beaumont, in his observa- tions of the stomach of Alexis St. Martin, that this fluid is not secreted till after food is introduced into the stomach. The cause of hunger is evidently in the state of the system. It is a state of want. Nutriment is needed by the formative vessels, the builders and repairers of the system, of which I shall speak particularly in the chapter on Formation and Repair. And they make their wants known as distinctly as the bricklayer does, when he calls for more brick and mortar. Through the nerves, an impression is communicated from these to the sto- mach, and the sensation of hunger is the result. That the sen- sation is seated there is evident from the fact, that it can be temporarily relieved by putting indigestible substances into the stomach. These produce the effect by causing other sensations there, which take the place, for the time being, of the sensation of hunger. After, however, the momentary effect is over, the sensation of hunger returns again in its full force. The cause, then, of the sensation is in the system at large, but its seat is in the stomach. Its degree of urgency depends upon this general state that causes it. If eating be delayed much beyond its usual time, or if the system has been exhausted by the wear and tear of severe labor, the sensation of hunger is very urgent. So, too, if disease has impoverished the system, as soon as the stomach is in a condition to respond to the call of the forma- tive vessels that set themselves to work to repair the waste, the hunger is often excessive. Observe, here, that in order to have the sensation of hunger, not only must there be a want in the system at large, but the stomach must be in a state fitted to receive the notice of this want. And fortunately it is seldom in this state except when in a condition to do its work. If it were otherwise, food would often be introduced into it when it could not be digested. The stomach is sometimes incapacitated DIGESTION. 55 Sensation of hunger affected by the mind. Thirst. for receiving the notice of the want of the system by mental impressions. In this case, an impression is communicated from the brain to the stomach, through the nerves, which counteracts the impression conveyed from the system to this organ, and so neutralizes the sensation of hunger. Grief thus often destroys the appetite for food. One thing more is to be observed in relation to hunger. Although this sensation is caused by the want of the system, it is removed long before the nutriment reaches its final destination, and supplies the want. How is this ? It is either because the new sensations produced in the stomach, by the commencing process of digestion, take the place of the sensation of hunger, or an impression is sent all over the system from the filled stomach, which, so to speak, stills the clamor of want with the immediate prospect of a supply. 88. Nearly the same remarks can be made in relation to thirst, that have been made in regard to hunger. The seat of this sensation is in the fauces or throat. Its cause is evidently not there; for the mouth and throat may be very dry, and yet there may be little or no thirst; while, on the other band, there may be much thirst, although the mouth and throat are moist. The cause of the sensation is like the cause of hunger, in the system, at large; and, therefore, no local cause, producing a dryness of the throat, can cause thirst independent of a general condition. 89. Before describing the remainder of the process of diges- tion, I will call your attention to the arrangement of the sto- mach, and the other organs of the abdomen engaged in this process. Fig. 15 exhibits them as they present themselves in a front view, except that they are somewhat separated from each other, instead of being as closely packed, as they are in the abdomen. The large end of the stomach, you see, lies to the left side,* and at this end is the spleen. The pancreas is be- hind the right end of the stomach. Above the stomach, and mostly to the right side, is the largest organ in the abdomen, the liver. It is represented as turned upward in the Figure. The stomach is directly connected with the small intestines at the pylorus. At the end of this long and winding tract begin the large intestines. The duct of the gall bladder, and that of the pancreas, empty their contents into the small intestine at its beginning. The office of the spleen has not yet been ascer- tained. Neither has that of the worm-like appendage at the * As this is a front view, the right side of the Figure is the left side of the body. 56 HUMAN PHYSIOLOGY. Arrangement of the digestive organs. FIG. 15. GALL BLADDER LARGE INTES TINES, BEGINNING OF LARGE IN- TESTINES. WORM-LIKE AP- PENDAGE. LARGE INTES- TINES. SMALL INTES- TINES. SMALL INTESTINES. DIGESTIVE ORGANS. beginning of the large intestines. The omentum, or caul, which hangs like a curtain from the front part of the sto- mach down in front of the intestines, is not represented in the Figure. 90. There is one arrangement in the abdomen which must not pass unnoticed. If the intestines were left to lie loose in this cavity, they would constantly be subject to displacement and injury. They are therefore fastened to the backbone by an arrangement, which secures them from any such accident, and at the same time allows of a sufficiently free motion of differ- DIGESTION. 57 The arrangement of the mesentery. Its offices. ent parts of this tube. It is this. The intestinal tube makes the margin of a broad sheet of membrane, the other edge of which is gathered up and fastened to the spinal column. The arrangement is like a ruffle with a puffed edging. The mem- branous sheet is called the mesentery. As the intestinal tube, the puffed edging, is much longer than the ruffle itself, the mesentery, it is gathered on to the ruffle, as a seamstress would express it. Now, the mesentery is composed of two folds of the peritoneum, the smooth, shining, outer covering of the in- testines. The arrangement will be easily understood by the diagram in Fig. 16, which represents a section of the intestine with the mesentery. The cav- ity of the intestine, a, is lined fig. 16. by the mucous membrane re- presented by the inner circle. Next comes the muscular coat, and next the peritoneal, the outer, which, instead of making a circular tube, as the other two coats do, passes backward on both sides of the intestine, to make the mesentery, b. After being attached to the spine by PLAN 0F THE mesentery. means of cellular tissue, it is re- flected off to pass over other portions of the intestine, as seen at c, c. Between the two layers of the peritoneal membrane, in the mesentery, is considerable space, as seen at b. This space is filled up with blood-vessels, nerves, lacteals with their small glands, soon to be described, all bound together by the com- mon packing material of the body, the cellular tissue. You see, therefore, that the mesentery subserves more than one use. Besides fastening the whole tract of the intestinal canal to the spine, so as to guard it against accident, it acts as a secure medium for the communication of the blood-vessels and nerves with the intestines. And, besides, as you will soon see, it con- tains the little tubes which convey all the nutriment into the blood for the growth and repair of the body. 91. I now go on to describe the remainder of the process of digestion. The chyme, (§ 83,) as it passes into the small intes- tine from the stomach, has mingled with it the bile and the secretion of the pancreas. These are poured into the intestine at the point represented at 6, in Fig. 14. These secretions un- doubtedly have some agency in separating the nutritious part 58 HUMAN PHYSIOLOGY. Chyme. Chyle. Luteals. Thoracic duct. of the chyme from that which is not so. When thus separat- ed, it is absorbed by the innumerable small vessels, called lac- teals, which are situated in the mucous membrane. This nutri- tious part of the chyme is a milky fluid, called the chyle. The lacteals which absorb it are little tubes or ducts. These enter certain glands, called the mesenteric glands, for the purpose of having some effect, we know not what, produced upon it. They then pass on, as seen in Fig. 17, to pour their contents into the FIG. 17. .'- ORIGINS OF LACTEALS. SECTION OF INTESTINE SHOWING THE LACTEALS. thoracic duct. This duct, which is about the size of a common quill, running up on the left side of the aorta, the great artery of the heart, pours its contents into the junction of two veins at the top of the chest. As the circulation of the chyle in the DIGESTION. 59 Mechanical continuance of the thoracic duct. Chyle makes blood. thoracic duct needs all the mechanical help that it can have, the mode of the joining of this duct with these veins is calculated to facilitate the freeness of the discharge of the chyle. As the two large currents in the veins, v and v, v, in Fig. 18, FIG. 18. JUNCTION OF THE THORACIC DUCT WITH THE VEINS. unite, there is created, by the forward motion of these cur- rents, a tendency to a vacuum at the angle at which they meet, the point where the thoracic duct, t, d, opens. There is, therefore, a suction power, as it is termed, exerted upon the fluid in this duct. The chyle, thus mingled with the blood, becomes a part of it. Or rather, I should say, that the blood is made from the chyle, and, as it is constantly used for formation and repair in all parts of the system, it is thus as constantly replenished. The material by which all the textures of the body are made and are kept in repair, is furnished to the system through this small duct, in the form of a milky fluid. You observe in Fig. 17, certain lymphatic vessels. These are trunks of absorbents, hereafter to be spoken of particularly, which bring a fluid called lymph, to be mingled with the chyle, and to be poured with it into the circulation. 92. The extent of surface on which the absorbent lacteals open can not be appreciated, if you look merely at the outside of the small intestines. It can be done only by looking at the inner mucous coat. This coat is really much more extensive than the outer coat, or the middle one, the muscular, and it is full of folds, as represented in Fig. 14, on page 52. The ob- 60 HUMAN PHYSIOLOGY. Extent of absorbing surface in intestines. Alimentary canal in different animals. ject of this is to offer a very large absorbing surface to the chyme as it passes, and also to prevent its passing along as rapidly as it would if the mucous surface were perfectly smooth, instead of having folds. Before leaving this subject, I would again call your attention to the analogy which exists between absorption in animals and in plants. The lacteals do for the animal in its stomach, what the absorbents do for the plant in the extremities of its roots. Both absorb and assimilate nutri- ment. The function is the same. It differs in the two cases only in the circumstances under which it is performed. 93. The digestive apparatus varies much in different animals, according to the kinds of food on which they live. As a gene- ral rule, the more the food differs in character from the animal itself, the more complicated and extensive is the apparatus. Thus, the herbivorous animals have a very long alimentary canal, and the beginning of it, the stomach, is a complicated organ. While, on the other hand, in the carnivorous, the flesh which they eat being very, much like their own flesh, and, there- fore, not requiring very much of a process of assimilation, the stomach is a simple organ, and the alimentary canal is very short. In the sheep, for example, the alimentary canal is about twenty-eight times the length of the body, but in the lion it is only three times its length. In man, who lives on a mixed diet, the alimentary canal is about six times the length of the body. 94. The stomach is more complicated in animals that chew the cud than in any other animals. It has four distinct cavities, and, as you will see, a singular mechanism is called into opera- tion in managing the .food as it passes through them. In Fig. 19, you have a representation of the stomachs of the sheep, as they appear exteriorly. The course which the food pursues is this. As the animal crops the food, it passes into the first sto- mach, which is little else than a great reservoir to hold it and to soak it. Then it passes into the second stomach, from which it is returned into the mouth. On being swallowed again, it passes from the oesophagus into the third, and thence into the fourth stomach. In Fig. 20, you see the interior of these four stomachs; and by the aid of this I will describe the process of digestion in the sheep more particularly. You see the very large first stomach, or paunch, in which the food is accumu- lated. It is not yet masticated thoroughly, for the animal has swallowed it as fast as he could, and packed it away in this reservoir. From this it is passed, in small quantities at a time, DIGESTION. 61 Digestion in the sheep. (ESOPHAGUS. _____ ORIFICE OF STOMACH. 3d STOMACH. STOMACHS OF THE SHEEP. FIG. 20. INTERIOR OF THE STOMACHS OF THE SHEEP. into the second stomach, the honey-comb, so called from the peculiar network of folds in it. Here the food is rolled up into balls by the action of the muscular fibres in this network. 6 62 HUMAN PHYSIOLOGY. Digestive apparatus in birds. Different in the grain-eating and the flesh-eating. Each ball of food is passed up through the oesophagus into the mouth, where it is chewed and thoroughly mixed with the saliva, in doing which the animal seems to have great enjoyment. Then it is swallowed, and, as it passes from the oesophagus, in- stead of going into the paunch, as it did when swallowed the first time, it is directed through the groove seen in the Figure into the third stomach, the manyplies. This has many folds, like the leaves of a book, so that the food is exposed to a large surface in this cavity. It passes from this to the fourth sto- mach, the reed. Here, and here only, it is acted upon by the gastric juice. This, therefore, is the true stomach, all the other cavities furnishing only preparatory steps to the true process of digestion. It is from this fourth stomach that what is called the rennet is taken. When fluid matter is swallowed, it goes directly into the second stomach, and not into the first, the paunch; so that, in the case of the sheep, the drink goes one way, and the solid food another. And, what is still more singu- lar, while the animal is a suckling, the milk passes directly into the fourth stomach through the third, which has its folds so closed together as to form a mere tube to conduct it to its des- tination. And the great paunch and the honey-comb are wholly useless until the animal begins to crop its food for itself. 95. In birds, the digestive apparatus is necessarily very peculiar, from the fact that they do not masticate their food. They have, on this account, an arrangement in the stomach itself for grinding the food. In the cavity called the gizzard are two opposing surfaces, made very hard, so that by rub- bing together they bruise the grains; and while they are thus ground, as between two millstones, the gastric juice is poured down upon them from above. This arrangement is seen in Fig. 21, which represents the digestive apparatus in the turkey laid open. At b is the gizzard, showing the two hard surfaces, which are rubbed together by the stout muscles that make the great bulk of the organ. Above, at a, are the glands which pour forth the gastric juice. And above this part of the stomach there is, in all grain-eating birds, a large sac bulging out from the oesophagus, called the crop, 'which is a reservoir for the food, just as the paunch is in the ruminating animals. In those birds that live on flesh or fish there is no such grinding apparatus; and the walls of the stomach are quite thin, and it presents no hard surfaces. 96. It would be interesting, were it consistent with the plan DIGESTION. 63 Digestion in the turkey. Digestive apparatus in different animals. FIG. 21. STOMACH OF THE TURKEY. of this book, to go into a further examination of the varieties in the digestive apparatus in different animals. They have a very wide range, being according to the wants of the animal in each case. The kind of food, the mode of life, and the pur- pose which the animal is designed to fulfill, are the circumstances which govern these variations. The proportion which the di- gestive apparatus bears*to^pther parts varies very much; and in some of the lower orders of animals, the body seems to be all stomach. In such cases, the only appendages are those which seize the food and direct it into the orifice of this organ. This 64 HUMAN PHYSIOLOGY. Apparatus of the circulation. Heart, arteries, veins, capillaries. is the case with the hydra, represented in Fig. 1. And, what is very singular, the outside of the body of this animal is just as capable of acting as a stomach as its inside. For you may turn it inside out, as you can a stocking, and yet it will go on to catch and digest its food as usual. But, wide as the varia- tions are in the digestive apparatus of animals, the same com- mon object is aimed at in all—the assimilation (§ 10) of nu- trient substances to the animal, to produce a material from which its structure can be built and kept in repair. There is, therefore, much that is common to them all in the modes in which this object is accomplished. And even the analogy which exists between the animal and plant, in regard to assimi- lation, does not relate to the fact alone, but in some measure to the modes in which the process is effected. CHAPTER VI. CIRCULATION OF THE BLOOD. 97. In the last chapter I described the manner in which the blood is made from the food. The blood, thus prepared, is circulated in every part of the body, that it may be used for the purposes of construction and repair. The apparatus by which this is done acts, as I have before said, as the common carrier of the material which is used everywhere in the body by the laborers, the builders, to whom it is thus brought. 98. This apparatus consists of several parts—a great central organ, the heart, situated in the chest; the arteries, the tubes by which the blood is conducted to all parts of the body • the veins, other tubes, which bring the blood back to the heart- and capillaries, a network of exceedingly minute vessels, throuo-h which the blood passes as it goes from the extreme arteries into the beginnings of the veins. The blood goes from the heart through a large artery, called the aorta, which sends forth branches; and these divide and subdivide, so that the extreme arteries, through which the blood flows into the capillary net work, are very minute. And the veins which receive the blood from this network to carry it back to the heart, are equallv minute; but joining together more and more, as they proceed THE CIRCULATION. 65 Heart a forcing and suction pump. Arteries firm tubes. Why. toward the heart, they are at length all united into two great venous trunks, one from above and the other from below, which pour their contents into this organ. The capillaries, taking their name from the Latin word, capilla, a hair, are so small that they can not be seen by the naked eye. In any small cut, the blood which oozes out conies from multitudes of these vessels. They serve to hold the blood, while the formative ves- sels, that construct and repair the body, may select from it such materials as they need for their purposes. 99. The heart is a great central forcing and suction pump, in the midst of this circulating apparatus. When it contracts, it forces the blood out through the aorta and its branching ar- teries into all parts of the system. And when it enlarges or dilates itself, it, by suction, as it is termed, receives the blood returning from the system through the veins. The blood never ceases to go these rounds. The necessity for this continual motion you will perceive as I proceed with the development of the subject. 100. The arteries differ from the veins in their structure and arrangement. The arteries are firm tubes, while the veins are lax in their structure. The object of the difference is obvious. As the blood is forced into the arteries by the powerful action of the heart, it is necessary that they should be strong and firm, else, they would be liable, to dilatation and rupture, and death would frequently result. As it is, it is not a common event to have an artery dilate and burst, though it does occa- sionally happen. When dilatation does occur in an artery, it is called an aneurism. But the arteries need to be firm, not only for the sake of security against rupture, but also that the force of the heart may propel the blood to the extremities of the arterial system. If the arteries were lax tubes, like the veins, the impulse would soon be lost in the yielding tubes, and > the blood would move very sluggishly in the small arteries at a distance from the heart. What we call the pulse, is caused by this impulse. If the arteries were lax tubes, the pulse would not be felt at any great distance from the heart. Instead of being distinct, as it now is, with every beat of the heart almost to the very extremities of the arterial system, it would be ren- dered confused by the yielding of the tubes, even quite near the heart, and at a distance from that organ it would be en- tirely lost. 101. Besides the firmness of the arteries, there is another circumstance which favors the freeness of the flow of blood 6* 66 HUMAN PHYSIOLOGY. Different arrangement of arteries and veins. through them. It is their mode of division. The branch of an artery leaves the main trunk at a sharp angle, making thus only a slight deviation from the direction of the current; while, on the other hand, in the veins where the current flows in an opposite direction, the branch unites with the trunk at nearly a right angle. This difference is represented in Fig. 22 ; 1 being the artery, and 2 the vein. FIG. 22. ARTERY AND VEIN. 102. The venous system has a much greater capacity than the arterial. That is, all the veins of the body are together ca- pable of holding more blood than all the arteries are. And the blood moves very rapidly and directly from the heart through the arteries, but it comes back to the heart quite slowly through the veins. Every thing is arranged to promote this rapid cir- culation through the arteries, while the venous system is calcu- lated for a slow but sure progress of the blood back to the heart. To secure this, valves, made of folds of the inner lining of the veins are so arranged as to prevent the blood from flow- ing in the wrong direction. Fig. 23 represents a vein cut open so as to show these valves. A shows the valves as they appear when the vein is laid open and spread out; B, as they appear when the vein is simply laid open; and C represents the ap- pearance of the outside of the vein where there are valves. THE CIRCULATION. 67 in veins. Dangerous to wound an artery. Therefore well guarded. FIG. 23. VALVES IN THE VEINS. The need which there is of this help to the circulation through the veins is obvious. The suction power of the heart is not competent, unaided, to move the blood throughout all the lax venous system. These pocket-like valves, therefore, are made in the veins to assist the circulation there. They do so in this way. Every motion of the muscles or other parts about the veins tends to keep the blood in motion, and the valves serve to prevent this motion from being in the wrong direction. The difference in force and velocity with which the blood moves in the arteries and in the veins, is made manifest when they are wounded. The blood flows from a wounded vein in a slow and steady stream. From an artery it flows rapidly, showing the impulse of the heart in its jets, which correspond exactly with the pulse. Hence comes the danger in wounding an ar- tery, while the wound of a vein is ordinarily attended with no danger. Accordingly, we find that the "Maker of our bodies" has so placed the arteries that they cannot easily be wounded, while many of the veins are quite freely exposed. The arteries are deeply seated, except in some few cases where this is im- possible ; but the veins are often superficially situated. You can see this, for example, in the bend of the arm. Some large veins appear there just under the skin, while the artery which supplies the arm is imbedded among the muscles and tendons. In eveiy part of the body, the most secure spot is chosen for an artery. Thus, at the knee joint, the artery, instead of run- ning over the surface of bone, where it would be liable to be 68 HUMAN PHYSIOLOGY. Few arteries superficial. Mode of stopping the bleeding of an artery._______ wounded, lies deep in the ham at the rear of the joint. The same is true of the elbow joint, just alluded to, and of other parts of the body. Although there are arteries everywhere, they are so uniformly deeply seated, that it is only in a few lo- calities that you can readily find one. You can feel one pul- sating at the wrist, and also on the temple. Here the arteries are superficial, only because it is impossible that it should be otherwise. 103. When the physician bleeds a patient, he commonly does it at the bend of the arm, as being the most convenient place for the operation. A ligature of some sort, as a ribbon, is tied around the arm above the elbow, with sufficient tightness to interrupt the flow of blood toward the heart in the super- ficial veins, but not so tightly as to prevent the free supply of blood to the arm by the artery. It is commonly tied as tightly as it can be without stopping the pulse at the wrist. An open- ing is then made in one of the veins; and, as the blood flows freely into the arm from the heart through the artery, on its return, so much of it as passes through the opened vein is dis- charged at that point. 104. It will be proper here to give some practical instruc- tion, in regard to stopping the flow of blood from a wounded artery, as many lives have been lost from the ignorance of by- standers when such accidents have happened. Enveloping the part in cloths, which is so commonly done at such times, does no good, but only serves to catch and conceal the blood as it flows. Pressure upon the artery, on that side of the wound which is toxoard the heart, will of course interrupt the supply of blood from this organ to the wound. Firm pressure with the thumb will do it. But the pressure must be made at the right point, that is, directly upon the artery. You may not, in all cases, press upon the right spot at once. If you do not, the blood will continue to flow. In this case, press at different points, until you find the point at which you see that pressure stops the flow of blood from the wound. But you may not be able to find the right spot. If you can not, you can tie a slip of strong cloth or a handkerchief around the limb, above the wound, and twist a stick in it until the bleeding stops. In one or the other of these ways, you can prevent the loss of blood until the surgeon arrives to take charge of the case. 105. Although there is no such free communication between arteries as exists between the capillaries, there is some amount of communication, and particularly in certain parts of the body. THE CIRCULATION. 69 Aneurism. Communication between arteries. And it is well that it is so, for it sometimes helps the surgeon to save a limb, when he could not do it if there were no com- munication. I have already alluded to a disease of the arteries called aneurism. An artery has three coats, one of which is a strong fibrous one. When this is thinned or ruptured, the other two coats bulge out, forming a pulsating tumour. And, as the blood is constantly pumped into this by the force of the heart, it enlarges, and at length it may burst, and the life of the patient will be destroyed by the loss of blood. When an aneurism formed in a limb, as for example in the ham, the sur- geon, in former times, used to save the life of the patient by amputating the limb above the aneurism. Putting a ligature round the artery above the aneurism would of course stop the flow of blood into it; but it was supposed that the limb would die, in that case, from the want of a proper supply of blood. But it was found, at length, that this was not so; and surgeons now, in such cases, cure the disease, and save the limb too, by tying the artery. Immediately after the operation the limb is cold, and there is plainly very little circulation in it. But in a few hours the circulation becomes free, and in a little time it is as well established as ever. This is effected by the communi- cations which exist between the branches which go off from the artery above the aneurism, and those which go off below it. It is obvious, however, that this would not be thoroughly effected if no change took place in the size of the communicat- ing arteries. But this change does occur. Some of them be- come enlarged to meet the necessity of the case. This is a most interesting fact; and so is also the fact, that these commu- nications between branches of arteries are very common in the neighborhood of those places in the body, where aneurism, from strains produced by violent and sudden motion, is peculi- arly apt to appear. This same provision avails, of course, when aneurism is cured by pressure made upon the artery above it, a measure which modern surgery has found in many cases to be as effectual as tying the artery. 106. There have been great differences of opinion among physiologists, in regard to the proportionate amounts of agency that the different parts of the apparatus have in carrying on the circulation. The heart manifestly exerts the chief agency, both by its forcing and its suction power. You can get a clear idea of the manner in which it exerts these two forces in this way. Fill a ball of India rubber, to which a tube is attached, with water, and immerse the tube in water in a vessel. If you 70 HUMAN PHYSIOLOGY.___________ Action of the heart illustrated. Agency of the capillaries in the circulation. press the sides of the ball together, some of the water is forced out into the vessel. This represents the contraction of the heart. If, now, you allow the ball by its elasticity to resume its round shape, the water rushes into it from the vessel. This represents the dilatation of the heart. The dilatation of the ball results from its elasticity; and so it is supposed by some that the dilatation of the heart results from the same cause, its contraction alone being produced by muscular action. Whether this be so or not, the dilatation is an active one, and the blood rushes into the heart from the veins by suction, as it is termed. The dilatation is so active that, as has been shown by experi- ments on animals, even a great amount of pressure is not able to prevent its taking place. 107. But, great as the agency of the heart is, it is not true that it is the only moving power, and that the arteries and veins are mere passive conducting tubes. There are various phenomena which show that the arteries, the capillaries, and even the lax veins, exert a considerable agency in circulating the blood. I will merely allude to some of these phenomena. Determina- tions of blood to particular parts show that the blood-vessels have an active agency in the circulation. In inflammation of any part, there is an increased activity of the particular portion of the circulating apparatus supplying that part. In the act of blushing, there is a local activity of the circulation somewhat independent of the heart. This is also true of the circumscribed flush of hectic. 108. There is one portion of the circulation in which the active agency of the capillaries is especially manifest. The veins, as I have told you, receiving the blood from all parts of the body, at length are all united into two veins, which empty their contents into the heart. But there is a very remarkable exception to this. The veins which collect the blood from the viscera in the abdomen unite in one large trunk, called the vena portae; and this, instead of pouring its contents into the large vein that goes up to the heart, divides, like an artery, into branches, which take all this Wood to the liver for the manufac- ture of bile. Fig. 24 represents this circulation of the vena portae. 1, 1, are the veins coming from the intestines ; 2 is the trunk of the vena portae ; and 3, 3, are the branches of it dis- tributed in the liver. Now, it can not be pretended that the suction power of the heart extends its influence through the veins that bring the blood from the liver, then through the capillaries of this organ, and then through all the veins that bring the THE CIRCULATION. 71 Circulation in the liver. Why the veins are full and the arteries empty after death. FIG. 24. CIRCULATION OF VENOUS BLOOD IN THE LIVER. blood to the liver, even to the capillaries of the abdominal vis- cera. There must be, in this case, some propelling power in the capillaries, and some, too, also in the veins. If there were not, another subordinate heart would obviously be needed in the vena portae, to pump up the blood from all the veins of the abdominal viscera, and then to send it through all its branches into the capillaries of the liver. 109. The veins have a less active agency in the circulation than any of the other parts of the apparatus. It is for this reason that commonly after death the veins are found quite full of blood, while the arteries are nearly empty. The appa- ratus of the circulation may be regarded as forming a circle of organs in this order—the heart, the arteries, the capillaries, and the veins. The blood is constantly going the rounds of this circle. It is plain that, as the apparatus is about to stop, there must be an accumulation in the weakest, least active, and most relaxed of this circle of organs. The arteries and capillaries force the blood into the veins to the last moment of life. This effect 72 HUMAN PHYSIOLOGY. The blood changed in the capillaries from red to dark._______________ probably extends no further than the smaller veins; but the heart, by its active dilatation, draws the blood from them into the larger veins. And as these two forces, at the two ends of the venous system, are at work up to the last moment, the whole of this system is filled with blood. 110. The fact, that the larger arteries are commonly found nearly empty of blood after death, gave the ancients the idea that air circulated in arteries, while blood circulated in veins. Hence, the name, artery, is derived from two Greek words, sig- nifying to hold air. And hence, also, by long established cus- tom, in common language, the blood is spoken of as running in our veins; and it would sound strangely, if, in common, and especially in poetical language, we should speak of it as running in our arteries also. Although there were from time to time some glimpses of the true idea of the circulation, it was not really developed and demonstrated till about two hundred and thirty years ago. Harvey spent eight years in maturing his ideas on the subject. When he published them, they en- countered much opposition; but he lived long enough to see them almost universally received by the medical world, although the profession was in a much less enlightened state than it is at the present day. 111. I will now take you a step farther in the development of the plan of the circulation. I have said that the office of the arteries is to conduct the blood to the network of capil- laries, and that in the capillaries the blood has reached its place of destination where it is to be used. The formative ves- sels, appended to the capillaries, take from the blood what they need for their various purposes, and at the same time there is added to the blood refuse matter from the waste of the tissues. The blood, then, is changed while it is in the capillaries. You see the change in its color. In the arteries it was red; but, after passing through the capillaries, it appears in the veins of a purple color. It is also as much changed in other properties. It is no longer fitted to nourish the body. It would even prove a poison to any organ if it should flow into its capillaries. If it should, for example, be sent to the brain, instead of bright ar- teriaj blood, that organ would cease to do its office ; insensibility would ensue, and life would soon be destroyed, if the flow of red blood could not be established. 112. This purple blood, which comes back to the heart from the capillaries by the veins, must, therefore, be in some way changed to red blood, before it is again sent all over the system THE CIRCULATION. 73 Change in the blood in the lungs. Course of the circulation. through the arteries. This change is effected in the lungs. As the purple blood returns to the heart, it is sent by the heart to the lungs, in order to be exposed to the air before it is sent again over the system. For this purpose there are two circula- tions, and the heart is a double organ; or rather, there are in effect two hearts for the two circulations, for the two sides of the heart have no communication with each other. The appa- ratus for all this is very complicated, but I think it can be made clear to you. 113. I present, first, a diagram, which is intended to repre- sent merely the course of the circulation, without regard to proportionate size, or to minutiae in the arrangement of the ap- paratus. Let a represent the right side of the heart, c the left side, b the lungs, and d the general system of the body. The arrows show the direction in which the blood flows. In all the shaded part the blood is venous or purple, and in the part not shaded it is arterial or red. We will now take some point of beginning, and trace on the Figure the course of the circulation. FIG. 25. DIAGRAM SHOWING THE COURSE OF THE CIRCULATION. We will start at or, the right side of the heart. The blood re- ceived here, of a purple color, from the whole body by the veins, is sent by the heart to b, the lungs. Here it changes to red blood, and passes by veins back to the heart—but, observe, it is to the left side of the heart, c. It is now sent by this left half of the heart to all parts of the system, represented by d. Here, in the capillaries, it ischanged to purple blood, and goes back by veins to the right side of the heart, a, the place where we started. 74 HUMAN PHYSIOLOGY. Two circulations and two hearts. Arrangement of valves. 114. You see, then, that there are two separate circulations, one through the general system, and the other through the lungs alone. In both circulations the blood is sent from the heart by arteries, and is brought back to it by veins. But notice that, while in the general circulation the red blood is in the arteries, and the purple in the veins, in the circulation through the lungs it is reversed—the red blood is in the veins, and the purple is in the arteries. So, also, while the change of the blood in the capillaries of the general system is from red to purple, in the capillaries of the lungs it is from purple to red. 115. There are not only two sides or halves of the heart, separated entirely from each other, but each of these sides has two apartments, with valves or folding doors between them, so arranged that the blood can pass one way through them, but not the other. There are also valves at the beginning of the great artery of the heart, the aorta. These are so arranged that the blood can go freely out of the heart into the artery, but not a drop can get back from the artery into the heart. There are similar valves, also, at the beginning of the great ar- tery, by which the purple blood is sent from the heart to the lungs. 116. In Fig. 26, is represented a section of the right side of the heart, for the purpose of giving you an idea of the arrange- ment and the relative size of the two apartments. The auricle, a, so called because a part of it has some resemblance to an ear, receives the blood from the whole system by two large veins, b, b, called the venae cavce. From the auricle it passes into the ven- tricle, v, which by its contractions sends it to the lungs through the pulmonary artery, /. The valve between the au- ricle and ventricle is composed of three membraneous sheets, which are held at their edges by small tendinous cords, d, just as a sail is held by the ropes at its corners. This valve permits the blood to pass from the auricle into the ventri- cle ; but when it attempts to pass back from the ventricle to the auricle, it section of the right pushes back the sheets of the valve, they si°e of the heart. beyig prevented from going too far back by the tendinous cords. There are also valves at e, the bee-inning of the pulmonary artery, which allow the blood to pass through THE CIRCULATION. 75 Relation between the auricles and the ventricles. them into the artery, but no blood can pass through them from the artery back into the ventricle. I shall soon call your atten- tion again to these different valves, that you may see more par- ticularly their structure and arrangement. 117. The auricle and ventricle act in this way in propelling the blood. When the auricle contracts, the ventricle dilates * to receive the blood from the auricle. The valves between them are open while this is taking place. But the next moment the ventricle contracts and the auricle dilates. You at once see, that if now the valves between them should be open, the blood would be forced back into the auricle. But the membranous sheets of these valves shut upon each other as the ventricle contracts, and thus prevent the blood from going back. It therefore is discharged through the pulmonary artery, /, the valves there being open. And when the ventricle dilates, you can see that the blood would, from suction, enter it from the artery as well as from the auricle, if the valves at the orifice of the artery should remain open. They are accordingly shut when the ventricle dilates. You see, then, that when the auricle dilates and the ventricle contracts, the valves between the auricle and ventricle are closed, and the valves at the mouth of the pulmonary artery are open; and, on the other hand, when the ventricle dilates and the auricle contracts, the valves between them are open, and the valves of the pulmonary artery are closed. 118. Dr. Carpenter has a very good illustration of the rela- tion of the actions of the auricle and ventricle, in a representa- tion given in Fig. 27. The apparatus which is represented consists of two pumps, a and b, the pistons of which move up and down alternately. These are connected with a pipe, c, /, in which there are two valves, d and e, opening in the direction of the arrows. The portion c of the pipe represents the venous trunk discharging its blood into the heart, and the portion /, the artery which is the outlet for the blood. The pump, a, represents the auricle, and the pump, b, the ventricle. When the piston in a is raised, the fluid enters through c to fill it by suction, as it is termed. When, now, its piston is lowered, the fluid is'forced through the valve d into the pump b, (which re- presents the ventricle,) whose piston is at the same time raised to receive it. And when the piston in b is lowered in its turn, • Thi« HilntHtion is an active one, as was stated in § 106, when speaking of the heart as aTJholt TheventrlckJ does not dilate because the blood is forced into .t, but the blood rushes into it because it dilates. 76 HUMAN PHYSIOLOGY. Ventricles larger and stronger than the auricles. Valves of the aorta. FIG. 27. the fluid being prevented from returning into a, by the closure of the valve d, is forced through the valve e into /, representing the discharging tube, the artery. At the same time, a fresh supply of fluid is received into a by the raising of its piston. 119. I have described the auricle and ventricle of one side of the heart, the right side. The left side is constructed very much in the same way. You will observe, in Fig. 26, that the ventricle is much more capacious than the auricle. The auricle is indeed the antechamber to the ventricle. The ventricle, too, you see, is much thicker in its walls. It is made very strong, because it does by far the principal part of the work. I remark here, in passing, that the size of the whole heart is about that of the closed hand of the individual. 120. I will now call your attention to a more particular view of the valves of the heart. We will take, first, the valves which are at the beginning of the aorta, the great artery of the body, going out from the left ventricle. These are very much like the valves of the veins seen in Fig. 23. There are three of them. They are like little pockets arranged around the ori- fice of the artery, and looking toward the tube of the artery. Of course, when the ventricle contracts, and forces the blood into the artery, these pockets are pressed by the blood flat against the sides of the artery. But when the ventricle dilates, and the blood attempts to go back from the artery into the ventricle, it gets into these pockets, and bulges them out toward the heart, and thus the mouth of the artery is closed. But you can see that if these pocket-like valves had plain curved edges they would not effect a perfect closure. There would be a THE CIRCULATION. 77 _________________Peculiar provision in the valves of the aorta. little space in the very middle of the orifice of the artery which would be left open. This is made plain by Fig. 28, which pre- sents the orifice of the artery with its closed valves, as it would appear seen from the interior of the heart, FIG- w- if the three valves had plain curved edges. There would be a space left between them. But this difficulty is remedied by a very simple contriv- ance. A little fleshy projection is placed upon the middle point of the edge of each valve, of such a size that the three projections together just fill the space A. When, there- fore, the valves are closed, no blood can go back from the artery into the ventricle. This arrangement is shown in Fig. 29, in which the aorta, a, is laid open and spread out, so as to show the three valves with their projections on the edges. At b and c, are the openings of the two arteries that supply the walls of the heart FIG. 29. VALVES OF THE AORTA. with blood for their growth and repair, for the heart is con- structed and repaired from its own blood. The valves at the orifice of the pulmonary artery are arranged in the same man- ner as those which are at the orifice of the aorta. 121. The valves which are between the auricles and the 7* 78 HUMAN PHYSIOLOGY._____________ Arrangement of the valves between the auricles and ventricles.__________ ventricles I have already partially described. They are folds of strong white membrane, their edges being held by numerous small tendinous cords. And these cords are manned, as we may express it, by muscles attached to the walls of the heart. The office of these muscles is to hold on to the cords that are fast- ened to the edges of the valves, and prevent these sheets of membrane from flapping back too far when the powerful ven- tricle contracts. It is by a nice adjustment of forces that these valves act with such exactness. They are of greater extent than the valves which are at the mouth of the aorta and the pulmonary artery, and, therefore, it would not do to leave them to act alone, as those valves do, upon simple mechanical princi- ples. The living muscular fibre must be introduced as the agent to control and regulate these principles in their applica- tion here. If it were not done, the consequence would be, that when the ventricle contracts with prodigious force, as it some- times does when the circulation is in a great state of excitement, the light tendinous fastenings would be ruptured by the pres- sure of the blood upon the valves. As it is now, the strong but yielding muscular bundles, to which these tendons are attached, regulate with great exactness the closing of the valves. Even if there were no need of any regulation, by muscular action, of the movement of these valves—if the tendons would, in all cases, let the valves go back to just the right point—as they are not extensible, and have no elasticity, it is manifest that there would be more danger of rupture than there is with the present arrangement. The tendons cannot be stretched, and, therefore, under great pressure they might break. In Fig. 30 is a representation of a portion of this valvular apparatus. The engraving was made from a drawing of the part taken from the heart, and pinned upon a board for the purpose. At m, you see the sheet of membrane; o, o, are two of the muscles attached to the inside of the ventricle, to hold on to the ten- dons, h, that are fastened to the edge of the membrane. This membrane is now in the position that it is when the valves are open, that is, lying in a line with the little tendons and their muscles. But when the ventricle contracts, the blood, pushing against the membrane m, carries up the free edge to which the tendons are fastened, which, meeting the free edges of the other valves, closes with them the communication between the auricle and ventricle. 122. In looking at Fig. 26, you observe that, while there are valves between the auricle and ventricle, and at the mouth of _______THE CIRCULATION. 79 No valves at the openings of the venae cavse. Why this. FIG. 30. PART OF THE VAVULAR APPARATUS BETWEEN THE AURICLE AND THE VENTRICLE. the artery going out from the ventricle, there are none at the openings of the two venae cavce, the veins that pour their con- tents into the auricle. Why is this ? Why is there no need of valves here to prevent a regurgitation into these veins when the auricle contracts ? It is because that, as the auricle con- tracts, there is at the same time the dilatation of the strong ventricle, making, of course, a suction in that direction so powerful as to counteract most fully any tendency to regurgita- tion into the veins. You readily see, that if the arrangement were reversed, and the auricle were stronger than the ventricle, then, when the auricle contracted, there would be regurgitation into the venae cavae, if there were no valves there to prevent it. The same remarks could be made in regard to the pulmonary veins, that pour their contents into the left auricle. 123. Having thus examined the heart in detail, you are now prepared to look at it as a whole. For this purpose, I present to you, in Fig. 31, a front view of the heart, in which a is the right auricle, receiving the purple blood from the whole body by the two large veins, h and i, called the vence cava ; b is the right ventricle, that receives the blood from the right auricle, and sends it to the lungs by the pulmonary artery, f; c is the left auricle, which receives the red blood from the lungs, by the pulmonary veins, g, g, g; d is the left ventricle that re- ceives the blood from the left auricle, and sends it all over the body through the aorta, e. You observe, that you see but a part of the left auricle and ventricle, they lying partly behind the right ventricle. You do not see the very beginning of the 80 HUMAN PHYSIOLOGY. General view of all the parts of the heart. FIG. 31. FRONT VIEW OF THE HEART. aorta, for, as it rises from the left ventricle it is at first con- cealed behind the top of the right ventricle and the beginning of the pulmonary artery. It then forms an arch, from which it sends forth branches to the head and upper extremities; and it afterwards passes down behind the heart, to supply with its branches the trunk of the body and the lower extremities. In the line of division between the two ventricles, b and d, you see one of the coronary arteries, as they are called, which, coming from the beginning of the aorta, as described in § 120, supply the walls of the heart with blood. 124. To make you quite familiar with the relations of the different parts of this complicated organ, and with the course of the blood through its different apartments, I give you, in THE CIRCULATION. 81 _________ Course of the blood through the different cavities of the heart. Fig. 32, a map of the heart, with the names placed upon the parts. I will describe the circulation with this map before you. The dark blood is received from all parts of the body by the vence cavce—from the parts above by the descending cava, and FIG. 32. MAP OF THE CIRCULATION. from the parts below by the ascending cava. These veins pour the blood into the right auricle. From this it passes into the right ventricle, which sends it by the pulmonary artery to the lungs. From the lungs it returns by the pulmonary veins to the left auricle. It then passes into the left ventricle, from which it is sent by the aorta to all parts of the body. 125. In Fig. 33 is represented the heart, situated between the two lungs, with the arteries which carry blood from it, and 82 HUMAN PHYSIOLOGY. Situation and connections of the heart. Its harmonious action.__________ the veins which pour their blood into it. The lungs are repre- sented as being drawn apart to the right and left m front, so as to expose fully the heart and its vessels. The sac containing the heart, and the packing cellular tissue are removed, so as to lay the heart and its vessels bare. At a is the trachea or wind- pipe ; on either side of it are the two arteries, the carotids, which go to the head; c is the artery which goes to the arm; 6, b, are the jugular veins coming from the head, d, d, the veins FIG. 33. LUNGS, HEART, AND PRINCIPAL BLOOD-VESSELS. from the arms, all empting their contents, as you see, into the descending cava; e is the right auricle, receiving the blood from the two cavae; / the ascending cava; g the right ventri- cle, i the left ventricle, and h the descending aorta. 126. I have been thus particular, and have led you through some repetitions in the description of some of the figures, in order that you may get a clear idea of the complicated mecha- nism of the circulation. And now, perhaps, you will inquire, in what way all these four apartments of the heart contract and dilate, so as to have the organ act as one harmonious whole. You have seen how the auricle and ventricle of one THE CIRCULATION. 83 The causes of the two sounds of the heart. Its forward impulse. side act in relation to each other—the auricle contracts when the ventricle dilates, and the ventricle contracts when the auri- cle dilates. Now, the harmony of action between the two sides is preserved by having the two auricles act together, and the two ventricles act together. And this action produces two sounds, which may be heard by applying the ear to the chest of any one on the left side. The first sound is rather a prolonged and heavy one, the second is light and quick. They are very well represented by the syllables lub-tup. The first sound occurs when the strong action of the heart is per- formed, that is, when the ventricles contract. It is owing to several causes. One of these is the impulse of the heart against the walls of the chest; the cause of which I shall speak of soon. Another is the flapping together of the valves between the auricles and the ventricles, to prevent the blood from regurgitating into the auricles, when the ventricles contract to force out their contents. The light and quick second sound is caused principally by the flapping together of the valves at the mouths of the aorta and the pulmonary artery when the ventricles dilate. The pulse (which I have already remarked upon in § 100) is produced by the impulse given to the blood by the contraction of the ventricles. There is, therefore, a pulse in the arteries of the circulation through the lungs, as well as in those of the circulation through the general system. Wherever there is an artery there is pulsation. 127. The impulse of the heart against the front wall of the chest on the left side is easily explained. The heart is so en- veloped by the lungs, that only a small portion of it comes near to the front wall of the chest, and such is the situation of the heart, that this portion comes to the left of the middle line of the chest. The position of the heart is an oblique one, its upper part being both farther back and more to the right than its lower part. Keeping in view this position of the heart, you will readily see how the impulse is produced against the front of the chest at its lower part. The aorta, in going from the heart makes an arch upward and backward, to go down in front'of the spine; and it is the tendency to straighten out, produced in this arch by the force of the blood thrown into it by the ventricle, that causes the throwing of the heart forward by a spring. This is easily seen as illustrated by Fig. 34, in which a is the spinal column ; b, the front wall of the chest; d the heart • and c, the arch of the aorta. When the heart throws the blood into this arched tube it tends to straighten it; 84 HUMAN PHYSIOLOGY. Arrangement of the sac of the heart. Its lubrication. FIG. 34. but, as the aorta is fastened to the fixed spine behind, there can be no impression made in that direction. The straightening of the arch must therefore occur in the other direction, to the front; and therefore the heart is thrown a little forward, as represent- ed by the dotted lines. The change of position thus produced is indeed but slight, but it is sufficient to cause the impulse. The entrance of the blood into the pulmonary artery per- haps aids in the result, but not very materially. 128. The heart, as I have already hinted, is inclosed in a sac, called the pericardium, which, at its upper part, is fastened all around the vessels that proceed from the heart. This sac is lined on the inside by a serous membrane, which also lines the outside of the heart, being reflected over upon it from the pericardium. This membrane forms, therefore, a sac without any outlet. This is made plain by Fig. 35. In this diagram, showing the plan of the serous membrane of the pericardium' a, a are the auricles; v, v, the ventricles; b, c, the vessels pro- ceeding from the heart; p the serous membrane lining the out- side of the heart; pf, the same membrane reflected from the FIG. 35. PLAN OF THE PERICARDIUM. CIRCULATION. 85 ^^^^^ Action of the heart involuntary. Number of its beats. upper part of the heart on to the inside of the pericardium. The arrangement of this membrane, as it fits on to the heart, is much like the common double nightcap, as it fits on to the head; and if it were dissected off whole from the outside of the heart and the inside of the pericardium, it would be like such a nightcap when taken off from the head—that is, a sac without an outlet. Now, this sac is kept moistened by a fluid exuding from its whole surface, so that, as that part of it which lines the outside of the heart, in the motions of that organ, rubs against that part which lines the pericardium, the lubrication prevents any injury from the friction. This lubri- cating fluid is continually renewed, the exhalents and the absorb- ents balancing each other in their action. When the exhalents secrete more fluid than the absorbents can take up, it accumu- lates, making what is called dropsy of the heart. 129. The heart, as you have seen, is a complex arrangement of muscles. And these muscles are wholly involuntary; that is, they are not at all under the direct control of the will. No one can by an exercise of the will make his heart beat slower or faster. As I shall show you in another chapter, this organ is kept at work by its nervous connection with the spinal mar- row. It has no repose, as the voluntary muscles have, unless you call the intervals between the contractions and dilatations of its several parts intervals of repose. The amount of work which it does is enormous, if we calculate it for a lifetime. The heart of an adult beats, that is, each one of the four cham- bers of this organ dilates and contracts, about 70 times in a minute. This would make 100,800 times in 24 hours, 36,792,000 times in a year, and 2,575,440,000 times in a life of 70 years. In children, the action of the heart is much more rapid, and in disease it sometimes reaches in them to 160 or even 200 beats in a minute. It is thus that this organ, situ- ated in the centre of the complicated apparatus of the circula- tion, labors continually, by night and by day, in keeping the blood in motion. The two circulations of the general system and of the lungs are ever going on. The blood is ever moving in all the cavities of the heart, in every artery, and vein, and capillary. It never stops till it is arrested by death. 8 86 HUMAN PHYSIOLOGY. Apparatus of respiration. Air-cells in the lungs. Their si: CHAPTER VII. RESPIRATION. 130. You saw, in the last chapter, that the purple venous blood is sent to the lungs to be changed into arterial blood. The great object of the apparatus of respiration is to introduce the air to the blood, so that it may act upon it, and produce this change. Another object is effected at the same time, viz., the production of the voice, by the striking of the air upon the vocal chords in the larynx, as it is forced out from the lungs. This will be made the subject of a future chapter, and I propose now to show how the chief object of respiration, which is so immediately essential to the continuance of life, is secured. 131. The lungs are spongy bodies, filling up a large part of the chest, and surrounding the heart. They are in common language, the lights ; and you can see what they are in man by observing the lights of other animals. They are composed chiefly of air-tubes, air-cells, blood-vessels, and nerves, packed together with the common packing material of the body, cellu- lar membrane. The spongy lightness of the lungs is owing to the air-cells or vesicles. You can get some idea of the propor- tion of these cells to the solid part of the organs if you inflate the lungs of some animal, as the sheep or calf, by blow- ing into the windpipe. These cells are exceedingly minute. It is in them that the change is effected in the blood. The capillaries holding the blood branch out on the walls of the cells, and the blood is acted upon by the air through the pores of the vessels. The object, therefore, of respiration is to in- troduce the air freely into these cells. The air enters through the windpipe, and this branches out into tubes called bronchi, which divide and subdivide, till they become very minute, and then end in the air-cells. These cells are estimated to be about the -rioth of an inch in diameter. Some calculations have been made in regard to the extent of surface which they would all make if they could be spread out in one sheet. There is of course no great accuracy in such calculations; but we can readily see that the aggregate surface must be immense, and, therefore, the blood is thus very extensively exposed to the action of the RESPIRATION. 87 Air-tubes. Relative situation of the lungs and the heart. air. In Fig. 36 is represented the lung of one'side, d; the branches of the bronchi of the other lung, c, at the lower part ot which, e, they are represented as they branch out minutely to open into the air-cells; b is the trachea or windpipe, and a LUNGS AND AIR-TUBES. is the larynx at the top of it. It is through a chink called the glottis, in the larynx, that all the air passes as it goes into and out from the lungs. This will be particularly described here- after. 132. In Fig. 33, in the last chapter, you see represented the relative situation of the heart and lungs, the lungs being some- what separated, however, from the heart, to the right and left, in order to show that organ fully. In their natural position they are close to the heart, and cover up all of it, except a small portion in front and to the left side, where its beating is so plainly felt. Both the heart and the lungs are suspended in the chest to the upper part of the walls of this cavity, and are fastened also to the spinal column in the rear. The large vessels of the heart, and the bronchi of the lungs, serve as the princi- 88 HUMAN PHYSIOLOGY. Pleura. Mechanism of breathing. Provision for expansion of the chest. pal means of suspending these organs, as you can readily see by the Figure. The lungs are covered by a white, shining membrane, which also lines the inside of the walls of the chest (§ 67.) called the pleura. This is always kept lubricated by a watery fluid, so that, as the lungs expand and the chest moves, the friction will be attended with no inconvenience or injury. You may perhaps ask why, as the lungs follow the walls of the chest in its expansion, they could not have been fastened to these walls throughout their whole surface. The principal reason probably is that, if this were the arrangement, the intimate vascular connection, which would in this case exist between the walls of the chest and the lungs, would ex- pose the delicate texture of these organs more frequently to injury from external violence. As it is now, the effusion, or the inflammation, consequent upon a blow on the chest, is not apt to affect the lung in the neighborhood, because it has no direct connection with it by nerves and blood-vessels. 133. You are now prepared to see by what mechanism the air is alternately introduced to and expelled from the lungs. The chest incloses a large space, which can be made much greater by certain movements of its walls. It is this expansion of the cav- ity of the chest, effected by certain muscles, which, by creating a vacuum, causes the air to rush into the chest through the tra- chea, just as air rushes into the bellows when the space within is expanded by the separation of its walls. That you may un- derstand how the expansion of the chest is effected, I now proceed to describe the chest. In Fig. 37 you see the frame- work of the chest. At b, b, is the spinal column, the grand pillar supporting the walls of this cavity. The ribs, c, c, go from this with a large curve round to the breastbone, a, in front. The ribs, however, do not join directly with the breast- bone, but there are cartilages intervening, as you observe in the Figure. The collar-bone goes from this breastbone across to the top of the shoulder. The ribs are twelve on each side. The lowest two are attached only to the spine, and are called floating ribs. The whole is so constructed as to allow a very considerable expansion of the cavity. As, in effecting this expansion, the ribs are carried upward and forward with the breastbone, the ends of the ribs at the spine move but very slightly. As the chest is kept in constant motion, lightness in its walls is an object of some importance; and, at the same time, it is necessary that the structure should be a strong one, in order to guard effectually the lungs from injury. Both of RESPIRATION. 89 Framework of the chest. Bones. Cartilages. Muscles. FIG. 37. these objects are secured, by having the walls in front and at the side composed of so many bones, well bound together by the muscles which move them. If these bones were all in one, it would be necessary that it should be quite thick, to answer as a defence, and then it would be a heavy and unwieldy thing to move. The cartilages which connect the ribs to the breast- bone are a great safeguard. They give elasticity to the struc- ture as a whole, and the ribs are not very liable to be broken, because of the yielding of the cartilages with which they are connected. 134. This framework is filled out with connecting material, chiefly muscles, which effect the expansion of the chest in in- spiration. First, there is a large expanse of muscle and tendon stretching across the lower part of the chest, separating its con- tents from the contents of the abdomen below. The edge of this muscle, which is called the diaphragm, is fastened to the spine behind, to the end of the breastbone before, and all around the lower ribs. It is arched upward ; and against its concave sur- face press upward the liver and stomach, while the lungs and 90 HUMAN PHYSIOLOGY. Diaphragm. Its action in inspiration and expiration. the heart press downward against its convex surface. The dia- phragm is represented in Fig. 38. The ribs are cut away in front, so as to give a front view of the cavity of the chest, U, c, the lungs and heart being entirely removed. D D is the diaphragm, very high in the central portion, which is tendin- ous, but descending very low at its edges at the sides and in the rear. FIG. 38. DIAPHRAGM. Front View. 135. You can see that, if all the muscular fibres in the dia- phragm contract, the arch will be flattened, and thus the room in the chest will be enlarged. To occupy this new room thus made, the air rushes in through the windpipe. This is inspira- tion In expiration, the reverse movement takes place—the arch of the diaphragm rises, and, compressing the lungs, forces the air out of them through the trachea. In inspiration, as the diaphragm is flattened, it pushes down before it the stomach, liver, &c and hence the pressing out of the abdomen, which is so sensibly felt if the hand be placed upon it during the act of inspiration. In expiration, on the other hand the RESPIRATION. 91 In expiration little muscular action. Elasticity the chief agent. abdomen retreats inward. These two opposite states of the arch of the diaphragm, and of the walls of the abdomen, are represented in Fig. 39. It is a side view, the ribs being cut away. C c is the cavity of the chest, and C a, the cavity of the abdomen. The diaphragm and the abdomen are represented FIG. 39. Side View. as they are in expiration. The dotted line marks the flattening of the arch of the diaphragm, and the projection of the ab- domen, as they occur in inspiration. It is supposed that in ordinary expiration, there is little, if any, muscular action— that as the diaphragm, which in inspiration pushed down the stomach and liver, and thus thrust out the walls of the ab- domen ceases to contract and relaxes, the mere elasticity of the parts below, and especially of the abdominal walls, restores the former condition of things, and so the diaphragm is car- ried upward, and expiration results. When, however, the ex- 92 HUMAN PHYSIOLOGY. Other muscles, besides the diaphragm, act in inspiration._____________ piration is at all forcible, it is produced in part by the action of the muscles of the abdomen and some of the muscles about the chest. 136. While this dome-shaped muscle, the diaphragm, is the principal agent by which the chest is enlarged, there are other muscles which do the same thing in another way. In Fig. 40, a is the spine; c, c, c, the ribs ; 6, the breastbone ; d, the col- lar-bone ; g, the diaphragm. You observe, on the right side FIG. 40. of the chest, certain muscles, i, extending from the spinal column in the neck to the first rib. When these contract, the effect will be to raise this first rib, and all the others, being attached to it, of course follow. And, as the ribs, as you see in Fig. 37, slant downwards from the spine toward the front, the result will be, that all the ribs will be carried together for- ward and upward. This result is the more effectually secured by muscles which pass from rib to rib, as seen at e, e, e, e. In this Figure, the ribs, c, c, c, are left bare on the left side, to show RESPIRATION. 93 ' Arrangement of muscles between the ribs. the arch of the diaphragm, g, the dotted line indicating it on the right side. 137. There are two layers of muscles connecting the ribs, the fibres of which cross each other, as represented at M, in Fig. 41. RR are parts of two ribs. The spaces between the FIG. 41. ribs are filled with muscular fibres, arranged as represented in in the Figure. If the fibres were straight, as at L, they could not bring the ribs as near together as the oblique fibres do. For, as muscles can not shorten themselves, at the farthest, more than one-third of their length, the straight fibres could bring the ribs only one-third nearer together, while it is obvious that the oblique fibres, with the same contraction, can do much more than that. These muscles between the ribs not only, then, help to raise all the ribs as a body, as mentioned in § 136, but they bring each rib nearer to the one above it. This in- creases the expansion of the chest, especially as the ribs are so joined to the spine, that if a rib be moved upward, it must be carried outward as well as forward. You can see, then, that by the operation of these muscles in the neck and between the ribs, the diameter of the chest will be increased from front to rear, and also from side to side. 138. The chest is expanded, then, in two ways—by flatten- ing the arch of the diaphragm, and by raising the ribs. In ordinary quiet respiration, this expansion is effected chiefly by the diaphragm. But when there is a call for more active respiration as in violent exercise, the muscles which raise the ribs act strongly, and hence the heaving of the chest, as it is called. Their action is violent when from disease, as in asthma for example, it is difficult to introduce sufficient air into the lungs. 139. The lungs, heart, &c, accurately fill the chest in all the variations of size to which its cavity is subjected in respiration. V 94 HUMAN PHYSIOLOGY. Change in the blood effected in the air-cells._____________________ For, when the chest is expanded, the spongy lungs swell out to follow its walls, and the air rushes in through the trachea to fill the expanding air-cells. If, now, there were an opening through the walls of the chest, communicating with the out- side of the lung, when the chest expanded, the air would rush in at this opening as well as through the trachea, and the lung would be compressed in proportion to the freeness of the open- ing. This has sometimes occurred from disease and from wounds. If a free opening were made at the same time in both sides, both lungs would be compressed, and death would be produced by suffocation, as really as if some obstruction in the windpipe prevented the air from entering the lungs. 140. I have said that the change in the blood, from purple to red, is effected in the air-cells. The blood and the air are brought very near together for this purpose; and yet they are kept entirely separate, except when, from disease, the blood escapes into the air-cells and air-passages, and is then expecto- rated mingled with air. It is supposed that the air in the cells acts upon the blood through the pores of the vessels containing it, which branch out on the walls of the cells; for if dark venous blood be inclosed in a bladder, the air will act through the pores of the bladder, and gradually change the outer por- tion of the blood to a red color. 141. These air-vesicles, then, do an important work. The change which is effected in them is immediately essential to the continuance of health, and even of life. If the air be in any way shut out from them death occurs at once. And so important is it that they should do their work well, that extraordinary provisions are made to secure an abundance of room for them under all circumstances. For the cavity of the chest, as you have seen in this chapter, can be expanded to a very great ex- tent. It would indeed be difficult to conceive how a greater range of expansion could be secured. As the air-cells are called upon to do more work at some times than at others, there are special provisions for a larger dilatation of the chest than is required in ordinary quiet respiration. Thus when, from violent exercise, the blood is coursing rapidly through the lungs, and more air is therefore needed to change it to red arterial blood, the chest is largely expanded by calling into action muscles, which do but little, if any thing, in ordinary breathing. 142. As the apparatus of respiration is so especially ar- ranged to secure room for the lungs under all circumstances, RESPIRATION. 95 Injury done to the air-cells by compression of the chest. it must be very deleterious to the health of the body to inter- fere with this arrangement. If the expansion of the chest in breathing be limited by any pressure, every air-cell must be embarrassed in doing its part in changing the blood. Either all of them must be unduly contracted, or some of them must become obliterated, so that there will not be as many vesicles as there should be. In either case, the organ is disabled in proportion to the amount of the compression. The blood is not as good as it would be if there were enough vesicles, and they could perform their work without constraint. The vigor of the system is therefore lessened. And, besides, the lungs themselves are especially liable to disease from this unnatural confinement. 143. Much injury is undoubtedly done to the lungs that are thus confined, when any strong exercise is taken. If the chest be left free to expand to its fullest extent when occasion re- quires, this injury is avoided. For when the strongly and rapidly contracting heart pumps the blood in such quantities into the lungs, the widely expanding chest draws in the due amount of air to change the extra amount of blood. All the air-vesicles are ready to do their duty, and, therefore, no violence is done to the delicate texture of the lungs. But if these or- gans be compressed, the dilatation of those vesicles that are not obliterated, in the midst of the commotion of the difficult res- piration, is very unequally effected, and some of them are stretched beyond their proper dimensions. At the same time, the blood must be here and there obstructed in its passage through the lungs, producing what is termed congestion. And if this violence be repeated from time to time, permanent dis- ease will after a while be the result. 144. From the considerations in the two last paragraphs it is manifest, that the interference with the due expansion of the lungs, which so commonly results from the modes of dress in the female sex, must be one of the prominent causes of con- sumption, to say nothing of other diseases arising from this cause. This interference is effected in two ways—chiefly by compression of- the chest directly, but also by the pressure which the load of clothing hanging from the girt waist must make upon the upper part of the abdomen. This latter cause interferes with that forward movement of the abdomen which, as you saw in § 135, is necessary to the flattening of the arch of the diaphragm in the act of inspiration. The extent to which compression of the chest is sometimes carried is seen by 96 HUMAN PHYSIOLOGY. Change of the form and capacity of the chest by compression.___________ comparing the two outlines in Fig. 42. One is an outline, the Venus de Medicis, the universally recognized beau ideal of beauty of form in the female, and the other is an outline FIG. 43. of the form of a lady with an artificially small waist. In Fig. 43 is represented the framework of the chest of its natural size, and as it is sometimes contracted by fashion. The Figures FIG. 43. representing the contraction of the chest may appear at the present time as caricatures, for a very small waist is not con- sidered now to be as essential to beauty in the female form, as it was twenty-five years ago. The truth, as uttered by medical men, has had some effect. But the evil is remedied only in RESPIRATION. 97 Cause of death in drowning. Singular provision in the whale. part. The chest of the female is still too much begirt, in obe- dience to the tyranny of fashion, to allow of the free expan- sion, to secure which such special pains are taken by nature. The evil begins in childhood. The chest is moulded during its growth to the shape which fashion prescribes. It could not be done after the chest has attained its full size. The torture of the compression necessary to do it could not be endured. In childhood, therefore, while the boy's chest is left to grow in its natural shape and dimensions, the girl is begirt so tightly as to embarrass her respiration, because nature is too ungen- teelly large in her patterns to suit her case. The subject is an important one ; but as this book is not designed to treat of hygiene, I can not go into it further. 145. It is the interruption of the change which is effected by the air upon the blood in the lungs, that produces death in drowning. The very common supposition, that considerable water gets into the lungs in drowning, is erroneous. Very little water ordinarily gets in—not enough to occasion any em- barrassment. The difficulty is, that the air is kept out, and not that the water gets in. The drowning person makes at- tempts to inspire, but the moment that the water reaches the epiglottis, the door of the windpipe, it causes at once, by its irritation, a spasmodic closure of the epiglottis, so that almost no water is introduced. In the mean time, the purple blood continues to be thrown by the right ventricle of the heart into the lungs. But the little air contained there soon parts with its oxygen ; and then the change in the blood ceases to occur, and dark blood is sent from the lungs to the heart, and thence to all the organs. These can not go on to do their duty with- out the stimulus of arterial blood. The brain, therefore, gives out, and there is insensibility. The muscles cease to act, and all motion is gone. If a good supply of arterial blood could be furnished to all the organs until breathing could be again commenced, life would be preserved. And there is provision for such a supply in certain animals that can remain under water for some time. For example, in the whale there are larov reservoirs for containing arterial blood, which can be used for°the supply of the organs while he remains under water. When the supply begins to be exhausted, the animal of course has those uncomfortable sensations which a predominance of purple blood is so apt to produce. He manifests his uneasiness by his puffing and blowing, as he rises to the surface, to get a fresh supply of air, and with it a fresh supply of arterial blood in tl)'' reservoirs. 9 98 HUMAN PHYSIOLOGY. Respiration in fishes. Arrangement of the gills. ^_____________ 146. The apparatus of respiration varies in different animals. It appears in three forms—lungs, gills, and trachea or air- tubes. The gills of the fish are arranged in fringed laminae, m order to present by all its minute divisions a large surface ; and these delicate organs are covered with a lid to protect them from injury. The blood-vessels which contain the blood to be changed branch out on the surface of the fringes of the lami- na, just as the blood-vessels in lungs branch out on the surface of the air-vesicles. The air which is to change it is mingled with the water. It acts upon the blood, as the water containing it, after being taken into the mouth of the fish, passes out through these la- minae, as through a sieve. That the air in the water is the cause of the change can be proved by experiment. If a fish be placed in a vessel with its orifice closed, so that no air can enter, it will soon die from suffocation, because the air in so small a portion of water is soon used up. Although the fish can not with his gills use air that is not mingled with water, it is supposed that it is merely because the gills soon become dry when exposed to the air, and that the air would act on the blood in the gills if they were only kept moist. Indeed, in the land crab, that has the power of living for some time out of the water, it has been found that there is a gland in the gill-chamber which furnishes a secretion to keep the gills moist. Gills differ much in their shape and arrangement in the various aquatic animals. In Fig. 44 is represented the arenicola or lob-worm. Here, the gills are in the form of tufts arranged along the outside of the body. They take a somewhat similar form in the larvae of many aquatic insects, as seen in Fig. 45. A large surface is presented to the air contained in the water by the delicate and beautifully arbor- escent gills of these animals. Tn insects, we find the respiration effected by tracheal or air-tubes. These go into all parts of the body, and the air contained in them acts upon the blood in the ves- sels which branch out upon their walls. The in- sect, therefore, has no distinct respiratory organs situated in any one part of the body, but the lob-worm. RESPIRATION. 99 Respiration in insects. Tracheae. Stigmata. air is carried into every part. This seems to be necessary on account of the feeble circula- tion in the insect. The trachea which, as Cuvier says, conduct the air in search of the blood, as the blood has no means of travelling in search of air, open on the surface by stig- mata, as they are called, which are of various shapes and number in different insects. In the grasshopper there are twenty-four, ar- ranged in four rows. You can kill an insect by suffocation by simply covering the stigmata with varnish. In Fig. 46 are represented the trachea in an insect, the nepa or water-scor- pion. The tracheae, as you see, send branches out in every direction, so that air is introduced FIG 46. LARVA OF THE MAY-FLY WING CUT OFF. 2d PAIR OF RESPTRATOKV APrARA'ITK OF THE WA 1 FR-?(OKTMON. 100 ITrMAN PHYSIOLOGY.___________ Respiration in birds. Apparatus for it extensive.__________________ into every part of the body. There are lungs, so to speak, everywhere in the insect. 147. The apparatus of respiration is largely developed in birds for two objects—to provide for the extensive change in the blood which is required by their great activity, and to give lightness to the body. To secure these objects there are air- sacs connected with the lungs, and located in different parts of the body; and in birds that fly rapidly and are long upon the wing, these sacs are very extensive, and even many of the bones are made hollow, and are connected with the air sacs. By this arrangement, the air is introduced extensively to the blood in the capillaries on the walls of these sacs, and at the same time the body is made very light. And the heat gener- ated by the effort of flying must expand the air in the air-sacs and swell them out, and thus make the body lighter. In Fig. 47 is seen this arrangement of air-sacs in the ostrich. The lungs, I, I, are quite small, but the air-sacs, c, c, c, are very large. The orifices by which they communicate with the lungs you see FIG. 47. * U'NGS OF TUl: OS'IT.U H. RESPIRATION. 101 Changes produced in the air in the lungs. in the Figure. In birds of great powers of flight, the air-sacs are much more extensive. This arrangement of air-sacs in different parts of the body of the bird bears some analogy to the trachea distributed in the bodies of insects. 148. You have seen that the object of the apparatus of re- spiration is to change venous blood into arterial, and you have also seen how the air is introduced to the blood in order to effect this change. And now the interesting inquiry arises, what are the actual changes which occur, both in the blood and in the air, in the lungs. If you take a tumbler filled with lime-water, and breathe into it through a tube, the lime- water will become turbid, and will soon deposit a sediment. This is chalk, or carbonate of lime, formed by the union of the carbonic acid gas exhaled from the lungs with the lime in the lime-water. Whence comes this carbonic acid gas, and how is it formed ? In order to answer this question satis- factorily, we must look at the chemical constitution of the air which we breathe. It is composed of two gases, oxygen and nitrogen. In every 100 parts of common air, there are 79 parts of nitrogen and 21 of oxygen. It is found that the oxygen is that constituent of the air which is necessary to life. If an animal be placed in a closed jar filled with com- mon air, he will soon die, and the oxygen will be found to have disappeared, while the nitrogen remains very nearly the same in amount. If, now, you place an animal in a jar of nitrogen, and another in a jar of oxygen, the one in the nitrogen will die immediately, while the other will be very lively until the oxy- gen is mostly used up by his lungs. The animal in the pure oxygen will breathe at first more rapidly than the animal in thenar of common air; and it is thought that oxygen is too stimulating for the lungs, and therefore needs to be diluted with the nitrogen, as it is in the air that we breathe. 149. In the case of both the animal in the jar of air, and that in the jar of oxygen, carbonic acid is found to have taken the place of the oxygen which has disappeared. This gas is made by a union of oxygen with carbon or charcoal. It was formerly supposed that this union is effected in the lungs— that carbon is thrown off from the venous blood in the lungs, and that the oxygen of the air there unites with it, and so car- bonic acid appears in the air expired from the chest. But it has been discovered that the exchange is made in a different man- ner. It is not made in the lungs. The oxygen is absorbed by the blood, and goes with it to the heart to be sent all over the 102 HUMAN PHYSIOLOGY. Changes produced in the blood by the air.________________ system. And it is in the capillaries that the oxygen unites with carbon to form carbonic acid. The union takes place while the blood is changing from arterial to venous, and is an essential part of the change. The carbonic acid thus formed in the capillaries, is brought back to the heart in the venous blood, and is discharged "from the system in the lungs. That the change takes place as stated has been abundantly proved in various ways. • It has been found by experiments which I will not detail, that carbonic acid exists in considerable amount in venous blood; while, on the other hand, there is much oxy- gen in arterial blood. The plain inference from this is, that oxygen unites with the blood as it passes through the lungs, goes with it to the capillaries, and there unites with the carbon, giving us the carbonic acid which we find in the blood in the veins, after it has passed into them from the capillaries. It has been found, also, that if frogs or other cold-blooded animals be placed in hydrogen or nitrogen, (gases which produce no in- jurious effect on them,) they will give off for some time nearly as much carbonic acid as they would have done in common air. In this case, as no oxygen is introduced into the lungs, the carbonic acid can not come from any union effected in these organs between carbon and oxygen, but it must be dis- charged by exhalation from the blood as it is passing through the lungs. Of course the discharge of the carbonic acid ceases after a little time; for, there being no new supply of oxygen by way of the lungs, as there is when the animal is breathing com- mon air, there can be no new formation of carbonic acid. But even cold-blooded animals can not live in these gases for any great length of time, although they are not positively deleterious to them, for oxygen is needed for the continuance of their func- tions. And in the warm-blooded animals, a constant supply of it is necessary—they will die if cut off from this supply even for a short time. 150. The change which takes place in the blood, as it passes through the lungs, occurs to some extent when the blood is ex- posed to the air in any way. Thus, if blood be drawn from a vein into a bowl, the surface of it becomes red by the action of the air upon it. Carbonic acid is discharged from it, and the oxygen of the air takes its place, uniting with the blood, just as the process occurs in the lungs. A larger part of the blood will be thus changed, if it be shaken so as to expose more of it to the air. The change takes place to some extent even if a membrane be interposed between, as when the blood RESPIRATION. 103 Quantity of carbonic acid given out by the lungs. Necessity of ventilation. is inclosed in a bladder. The oxygen of the air, in this case, is introduced through the minute pores of the bladder, and the carbonic acid gas escapes through them. Precisely in this way is the change effected in the lungs, as already stated in § 140. The blood is separated from the air by being con- fined in blood-vessels, and the air in the vesicles acts upon it through the minute pores of these vessels. And, as the blood is divided into innumerable little streams, every part of it is acted upon by the air in the vesicles. Though the texture of the lungs is exceedingly delicate, and the separation between the air and the blood is almost as nothing, yet the blood is confined to its limits, even though it courses through these organs with great, rapidity, and it never mingles with the air except as a consequence of actual disease. 151. The quantity of carbonic acid gas discharged from the lungs in the course of twenty-four hours is very great. Many experiments have been tried and calculations made to ascertain its amount, and I am within bounds when I state, that there is at least three-quarters of a pound of charcoal in the carbonic acid thrown off from the lungs of a common-sized adult in the course of twenty-four hours. This gas is a deadly poison. When accumulated in a considerable amount, as when char- coal is burned in an open furnace in a close room, it may prove immediately destructive to life. And in the very prevalent neglect of ventilation, the frequent accumulation of this gas from the respiration must prove more or less injurious to the health. Whenever the proper amount of oxygen gas is with- held from the lungs, and carbonic acid takes its place, the quality of the blood is impaired from incompleteness in the change effected in the lungs, and the vigor of the body must in this way be lessened, to say nothing of the deleterious influ- ence of this gas upon the nervous system. Though the results are not immediate and palpable, great injury is continually done to the health of multitudes by the accumulation of this gas, in small close apartments, and in crowded assemblies. A congregation of twelve hundred people in two hours throw off from their lungs an amount of carbonic acid that contains seventy-five pounds of charcoal. And yet little pains is com- monly taken to carry off this vast quantity of poisonous gas, and replace it with pure air. 152. As so much oxygen is absorbed in the lungs of all ani- mals, and so much carbonic acid is thrown out from them, the inquiry arises how the air is replenished with oxygen, and is 104 HUMAN PHYSIOLOGY. Carbonic acid exhaled from the lungs of animals absorbed by plants.________ cleared of the carbonic acid which is thus so largely mixed with it. It is found that this is done, to a great extent at least, by the leaves of plants. The process which goes on in these lungs, as they may be called, of the plants, is quite the reverse of that which is going on in the lungs of animals. The carbon of the car- bonic acid which is thrown off from the lungs of animals is ab- sorbed by the leaves of plants, and the leaves replenish the air with the oxygen, which is so constantly and abundantly ab- sorbed in the lungs of the animal creation. Thus, the animal and vegetable kingdoms are sources of supply to each other. But it may be thought that there would be apt to be a surplus of oxygen in the atmosphere in warm climates, where the vege- tation is so luxuriant; while, on the other hand, there would be an accumulation of carbonic acid gas in the colder regions. This would be so, if the air were not so movable that the equi- librium is readily secured in either case. 153. It is an interesting fact, that the presence of light is necessary to the process which I have described as going on in the leaves of plants. Each leaf may be considered as a labor- atory, and the light as the chief agent in effecting the chemical changes that occur in it. And it is found that no artificial light can do the work. It is only the light of the sun that is competent to this chemistry. And as these innumerable labor- atories are everywhere at work, absorbing the carbon and ex- haling the oxygen, to purify the air rendered noxious by the laboratories of the animal creation, we must confess it to be a mystery as to how the chemistry of the lungs of animals, and that of the leaves of plants should be kept so nicely balanced. The balance is so strictly maintained, that the chemical composition of the air is always found to be almost exactly the same. 154. The heat of the body is maintained by the union which takes place in the capillaries between the carbon and hydrogen of the system, and the oxygen which is introduced into the blood through the lungs. It is a process analogous to combustion. When carbon or charcoal is burned in a ves- sel containing air, the oxygen disappears, for it unites with the carbon, and carbonic acid gas, therefore, appears in its place. The same union occurs in this case between carbon and oxygen, as we find occuring in the capillaries. A sort of com- bustion, then, is going on in every part of our bodies. And, as heat is evolved in the one case, so it is in the other. The same can be said of the burning of hydrogen and oxygen together. Heat is caused by the union thus produced between them, and RESPIRATION. 105 Animal heat. Produced by a sort of combustion. Three sources of fuel. so it is when they unite in the body. The water which is ex- haled from the lungs comes from this union of oxygen and hydrogen. It was formerly supposed that the union between the oxygen and the carbon and hydrogen takes place in the lungs, and that the heat is made there, and then is distributed over the whole system. But it was objected to this supposi- tion, that it made the lungs a sort of furnace for the rest of the body, and that, if the supposition were correct, there ought to be a much higher degree of heat in these organs than any- where else, which is not the case. Ingenious theories were broached to get over this difficulty; but it was at length dis- covered that the union between the oxygen and the carbon and hydrogen occurs in the capillaries of the body, instead of the lungs, and that the combustion, therefore, that produces the heat is everywhere, instead of being in one locality. 155. The fuel for this combustion comes from three sources. One of these is the waste of the tissues. This furnishes a con- siderable amount of the carbon and hydrogen for the union with the oxygen, in all animals that are subjected, from their activity, to much wear and tear of the system. I barely al- lude to this now, and shall enlarge upon it soon. Another source of the fuel for combustion is food. The oily, sugary, and starchy kinds of food are devoted in a great measure to this particular purpose. These furnish a sort of floating fuel, as we may express it, which is carried about in the blood. Hence, we see, that our diet must necessarily be varied accord- ing to the weather and the climate. In cold weather, the heat of the body is more rapidly abstracted than in warm weather, and, therefore, we need then more of that food which affords a supply of carbon and hydrogen. And so as to climate. The enormous quantity of oily food often consumed by inhabitants of very cold climates is used up by being burned, as we may say, in the capillaries to keep up the animal heat. Of course, keeping the body warm by fire and clothing relieves from the necessity of taking any large quantities of fuel-making food. Still under the most favorable circumstances in this respect, there is a need of variation in diet to suit the weather and the climate and we make this variation for the most part instinct- ively. Indeed there is a marked provision in nature for it. I will mention but a single example of this provision. While there is a laro-e amount of fat in the bears and seals and whales which afford food for the Esquimaux and Greenlander, there is very little in the animals which furnish a part of the diet of the 106 HUMAN PHYSIOLOGY. Animal heat differs in cold and wurin-bloocled animals. Why. inhabitants of tropical climates. Another source, still, of ani- mal heat is the store of fat which is laid up in the body. One design of this accumulation of fat in different parts of the body seems to be to provide for the heat when other sources fail. Thus, when disease destroys the appetite, and thus cuts off the supply of food, the fat wastes away, or rather is burned up, to keep up the temperature of the body. The fat is the great means of maintaining the requisite temperature when hi- bernating animals become torpid for the winter. They become very fat in the autumn, before crawling into their winter quar- ters, and in the spring they come out very lean, their fat having been consumed in keeping up the low degree of temperature re- quired during this time. 156. As the amount of heat produced, when charcoal is burned in air, or when oxygen and hydrogen are burned together, depends upon the quantities of carbon and hydrogen that unite with the oxygen, so, also, the degree of animal heat depends upon the quantities of carbon and hydrogen that unite with the oxygen in the capillaries. This may be illustrated by referring to the effects of exercise on the heat of the body. When the circulation is quickened by exercise, the blood passes more rapidly than usual through the lungs, the respiration is consequently quickened, more air is introduced into the lungs, and therefore oxygen is more rapidly absorbed by the blood. At the same time, the action of the muscles effects a waste in their structure by the wear and tear, so that more carbon and hydrogen are ready to be released to be united with the in- creased oxygen. Hence comes the heat produced by exercise. So, too, those animals which are the most active, ordinarily have the most animal heat, and have the most extensive respi- ratory apparatus, so that there may be a free supply of absorbed oxygen to unite with the carbon and hydrogen of the changing tissues. It is in birds and insects that this union takes place most largely, and in them, therefore, the respiratory apparatus is very largely developed. This is to be attributed to their mus- cular activity, which produces so much waste matter that must be removed from the system. Cold-blooded animals, on the other hand, are very inactive. There is not, therefore, much wear and tear of the tissues. There is comparatively little waste, therefore, to be thrown off. And so but little oxygen needs to be introduced into the lungs, and consequently little heat is generated. To realize fully the contrast between the warm and the cold-blooded animals in these respects, observe, as RESPIRATION. 107 Uniformity of animal heat in the warm-blooded. Interesting experiments. the representative of the one class, a canary bird, and a frog as the representative of the other. The frog is generally quiet, and only now and then takes a leap or croaks; but the bird is ever in restless motion, and sings much of the time with all his might. The bird is warm with the heat generated by the constant union of oxygen with carbon and hydrogen in its capillaries ; but the frog is nearly as cold as the water in which he is immersed. The bird breathes rapidly, to let the oxygen of the air largely into his lungs ; but the frog scarcely seems to breathe at all, so scanty is the supply of oxygen which he needs. 157. Cold-blooded animals are very nearly of the same tem- perature with the substances that are around them; but warm- blooded animals have a certain degree of temperature, which they maintain with considerable uniformity under all variations of temperature in the atmosphere. This in man is about ninety- eight degrees of Fahrenheit. This, you observe, is above the temperature of the surrounding air, except in exceedingly hot weather. The human body is therefore always giving off heat. Indeed it is essential to comfort that it should part with con- siderable heat, f<5r any near approach of the atmosphere to ninety-eight degrees produces an uncomfortable sensation of heat. But the amount of heat which the human body can bear for a short time is much greater than the facts above alluded to would lead us to suppose. It was long taken for granted, that it could not safely bear, even for a short time, a heat much higher than that which is endured in hot climates. The truth on this subject was at length discovered by accident. Two Frenchmen were employed by government, in 1760, to devise some method of destroying an insect which infested the grain at that time. The result of their experiments was the discovery, that by subjecting the grain to a certain degree of heat in an oven the insect was destroyed, and the grain not injured. While they were trying their experiments, a girl offered to go into the oven and mark the height of the mer- cury in the thermometer. It stood at 260° ; and, after remain- ing there for ten minutes, which she found that she could do without any great inconvenience, she marked it at 288°, that is 76° above the boiling point of water. These facts led to the famous experiments of Dr. Fordyce and Sir Charles Blagden, in England. With wooden shoes, tied on with list, they went into a room in which the thermometer showed the air to be at 260°. Their watch chains were so hot that they could scarcely 108 HUMAN PHYSIOLOGY. Different degrees of torpor in hybernating animals. touch them, and eggs were roasted hard in twenty minutes, and beefsteak was cooked in thirty-three minutes. And yet the same air that produced these results was breathed by them with impunity, and it raised the heat of the body but very little. The air which was breathed out from the lungs was so much cooler than the air of the room, that it was refreshingly cool to the nostrils, and to the fingers as they blowed upon them. In such cases, the evil effects of the heat are prevented chiefly by the great amount of perspiration that occurs, the vaporization of this abstracting the heat, which would otherwise accumulate in the body and produce disastrous results. The exhalation from the lungs, also, has some influence. 158. In the state of hibernation, to which I have several times referred, the torpidity varies in degree in different ani- mals. In cold-blooded animals, respiration and circulation may cease altogether in this state. In them the movements of life are often, perhaps we may say generally, as fully suspended as they are in the seed that is kept from heat and moisture. They may be preserved in this state for a long time and yet revive. Serpents and frogs have been kept in an ice-house for three years, and then have been revived on being brought out into a warm atmosphere. In the warm-blooded animals that hiber- nate the torpidity is less deep than in those which are cold blooded. In them the respiration and the circulation become very slow, but never entirely cease. Indeed some species take food with them into their winter quarters, and occasionally wake up sufficiently to eat. But most of them are in a quiet, deep sleep, from which they do not arouse at all till the winter is past. In this state, as life is nearly, sometimes quite at a stand, there is no wear and tear, and therefore no change in the tissues, and so there is no need of the introduction of oxygen by the respiration. Dr. M. Hall, in his experiments and observations, found that the bat, when completely torpid, consumed no oxy- gen, and discharged no carbonic acid from the lungs, although its circulation was not entirely suspended. 159. The more active is the respiration of animals, the less able are they to bear a deprivation of air. A warm-blooded animal will die if it be under the water only a few minutes; but a cold-blooded animal can live under the water for some time, because it is not in so urgent need of oxygen. And, for the same reason, a warm-blooded animal, in a state of hiberna- tion, may be kept under water for a long time without destroy- ing life, although when in its active state it would die on being FORMATION AND REPAIR. 109 Formative vessels appended to the capillaries. kept under water for only a few minutes. And this suggests a probable explanation of those cases, in which individuals have been restored, after having been under the water longer than the usual time that suffices to destroy life in drowning. In such cases, the condition is not simply that of a drowned person. A blow, or the shock of body or mind, or both, may have in- duced a suspension of active vitality, like that which we see in the animal in a state of hybernation. The bare fact of immer- sion in the water may have but little or even nothing to do with the actual condition. Such a state of things is especially to be suspected in those cases in which the countenance does not exhibit the usual dark and full appearance of drowned per- sons. 160. I have thus shown the extensive play which the respi- ration has in the vital operations of the system. I have shown what the chemical changes are, which it effects directly in the lungs, and indirectly in the system. And you have seen how the animal heat is produced by these changes, and how unac- countably it is so regulated, that it seldom varies to any ex- tent from its fixed standard. But it is to be remembered that, while the lungs, and even the capillaries, everywhere are thus chemical laboratories, the nervous system exerts a constant influence upon this chemistry of the body. This is especially seen in regard to the production of heat, but it is true of the whole range of the chemical operations. The laboratories would all cease their work if their nervous connections were destroyed. CHAPTER VIII. FORMATION AND REPAIR. 161. The building and the repairing of the various struc- tures of the body are done by vessels appended to the capil- laries. The capillaries having received from the arteries the blood, the building material, the formative vessels select from it while it is in these capillaries, whatever they need for their purposes. The selection is made according to the tissue or structure to be formed. Those vessels which, for example, form r bone, select from the blood very different constituents from those which make nerve or muscle. 10 110 HUMAN PHYSIOLOGY. Selecting power of the formative vessels. Their concert of action. 162. It is wonderful that the blood can be formed from such a variety of food as is often taken into the stomach. But it is far more wonderful that from the blood can be made so many and such different structures. How different are the teeth from the gums which surround them ; and yet both are made from the blood. Observe, in some particular part of the body, how many different structures there are which are all made from the same common material. Take, for example, those which are in and around the eye. There are, the skin of the eyelids ; the eyelashes; the vascular lining on the inside of the lids; the cartilages of the lids; the firm, white coat of the eye, giving to the eyeball its firmness; the thin, transparent window in front, setting into the firm, white coat, like a watch-glass into the case ; the beautiful iris, a round moving curtain with a cen- tral opening; the lens behind this opening; the optic nerve ex- panded on the inside of the cavity of the eye; the muscles that move the eye, with their tendons; the tear-gland; the cushion of fat on which the eye reposes; the bone which forms the socket, &c. All these various textures are formed from the blood ; and the different workmen are as unerring in their se- lections from this common material, as if they were intelligent beings. Indeed, no ordinary intelligence could accomplish such a selection. It is effected, inscrutably to us, under the direction of an all-wise Intelligence, and by Almighty power. 163. But these builders of the body not only have the power of selecting their building materials from the blood, but they work in concert. Each company of builders work together in harmony, as if they were under intelligent leaders. And though different companies may be in close proximity, there is no disagreement nor interference. For example, the builders of a tooth and the builders of the gum around it, do not en- croach on each other; but each do their appropriate work with- in their assigned limits. Even when different structures are intermingled, as when tendon and muscle mingle together at their place of union, there is no confusion in the work of the two sets of laborers. In Fig. 48 you see the difference in struc- ture between the transparent cornea in the front part of the eye and the white coat, the sclerotic coat, into which the cornea is set like the crystal of a watch. It is represented as seen magnified 320 times. The dotted lines mark the place of union. The cornea, a, is a much more open structure, you ob- serve, than the sclerotic coat, b. The builders of these two structures, though some of them are in such near neighborhood FORMATION AND REPAIR. Ill Concert of action shown in producing different shapes. FIG. 48. never encroach on each other, but each set adheres strictly to its own kind of work. The sclerotica-makers never go to mak- ing the open work which you see in the cornea. If they should do so at any point there would be a little transparent window at that point in the white of the eye ; and if the cornea-makers should at any point make close work like that in the sclerotica, there would be a white spot in the cornea. 164. The concert of action which we observe in the different sets of formative vessels is to be looked at in another point of view. It is such that they give a definite and peculiar shape to the structure which they make. Each bone differs in shape from every other bone, each muscle from every other muscle; and so of other parts. There is very great variety of shape in the structures of the body ; and each shape can be determined only by a certain concert among the builders. That you may realize in some measure the extent of this variety,. observe again the numerous different textures which I have mentioned as making up the eye. Each of these has its own peculiar shape, and its definite limits. Its builders work after a fixed plan, and within fixed bounds. 165. This concert of action may be looked at in still another point of view. If the different structures in the body were made, as a crystal is, by layer after layer of particles deposited upon the outside, wonderful as the concert among the little builders would be in that case, it would not be any thing like as wonderful as it is now. In the growth, that is the construc- tion of any part, the addition is made by the formative vessels at every point of the part, and not upon the outside merely. As these builders are at work enlarging the part in the growth from infancy to childhood, they must so act in concert, as to 112 HUMAN PHYSIOLOGY. Change of action. The teeth. Tadpole and frog. preserve the same general form in the part during all the suc- cessive stages of growth. And, as all the different structures of the body enlarge together, there must be agreement between different sets ; else there would be encroachment and confusion. Thus in the growth of the tiny arm of infancy to the sturdy arm of manhood, each set of builders must during all this time keep within its proper limits, so that there may be just the right proportion, and the right position of bone, and muscle, and tendon, and ligament, and cellular membrane, and skin, and nail, &c., that make up the arm. 166. But this concert of action appears the most wonderful when a new action, or change of action is called for. In the transition from childhood to youth, for example, the builders of the apparatus of the voice, the larynx, all at once become unusually active in their work, and a great enlargement of this musical instrument, for such it is, takes place, so that it may now utter the grave notes of manhood. Soon, too, the beard- builders begin their new work upon the face. And during the period of childhood new operations have been continually insti- tuted among the builders of the teeth, as one tooth after another has made its appearance, and as the new set have re- placed the old. To produce in the enlarging jaw a new set of teeth to take the place of the smaller and less numerous first set, and to bring them out in a symmetrical arrangement, re- quire a very complicated series of operations. To effect each one of these, there must be concert of action among the forma- tive vessels; and there must be a most wonderful concert among the different sets and succession of builders, to make all these series of operations work out at length the general result. 167. This change of action in the formative vessels is strikingly exemplified in some animals. I refer to those that so entirely change their forms during the period of their exist- ence. I will give two examples. The first is the common frog. He is at first what is termed a tadpole, and goes through many successive changes to become a complete frog. These changes are represented in the following figures. The relative sizes are not preserved, the tadpole state being represented re- latively much too large, for the purpose of showing more clearly the development of the legs. The young tadpole is represented in Fig. 49. It has a large head and body, and a long flat tail by which it swims easily. There are no promi- nences to indicate the putting forth of any thing like limbs. It FORMATION AND REPAIR. 113 Change of action in the silk-worm. Concert preserved in these cases. FIG. 50. has gills, which are loose fringes on each side of the head. These gills after a time disappear, and it has another set of gills arranged under a fold of skin very much like the gills of a fish. The form is then-as in Fig. 50. The next change is this. The hind legs begin to grow out as seen in Fig. 51. Next, the fore legs appear as seen in Fig. 52. The tail is still very large. This now gradually disappears while the legs grow as represented in Fig. 53. In Fig. 54, representing the perfect frog, the tail has entirely disappeared. With these exterior changes interior ones have been going on also. The animal, which was at the first a real fish, breathing with gills and swimming in water, has lost its gills, and has now a pair of lun