CLASS-BOOK OF PHYSIOLOGY; FOR THE USE OF SCHOOLS AND FAMILIES. COMPRISING THE STRUCTURE AND FUNCTIONS OF THE ORGANS OF MM, ILLUSTRATED BY COMPARATIVE REFERENCE TO THOSE OFy^NFERIOR ANIMALS, BY B. N. COMINGS, M. D. AUTHOR OF PRINCIPLES OF PHYSIOLOGY, OUTLINES OF PHYSIOLOOY, THE PRESERVATION OF HEALTH, ETC., ETC. WITH TWENTY-FOUR PLATES, AND NUMEROUS ENGRAVINGS O NW OOP, COMPRISING IN ALL ABOVE TWO HUNDKEQyflGUA^S,'i'L •'.< .<;'"•• NEW-YOEK: £ '- l/ D. APPLETON & CO. 200 BROADWAY."' 18.13. ENTERED, ACCORDING TO ACT OF CONGRESS, IN THE YEAR 1852, BY B. N. COMINGS, IN THE CLERK'S OFFICE OF THE DISTRICT COURT OF CONNECTICUT. FOUNDRY OF S. ANDRUS AND SON, • HARTFORD. W. C. Armstrong, Typographer. PREFACE. The '• Principles of Physiology" which has been so highly commended by all literary men, who have examined its merits, although well adapted to more advanced institutions, has been thought too large and expensive for the major portion of our schools. To meet the demand for a lower priced book, and comply with the desires of numerous friends of edu- cation, the Class-Book of Physiology is now offered to the public. The unanimous verdict of teachers in favor of the general plan of he "Principles of Physiology," has induced the author to pursue the same method of illustration, and, if possible, bring to a higher perfection the original plan of that work. It has been his constant study in this treatise to explain and illustrate to the student, fully and clearly, as many of the most practical principles of Physiology, as could be included in the moderate limits of the present volume. That Human Physiology can be made more easy of comprehension— more profitable, and more attractive to the young, by appropriate refer- ences to the Comparative Physiology of the inferior animals, than by any other method—is an established fact in the mind of the author, though he must leave it for others to judge of the success with which this plan has been developed. Physiology owes much of its present eminence, as a deeply interest- ing and progressive science, to observations made on the structure and functions of the same organs in the lower orders of animals as are found to exist in man. Take from Human Physiology the light derived from Comparative Physiology, and it would become what it was centu- ries ago, an obscure and comparatively uninteresting science. The young universally feel a deep interest in the animated beings which they see around them. The horse, the ox, the dog, and birds and insects, are all objects of admiration and wonder to the philosophic mind of youth; and when they learn that each possesses organs approximating more or less in structure to their own, their sympathies are enlisted, they feel a deeper interest in their own organization, acquire more noble and exalted views of the position they occupy in the scale of being, and learn to appreciate more truly the evidences of an all-wise design in the works of their Creator. 4 PREFACE. Physiology cannot but be considered, by every intelligent and reflect- ing mind, a deeply interesting and practically important study. It makes us acquainted with the structure and uses of the organs of life, and the laws by which they may be preserved in health and vigor. When a knowledge of the principles of Physiology shall be diffused through all classes of society, the sum-total of human happiness will be greatly increased, and more permanent enjoyment of health, and more exalted exercise of all the physical and intellectual .powers, will be secured to each individual. This science has peculiar claims to the attention of every teacher who would be successful in his profession. It presents, in their true light, those facts which lie at the foundation of mental development, and suggests the best means of promoting intellectual growth. For the convenience of those who have not had sufficient experience to dispense with questions, they have been put at the bottom of each page, but there is left ample scope for teachers to propose many more. For those who may be advanced in study, or who may review the book, a description of the figures and plates are recommended as an useful exercise. Carpenter, Roget, Wilson, Pereira, Bourgery, Aitken, Kirkes and t Paget, Horner, Hussall, Aggassiz, Wyman, Chambers, Griscum, and vari- ous other authors, have been consulted in the preparation of this work. The author is under obligations to the publisher of "Principles of Physiology" for permission to use the figures and such portions of the text of that work as he found suited to his purpose. New Britain, May, 1853. CONTENTS. PAOIC INTRODUCTION,............................................................ 9 CHARACTERISTICS OF ORGANIC AND INORGANIC MATTER,............. 9 DEFINITIONS,............................................................... 10 CHAPTER I. CHARACTERISTICS OF PLANTS AND ANIMALS,.......................... 15 CHAPTER II. STRUCTURE AND COMPOSITION OF ANIMAL BODIES,.................... 19 CHAPTER III. THE BLOOD,................................................................ 27 CIRCULATION OF THE BLOOD,............................................ 29 CHAPTER IV. RESPIRATION,.............................................................. 47 CHAPTER V. ANIMAL HEAT,............................................................. 53 CHAPTER VI. DIGESTION,................................................................. 67 MASTICATION,............................................................... 67 INSALIVATION,............................................................. 72 CHAPTER VII. ABSORPTION,............................................................... 93 CHAPTER VIII. NUTRITION,................................................................. 98 CHAPTER IX. SECRETION,.................................................................. 101 CHAPTER X. THE SKIN,..........................•......................................... 109 6 CONTENTS. CHAPTER XI. paok THE NERVOUS SYSTEM,.................................................... 119 NERVOUS SYSTEM OF MAN,................................................ 12, SPINAL CORD,............................................................... 12a MEDULLA OBLONGATA,.................................................... 131 THE BRAIN,.................................................................. n'2 FUNCTIONS OF THE CEREBELLUM,......................................... 141 FUNCTIONS OF THE CEREBRUM,........................................... 1J" CONNEXION OF THE MIND WITH THE BODY,.............................. 149 CHAPTER XII. THE SENSES,................................................................ 153 SENSE OF TOUCH,...................................................... 153 SENSE OF TASTE,....................................................... 155 SENSE OF SMELL,...................................................... 161 SENSE OF HEARING,................................................... 161 SENSE OF SIGHT,....................................................... 170 CHAPTER XIII. ANIMAL MOTION,.......................................................... 192 THE MUSCLES,.............................................................. 219 CHAPTER XIV. THE VOICE,.................................................................. 234 CHAPTER XV. THE HUMAN FORM,....................................................... 239 SYNOPSIS OF LAWS OF HEALTH,.......................................... 251 INDEX AND GLOSSARY,........ ........................................ 253 LIST OF FIGURES. FIQ piOg F1Q l'AQ.T. 1. Parent Cells, 20 36. Single Gland of Small Intestine, . 90 2. Fasciculi and Fibres of Cellular Tissue, 21 37. Lymphatics, .... 94 38. Diagram of Mucous Membrane, 95 4. Corpuscles of Human Blood. 28 39. Appearance of Lymphatics, . 95 5. Dorsal Vessel of the Spider, 30 40. Section of Kidney, 104 6. Circulation in Crustacea, 30 41. Structure of Parotid Gland, . 104 7. Circulation in Fishes, , 31 42. Nervous System of Ascidia, 121 8. Circulation in Reptiles,. 31 43. Nervous System of Pecten, . 122 9. Circulation in Man, . 32 44. Nervous System of an Insect, . 123 10. Diagram of the Pericardium, 32 45. Nervous System of Sphinx, ., 124 11. Section of Right Side of the Heart 33 46. Brain of a Cod—of a Shark, 126 12. Valves of a Vein, .... 36 47. Brain of a Bird, .... 12G 13. Capillaries in a Frog's Foot, 36 48. Portion of Spinal Cord, 129 48 49. Skull of European, 144 15. The Serpula, .... 49 50. Skull of Negro, .... 144 16. Respiratory Apparatus of Insect, . 50 51. Capricorn Beetle, .... 153 17. Interior of a Snail, . 51 52. Vertical Section of the Nasal Cavily 162 18. Lungs of an Ostrich, 52 53. Bones of the Ear, .... 167 19. Air Cell,..... 52 54. Cavity of the Tympanum, . 167 20. The Larynx, Bronchial Tubes, &c. 53 55. Interior of the Eye, 181 21. Human Thorax, 54 56. Section of the Eye, . 181 22. Development of Teeth, . 69 57. Refraction of Rays of Light, 184 23. Dental Capsule, 69 58. Image on the Retina, 189 23'. Human Teeth, .... 69 59. Bones in the Foot and Leg of a Deer ,213 24. Section of a Human Tooth, 70 60. The same of a Horse, 213 25. Jaw and Teeth of Rabbit, . 70 61. Bones of the Arm,. 220 26. Teeth of Herbivorous Animal, 71 62. Bones and Muscles of the Humerus ,225 27. Teeth of Carnivorous Animal, 71 63. Tail-Fin of a Whale, . 226 28. Teeth of Insect-eating Animal, 71 233 29. Teeth of Frugivorous Animal, 71 65. Larynx of a species of Duck, 235 30. Teeth of a Rattle-Snake, . 71 66. Larynx of a Rook, . 235 31. Section of Mouth and Throat, 73 67. Vertical Section of the same, 235 32. Stomach of the Sheep, . 76 68. Human Larynx, viewed sideways, 236 33. Section of the Stomach, 80 69. Vertical Section of the Larynx, . 236 34. Mucous Membrane, . ... 81 70. Front View of the Larynx, 236 35. Tubular Follicle of the Stomach, . 82 71. Bird's-eye View of the Human do. 237 LIST OF PLATES. PLATES. PAGB. I. Comparative View of the Organs of Circulation,.....13 II. General View of the Circulating Apparatus of Man, .... 17 III. Organs of Circulation,..........25 IV. Organs of Circulation.—Heart and Lungs,......39 V. Organs of Respiration, ,.........45 VI. Organs of Respiration, ,.....57 VII. Organs of Digestion,...........77 VIII. Organs of Digestion,..........87 IX. • The Skin,.............109 X. The Skin,.............117 XI. The Brain and Spinal Cord,.........139 XII. The Nerves,............147 XIII. Organs of Sense,............159 XIV. Organs of Vision.—The Eye,.........173 XV. Organs of Vision.—The Eye,.........179 XVI. Organs of Vision.—The Eye,......... 187 XVII. Organs of Motion.—The Bones.—Anterior View,.....195 XVIII. Organs of Motion.—The Bones.—Posterior View, .... 201 XIX. The Extremities,............209 XX. Organs of Motion.—The Bones.—Comparative View, .... 217 XXI. Organs of Motion.—The Muscles.—Lateral View,.....223 XXII. Organs of Motion.—The Muscles.—Posterior View, .... 231 XXIII. The Human Form, in various Positions,.......240 XXIV. The Human Form in various Positions, and Public School Play-grounda, 253 CLASS-BOOK OF PHYSIOLOGY. INTRODUCTION. 1. All bodies in which evidence of life has been observed, possess certain parts or organs, which are essen- tial to the existence of the individual. A plant or an animal exists by means of its appropriate organs, and the matter of which they are composed is therefore called organized or organic matter. 2. Minerals exhibit no signs of life, but they continue to exist without any distinction of parts or organs, and belong to the kingdom of unorganized or inorganic matter. A piece of quartz, for instance, may be broken into any number of fragments, each of which will retain all the characteristics of the original mass. 3. Minerals may grow or increase in size by the super- addition of new particles to their surface; but they are not nourished, like organized beings, by interposing new par- ticles between those which already form their substance. 4. In plants and animals, each part is essential to the completeness of the individual being, and no part or organ can maintain an independent existence. If a rose be cut from its stem, or an arm be amputated from the body, it immediately dies, and loses its original characteristics. 5. A plant is dependent upon its roots and its' leaves for vitality, and an animal cannot live without its heart, its lungs, and its stomach. What are essential to the existence of all living bodies'? What is the matter of which such bodies are composed called? What is the matter found in minerals called ? Do minerals growl How? Are they nourished? What is essentia! to the completeness of a plant or animal? Can any organ main- tain an independent existence? Upon what parts is a plant dependent for its vitality? Upon what parts is an animal dependent? 10 CLASS-BOOK OF PHYSIOLOGY. 6. There are two conditions in which we may study or- ganized matter: namely, as living beings and as dead bodies. 7. The science of Physiology is derived from the first method, and the science of Anatomy from the second. 8. Physiology makes us acquainted with the uses to which the different parts are subservient, and the laws by which they are governed. 9. Anatomy teaches the number, size, situation, form, texture, and composition of the various parts, with their relations to each other. 10. Anatomy and Physiology, in their most extended use, apply to all organized beings, though they are naturally divided into the several branches of Vegetable Anatomy and Physiology, and Animal Anatomy and Physiology. 11. Animal Anatomy and Physiology are again divided into Comparative Anatomy and Physiology, and Human Anatomy and Physiology. 12. Comparative Anatomy and Physiology are devoted to the lower orders of animals. 13. Human Anatomy and Physiology are limited in their application to man. 14. The divisions of Physiology into Comparative and Human are usually made as a matter of convenience in study, and not because the Physiology of man is peculiar to him alone: for the same organs which exist in man, may be found in greater or less perfection in nearly all the lower orders of organized beings. 15. Comparative and Human Physiology naturally illus- trate each other, and thereby become more interesting and more easily comprehended. In what conditions may we study organized matter? What science is derived from the first, and what from the second method? With what does Physiology make us acquainted? What does Anatomy teach? How are Anatomy and Physiology naturally divided? How are animal Anatomy and Physiology divided? To what orders of animals are Comparative Anato- my and Physiology devoted? Why is Physiology divided into Comparative and Human? \ PLATE I. COMPARATIVE VIEW OF THE ORGANS OF CIRCULATION. Fiourb 1.---Circulation in the Insect—a, The dorsal vessel, divided into valvular partitions, by the successive contraction of which, the blood is propelled forward, J, 4, Canals which carry the blood to the head, c, c, Canals passing backward for the supply of the body, and returning the blood to the posterior end of the dorsal vessel.—The course of the circulation is indicated by the direction of the arrows. Fiourk 2.---Circulating Apparatus of the Lobster.—In the lobster and crab, the heart has but a single cavily, and the veins are indistinct, consisting merely of irregular channels excavated in the tissues, a, The heart; b, c, Arteries which go to the head and to the antenna or feelers; d, The hepatic artery, or artery of the liver; e, /, Arteries which sup- ply the thorax and abdomen.----After the blood has been propelled through these arteries by the heart, it passes into the great vein, g, g, from all parts of the body. Thence it passes to the gills, h, where it is exposed to the action of the air, and is then returned to the heart by the branchial veins, i, which correspond to the pulmonary veins of Man. Figure 3.---Sectional View of the Circulating Apparatus of the Lobster.—a, The heart. o, b, Venous sinuses, c, c, Branchial arteries, d, d, The gills or branchiae, e, e, Branchiul veins, terminating in the heart. Fiqurz 4.----Circulating Apparatus of Fish.—a, The auricle, having a single cavity, to which the blood is sent from all parts of the body. 4, The ventricle, which receives the blood from the auricle, and propels it to the arterial bulb, e, into the branchial artery, d. The branchial artery is sub-divided into the arteries of the gills, e, in which the blood is aerated. /, /, The dorsal artery, or aorta, which receives the aerated blood from the gills, and distributes it to all parts of the body, g, The vena cava, or great hollow vein, which conveys the blood back again to the auricle. A, Vena porta?, that branch of tho vena cava which conveys the blood from the abdominal organs, i, The intestine, k, The kidneys. Figure 5.---Circulating Apparatus of Lizard.—a, Left auricle, b, Right auricle.— One of these receives the venous blood from the system, and the other receives the arte- rializcd blood from the lungs, c, The single ventricle, which'receives the blood from both auricles, and transmits it partly into the lungs and partly into the aorta, d, d, Arches of the aorta, e, Carotid artery, which distributes the blood to the head, f, Pulmonary vein, which conveys the blood from the lungs to the heart, g, Brachial artery, which goes to the fore-legs. A, A, Pulmonary artery, in which the blood is submitted to tho inlluenct! of air in the lungs, t, The lungs, j, The stomach, k, Vena portae. /, Intestines, m. Ventral aorta, or that portion,of the aorta contained in the abdomen, n, Kidneys, o, Liver and vena purtae. p, Inferior vena cava, which conveys the blood from all the lower parts of the body to the heart, j, Superior vena cara, through which the blood of the upper parts of the body is sent to the heart. Figure C.---Heart of Tortoise.—a, Right auricle, 4, Single Ventricle, c, Left auri- cle., d, d, Pulmonary artery, e, e, Pulmonary vein. /, Vena cava, g, Right aorta. A Left nnrtn. PI 1 PLANTS AND ANIMALS. 15 CHAPTER I. CHARACTERISTICS OF PLANTS AND ANIMALS. 16. Plants and animals are distinguished from alf unor- ganized bodies by the process of nutrition and the property of reproducing their kind. Plants are nourished by the inorganic elements found in the earth and air around them. The materials of their growth are received in the form of a liquid or a gas, already prepared for their use. 17. Animals are nourished by the organic materials of vegetables or of other animals. Animals always possess a stomach or a digestive cavity, in which their food is received, to undergo a process of preparation before it can be absorbed into their tissues. 18. Sensation and voluntary motion are peculiar to animals alone, and are therefore called animal functions. 19. Plants and animals both have a limited period of existence, which varies with every species. In some, this period is confined to a single day; in many plants, to a single summer: while some animals live more than a cen- tury, and some trees—as the oak and olive—are supposed to live a thousand years. 20. The natural age of man is usually estimated at about seventy years, though instances occasionally occur of indi- viduals who survive an hundred or more years. The duration of man's life is less uniform than that of any other class of animals. In some communities, the average age at death is only ten or twelve years; in others, it is nearly forty. In all cases, individuals and communities are long-lived just in proportion as their localities, occu- How are plants and animals distinguished from unorganized bodies? How are plants nourished ? How are animals nourished ? What do animals Hiwaya possess? What functions are peculiar to animals alone? What is snid of the period of existence of plants and animals? What is the natural age of man? How does it vary in different communities? In proportion to what are com munities and.individuals long-lived? 2 16 CLASS-BOOK OF PHYSIOLOGY. pations, and habits of life are favorable to health; for health and life are blessings lent to man only as conditions of obedience to the laws of his organization PLATE II. GENERAL VIEW OF THE CIRCULATING APPARATUS OF MAN. The course and relative positions of the principal arteries and veins of the Systemic circulation are shown in this plate. The arteries commence from the great arterial trunk called the aorta, and their branches are distributed to all parts of the system. The venous branches, which accompany the arteries, unite into two great veins, the superior and infe- rior vena cava, which convey the blood back to the heart. a, The left ventricle of the heart, b, The right auricle, c, The superior vena cava, d, The root of the pulmonary artery, e, e, The aorta, which is seen arching backward over the heart, and passing downward into the abdomen, where it divides into its two great branches, the iliac arteries, through which the blood passes to the lower extremities. /, The inferior vena cava, which accompanies the descending aorta and its branches, and returns the blood from the lower extremities. The dotted lines represent the outlines of the kidneys. PRINCIPAL DIVISIONS OF THE AORTA AND VENA CAVA. It should be remembered that most of the branches which spring from the great artery and vein, are double—that is, each right branch has a corresponding one at the left side —so that there are, for instance, the right and the left carotid arteries, the right and the left jugular veins, &c. From the arch of the aorta are sent off those arteries which are distributed to the head and arms. The principal ones among these are named as follows: g, The carotid artery, which nscends in the side of the neck, and divides into the tem- poral artery, A, which is distributed in the temple, and the facial artery, i, which supplies the face; and also sends a branch, called the internal carotid, to the parts within the skull. j, The sub-clavian artery, lying beneath the clavicle or collar-bone. That part of the continuation of this artery which passes through the axilla or arm-pit, is called the axillary artery, k ; that which lies in the upper arm, the brachial artery, /; and in the fore-arm it divides into the radial and ulnar arteries, m, n, which are distributed to the hand and fingers in the manner indicated in the plate. The principal branches of tho descending aorta are named as follows. The iliac artery, o, which, on passing into the thigh, becomes the femoral artery, p, and in the leg divides into the tibial and peroneal arteries, q, r, which form numerous brunches for the supply of the leg and foot. Before dividing into the iliac arteries, the descending aorta gives off several important branches; as the coeliac artery, from which the stomach and liver are supplied ; the renal artery, which goes to the kidneys, and the mesenteric artery, to the intestine's • besides many other sub-divisious in various parts of its course. The branches of the vena cava generally accompany those of the aorta in their distribu- tion, as shown in the figure, and are often called by the same names. Tho principal divisions of the ascending vena cava are:-The jugular vein, s, which accompanies tho carotid artery. The sub-clavian vein, t, which accompanies the artery of tho same name and receives tho blood from the arm and hand. ' ' The inferior vena cava, like the aorta, divides into two great branches, the iliac veins « the sub-divisions of which accompany those of the arteries, and are called by the same names. The manner in which the superficial veins ramify and anastomose with each othP, is shown on the upper and lower extremity of the left side. OF ANIMAL BODIES. 19 CHAPTER II. STRUCTURE AND COMPOSITION OF ANIMAL BODIES. 21. The component substances of animal bodies may be divided into fluids and solids: 22. The fluids are found mostly in the form of chyme, chyle, lymph, and blood. About thirty pounds of fluid can be drawn directly from a man who weighs one hun- dred and fifty pounds. By exposure to a process of evap- oration, the body may then be reduced to twelve or fifteen. Perfectly dry mummies are sometimes found to weigh only seven or eight. Hence the fluids may be said to constitute by far the largest proportion of the body. 23. The fluids'vary exceedingly at different periods of life. In youth, they are very abundant, making the form plump and round. In old age, they are greatly dimin- ished, leaving the form shrunken and wrinkled. 24. The fluids contain in solution the materials for the formation of the solid tissues, and are also the medium through which all the waste particles are carried out of the system. The fluids and solids, being alternately con- verted into each other, do not differ essentially in their chemical elements. 25. The human body has been found, by chemical analysis, to contain the following elementary substances: Oxygen, hydrogen, nitrogen, carbon, sulphur, phosphorus, silicon, chlorine, fluorine, iron, and sometimes mangane- sium, aluminum, and copper. 26. The first four are most constant and most abundant, and are named essential elements. 27. The solid portions of the body are called tissues. How are the substances which compose animal bodies divided ? How many pounds of fluid may be drawn from the body of a man weighing one hundred and fifty pounds? How can the weight of the body be farther reduced? When are the fluids most abundant and when least abundant? How many elementary substances are there in the human body? WhicrFare the foui essential elements? What are the solid portions of the body called? 20 CLASS-BOOK OF PHYSIOLOGY 28. The tissues, in their primary formations, are made up of granules, nuclei, cells, filaments, and fibres. 29. Granules are particles of various sizes, from immea- surable minuteness to the ten-thousandth of an inch in diameter. Granules are found floating in milk, chyle, and other animal fluids, and imbedded in most of the tissues. 30. Nuclei are the germ and cen- tre around which cells are formed. (Fig. 1.) 31. Cells are minute bubbles, vesi- cles or scales. Their natural shape is oval or spheroidal, but they often become flattened or many-sided by pressure, as may be illustrated by ng. i.-parent cells.— filling a vial with a strong solution a, a, Parent cells. n • , j ,-\ b, b, secondary cells. of soap m water, and then inverting c, e, Nuclei. ^ whe]Q if. ^ be fi]]ed ^^ bubbles> each one of which will assume a form in accordance with the position it occupies. The cells perform a highly im- portant process in all animal and vegetable structures. The solid portions of plants, and the tissues of animal bodies, are formed directly by the deposition of cells, or by the indirect elaboration of their fluid contents. They are concerned, not only in the functions of nutrition, in the development and restoration of parts, but in absorp- tion and secretion. They are developed into tissues in various ways. In some, the cell-membranes become elon- gated, and are folded and divided into threads or filaments of exceeding fineness. 32. Tubules, or little tubes, are several cells elongated, and placed end to end—the partitions being removed. 33. Filaments, or fibrils, are exceedingly delicate threads, composed of minute particles, usually arranged in parallel bundles or fasciculi. {Fig. 2.) Of what are the tissues made up? What are granules? Where are gran- ules found ? ^Vhat are cells? How are cells developed into tissues? Whal are tubules? What are filaments? OF ANIMAL BODIES. 21 Fig. 2.—Fasciculi and fibres of cellular tissue. A, White fibrous element, with cell-nuclei visible in it B, Yellow fibrous element, showing its branching fibrils. C, Finer fibrils of the yellow element. 34. Fibres are larger than fibrils, but are similar in other respects. 35. A tissue is the union or interlacement of one or more of these primary structures. 36. An organ fs an instrument composed of tissues, and designed for action. Its action is called its function or use. Thus, the liver is an organ, and the secretion of bile its function. 37. An apparatus consists of a number of different organs, arranged for the performance of some one office. The teeth, mouth, stomach, intestines, &c, belong to the digestive apparatus. 38. A system is a connected series of similar parts, such as the muscular or the nervous system. 39. The number of tissues which make up the different organs of animals is variously stated, according to the minuteness of description which different anatomists adopt. What are fibres? What is a tissue?—an organ? What is an apparatUB? —a system ? 2* 22 CLASS-BOOK OF PHYSIOLOGY. ^.3._A magnified representation of a portion ln futures of the ribs, the of areolar tissue. external air passes from the lung into this tissue, and continues inflating it till the whole body becomes enormously distended with air, caus- ing suffocation and death. In the progress of disease, the watery portions of the blood are sometimes effused into this membrane, causing dropsy. When the finger is pressed on a dropsical limb, a hollow or depression is pro- duced by the forcing out of the fluid from this tissue at that particular point. 41. The cellular tissue is found in every part of the sys- tem, except in compact portions of bone, teeth, and cartilage. Its chief use seems to be to connect together organs and parts of organs which require a certain degree of motion on each other. It possesses great power of extensibility and elasticity. 42. Yarious names have been assigned to the cellular membrane, corresponding to the different positions in which it is found. When inclosing those organs not exposed to What tissue is the primary form of all others? How is the cellular tissue formed ? How is the cellular tissue sometimes inflated ? Where is the cellular tissue found? Wha't is its chief use? What different names have been given to the cellular tissue ? OF ANIMAL BODIES. 23 the air, it receives the name of serous membrane, from a fluid secreted in*it, called serum. In the lining of the respiratory passages and of the alimentary canal, it is called mucous membrane, from a secretion of mucous which is poured out from numerous glands beneath its surface. Where it forms a covering for the body, it is known as the dermoid membrane, or skin. PLATE III. ORGANS OF CIRCULATION. Figure 1.----Front View of the Heart.—a, Right auricle, b, Right ventricle, c, Left auricle, d, Left ventricle, e, Aorta. /, Pulmonary artery, g, Superior vena cava. A, A, Coronary veins and arteries—the nutrient vessels of the heart. Fioure 2.----Section of the Heart, showing its Cavities and Partitions or Septa.—a, Right auricle, b, Right ventricle, c, Left auricle, d, Left ventricle, e, Aorta. /, Pul- monary veins, g, Superior vena cava, j, Right pulmonary veins, k, Left pulmonary veins. Figure 3.----Ideal View of the Circulation.—a, Right auricle. b\ Tho entrance of the Buperior vena cava, c, The inferior vena cava, d, Tricuspid valves, e, Right ventricle. /, Pulmonary artery, g, g, Branches of pulmonary artery. A, A, Capillary vessels of tho lungs, i, Pulmonary veins, j, Left auricle. A, Bicuspid valve. I, Left ventricle, m, Arch of the aorta, n, n, Arteries which distribute the blood to the head and upper ex- tremities, o, Descending aorta, p, p, Capillary vessels of the systemic circulation. Figure 4.----Capillaries.—This figure represents a highly magnified view of the ar- rangement of the capillaries, between the branches of the arteries and veins, as found in an intestinal villus, a, Arteries, b, Veins. Hartford, Conn THE BLOOD. 27 CHAPTER III. THE BLOOD. 43. Ix all organized beings, the process of nutrition is carried on by means of a circulati ng fluid. In plants, this fluid is called sap; in animals, it is called blood. 44. The blood of insects is white or colorless. In fishes, it is red in the gills, heart, and liver; but nearly colorless in the main bulk of the body. In the mammalia, birds, and reptiles, it is of a dark purple color when drawn from a vein, and of a bright scarlet when it comes from an artery. It emits an odor peculiar to the animal from which it is taken. 45. In a few minutes after blood is taken from a living animal, it begins to coagulate, and become solid, like a soft jelly. If allowed to stand for a few hours, the clot will be found diminished in size, but firmer than before, and floating in the midst of a yellowish fluid, called serum. 46. The serum, or liquid, which remains after coagula- tion, is composed principally of albumen and water. 47. Albumen forms a very large proportion of the brain, the spinal cord, and nerves. It is perfectly colorless when pure, and is coagulated or hardened by either heat or acid. An example of it may be found in the white of an egg. 48. The ivater of the blood is always one of its most important constituents, and always forms by far the greater proportion of its bulk—one thousand parts of the blood containing from seven to eight hundred parts of water. The amount given off in perspiration, the state of the at- mosphere, the fluid drank, and many other circumstances, How is the process of nutrition carried on in organized beings? What is this fluid called in plants?—in animals? What is the color of the blood in insects?—in the mammalia, birds and reptiles? How does the blood coagu- late? Of what is the serum composed? Of what tissues does albumen form a large proportion ? What are some of the characteristics of albumen ? What proportion of the blood is water? What circumstances influence the quantity of the blood ? 28 CLASS-BOOK OF PHYSIOLOGY. increase or diminish this portion of the blood, and make it subject to frequent variations in quantity. 49. The clot or crossamentum is composed of fibrin and numerous red particles, called blood discs or corpuscles. 50. Fibrin is white when obtained pure, as it may be by repeated washings. Its peculiar property is to coagu- late spontaneously. It forms the basis of the muscles, and is found in lymph and chyle. It is also found in solution in the serum, as the blood flows in the vessels of a living part, but coagulates soon after the blood is exposed to the air, and forms first a jelly-like mass, and then, as the con- traction progresses, it entangles the coloring matter, and presses out the serum—thus forming an imperfect analysis of the blood. 51. The blood discs, which contain the coloring matter, have a size and form peculiar to each species of animal in which they exist. In man, they are little round cells, flat- tened like a piece of money, and from T7Votns to ^tot^s Fig i.—A, Corpuscles of human blood, magnified five hundred diameters, a, Particles collected in a columnar form. B, Red particles of the blood of the common fowl, b, A particle seen edgeways. of an. inch in diameter. {Fig. 4.) In birds, reptiles, and fishes, they become necessarily larger, till, in some species of fish, they present a surface about six times as large as those found in man. The number of the blood discs cor- respond very much to the temperature of the animal. In warm-blooded animals, they form from twelve to fifteen Of what is the clot composed? What i3 the color of fibrin when pure? What is its peculiar property? In what is it found? How does the coagu- lation of the blood form an imperfect analysis of the blood? What is the color and form of the blood discs? What is the size of the blood discs in different animals? THE BLOOD. 29 per cent, of the whole mass of the blood; in birds, fifteen per cent.; in man, twelve or thirteen per cent. In some of the cold-blooded animals—in fishes, for example—they form only five or six per cent, of the whole weight of the blood. The coloring matter of the blood discs contains nearly seven per cent, of iron. In certain diseases, in which the proportion of iron is diminished below the natu- ral standard, the capacity of maintaining animal heat has been found correspondingly reduced. In such cases, some preparation of iron has been found the best remedy. From these facts it is inferred that the blood discs perform an important office in maintaining animal heat. 52. By chemical analysis, the blood is found to have nearly the same elements, combined in the same propor- tions as they exist in the flesh of the animal. It is there- fore better fitted to renovate the tissues, and to carry.nu- triment to every part of the system—to furnish at one point the elements of bone; at another, those of muscle; at another, those of brain, and so on. The blood also takes up and carries off, through appropriate organs, all waste particles; thus maintaining in the body a continuous round of organization and decomposition—of growth and decay—a perpetual change of particles, new and old. CIRCULATION OF THE BLOOD. 53. The organs by means of which the blood is carried from one part of the body to the other, constitute the cir- culatory apparatus, and the course of the blood through the organs is called its circulation. 54. If we commence with those animals which are lovv- What the number? Under what circumstances is the iron found to be diminished ? What is the remedy in such cases ? In what vital process do the blood discs perform an important office? What elements is the blood found to possess by chemical analysis? What does the blood carry to every part of the system? What does the blood take up and carry off? What is the circulating apparatus? What the circulation ? What is the condition of the circulatory apparatus in different orders of animals? 30 CLASS-BOOK OF PHYSIOLOGY est in the scale of organization, we shall find the apparatus for circulating the blood exceedingly simple, though it becomes more and more complicated as we ascend to higher orders. 55. In insects, the blood is sent to various parts of the body by the alternate contractions of different portions of a central vessel which runs along the back, and thus forms a rudimentary heart. (Fig. 5, and fig. 1, Pl. I.) Fig. 5.—Dorsal Vessel of the Spider.—a, the abdomen ; i, the dorsal vessel or heart; c, u trunk passing to the head ; (/, commence secreting actively an acid fluid, which exudes in drops, running down the walls of the stomach, and soaks into the substances within it. 169. The gastric fluid is a very powerful solvent of all proper food of every kind, whether animal or vegetable. It is indis- pensable to digestion, and is found do where but in the living stomach. 170. The amount of gastric fluid se- creted at one time, corresponds with the Fig. 35.—onb or the quantity of food which is then needed by Tubular Follicles t . "-, ■ i t -i • or the stomach, the body as nourishment. It begins to flow as soon as the first mouthful of food is introduced into the stomach, and continues to be poured out till the demand for nourishment is supplied, and then ceases. If more food be taken than is sufficient for the wants of the system, it will remain undigested, and be- come a source of irritation and oppression; or, being mixed with that previously received into the stomach, the digestion of the whole is retarded. 171. The sense of hunger is felt when there is a demand for a fresh supply of nourishment, and the stomach is in a condition to pour out its secretion. If the food be then swallowed no faster than the gastric fluid is prepared to be mixed with it, hunger or the desire for food will cease when the secretion ceases, or when just food enough has been taken; but if the food be swallowed twice as fast as it can be supplied with gastric juice, the sense of hunger will continue till twice as much is taken as is actually re- How does the gastric juice act on all kinds of food"? With what does the amount of gastric juice correspond? When is it secreted ? How is the food disposed of when more is taken than is required ? When is the sense of hunger experienced? When does it cease to be felt? What is the conse- quence of swallowing the food twice as fast as it should be? DIGESTION. 88 quired. Hence, rapid eating creates an unnatural appetite, frequently causing nervous irritability, and dyspepsia or disease of the stomach. In this country, where men are governed more by the excitement of business than by any regard to health, rapid eating is a prevailing sin, the con- sequences of which are apparent in a greater proportion of dyspeptic complaints than are to be found in any other country. 172. AVhen the proper kind of food has been taken in proper quantities, the fibres of the muscular coat of the stomach alternately contract, pressing the mass of food backward and forward, and from side to side, exposing every part of it to the action of the gastric juice, until its solution is complete. This process lasts from two to five hours, or even longer, according to the kind of food and the thoroughness with which it has been masticated. The agitation of the food is assisted by the action of the respir- atory organs, in alternately raising and depressing the diaphragm whenever the air is inhaled or exhaled from the lungs. 173. "The readiness with which the gastric fluid acts upon the several articles of food is in some measure deter- mined by the minuteness of division, and the tenderness and the moisture of the substance presented to it. By minute divisions of the food, the extent of surface with which the digestive fluid can come in contact is increased, and its action proportionably accelerated." Hence, when the food is thoroughly masticated, it digests with greater facility. Tender and moist substances also digest more easily than those that are tough, hard, and dry; because they are more easily as well as more thoroughly penetrated by the gastric fluid. 174. The value of any particular substance, as an article What evils result from rapid eating? Describe the manner in which the food is moved backward and forward in the stomach. How long does this process last? What circumstances facilitate the action of the gastric juice on the food? 84 CLASS-BOOK OF PHYSIOLOGY. of diet, is not in all cases in proportion to its digestibility; for many substances that are easily soluble in the gastric fluid afford only a small amount of nourishment, while others that are more difficult of solution are highly nutri- tious. Fresh fish, for instance, may be digested in less than half the time required for beef-steak, though the beef- Rteak is decidedly the most nutritious. A substance, to be nutritive, must not only .contain an abundance of those elements which go to form the tissues of the body, but it must be capable of being digested by the gastric fluid or some other secretion in the alimentary canal, and of being assimilated to the blood. 175. When digestion is completed in the stomach, the food and gastric juice are thoroughly mixed, and converted into a pulpy mass, called chyme. There are many circum- stances, besides the nature of the food, which affect this process. 176. Only a sufficient quantity of food should be taken to fairly fill the stomach, and not to distend it. 177. Sufficient time should elapse after each meal to allow the stomach to become empty before a fresh supply is taken. This interval in adults should generally be five or six hours, though it varies with the kind of food, the condition of the stomach, and the constitutional peculiari- ties of each individual. In young persons, where all the functions are performed with more activity and vigor, the intervals between the meals may be much shorter than in adults. 178. Gentle exercise, both before and after each meal, is favorable to digestion; while excessive exertion, whether bodily or mental, retards it. Is the value of a particular article of diet in proportion to its digestibility I What properties do render an article of diet nutritive ? What is the food converted into when digestion is complete ? What should be the quantity of the food? What time should be allowed between each meal? Why should the intervals between the meals be shorter with young persons than with adults? How does exercise affect digestion? PLATE VIII. ORGANS OF DIGESTION. Figure 1.----General View of the Digestive Organs of Jl/an.—This figure is intended to give a general idea of the forms and relative positions of the organs of digestion.—a, The oesophagus. J, The stomach, c, The duodenum, d, d, d, Convolutions of the small Intestine, e, The coecum. /, Appendix of the coecum. g, Opening of the small into tho large intestine, h, The ascending colon, i, i, Transverse arch of the colon, j, The de- scending colon. /;, The liver. /, The gall-bladder, to, The pancreas', mostly covered by the stomach, o, The spleen.—In this figure, the liver is raised up and the transverse arch of the colon drawn down, in order to show parts which they cover when in their natural situation. Figure 2.----General Aspect of the Abdominal Viscera.—In this figure, the anterioi walls of the abdomen are removed, so as to show the organs in their natural positions. The small intestine is removed.—a, The liver, situated beneath the right arch of the dia- phragm, b, The stomach, c, Epiploa, or floating folds of the peritoneum, d, Summit of the gall-bladder, e, e, Large intestine, showing all its courses. Figure 3.---The Chyle-Vessels and the Thoracic Duct.—a, A portion of the Bmall intestine. 6, b, Origins of lacteals. c, Mesentery, d, Mesenteric glands, e, Lymphatic vessels. /, Thoracic duct, g, Aorta, h, Thoracic duct, curving downward and forward, to empty its chyle at the junctions of the left jugular and sub-clavian veins. Figure 4.----The Pancreas.—This figure is given to show more clearly the situation and connexions of the pancreas, a, The pancreas. A, The duodenum, c, The gall-blad- der, d, Duct of the gall-bladder, which communicates with the hepatic duct, «, which leads from the liver. /, Duct of the pancreas, which opens into the common bile-duct,"', through which the combined secretions of the pancreas and the liver are poured into the duodenum. Figure 5.----The Liver.—Section of the liver, showing the ramifications of the vessels. The hepatic vena porta? is a division bf the abdominal vena portse, into which empty all the veins of the digestive organs. It sends branches into ull parts of the liver. After these branches have reached the capillary state, they are succeeded by the roots of the hepatic veins which unite into three large veins which empty into the inferior vena cava. a, The right or greater lobe, b, The left or smaller lobe, c, Groove which lodges the umbilical vein, d, Hepatic vena portse. f, Hepatic artery. /, Inferior vena cava. Figures 6 and 7.---Liver.—Figure 6 represents a horizontal section of two superficial lobules of the liver, showing the probable arrangement of the biliary ducts, a, a, Inter lobular branches of the hepatic vein*, b, b, Trunk of biliary ducts. Figure 7 represents a horizontal section of three superficial lobules, showing the two principal syetems of blood-vessels, a, a, Inter-lobular veinB proceeding from the hepatic veins, b, b. Inter-lobi lar plexuses formed by branches of the portal veins. P. (:.KrllU£-y Lith DIGESTION. 89 179. A quiet and tranquil state of mind is also essential to good digestion. 180. Drinks taken in large quantities materially inter- fere with the process of digestion. The juices secreted by the stomach itself are sufficient for the solution of most articles of diet; and whenever a superabundance of liquid is swallowed, the first effort of the stomach is to get rid of it. This it does by absorbing the liquid at once and without change through the blood-vessels of the mucous coat. The digestion of the more solid materials is not commenced until the liquid is disposed of. 181. As soon as any portions of the food become suffi- ciently digested to mix Avith the gastric fluid, and form chyme, they are sent through the pyloric orifice of the stomach into the duodenum, or first portion of the aliment- ary canal. (Fig. 33.) Around the pyloric orifice (d), at the inside, is a thick band or valve, called sphincter. When digestion is not going on, this sphincter, as well as a similar one at the cardiac orifice (b), is so completely closed that none Of the contents of the stomach can escape. But towards the termination of the digestive process, the pyloric offers less resistance. First it yields to allow the successively digested portions to go through it, and after- wards it allows the passage of even undigested portions, which are always retained in the stomach much longer than those portions that are readily digested, and produce the irritability and suffering so often experienced after eat- ing indigestible substances. 182. In the intestinal or alimentary canal the food is further acted upon, and undergoes other changes. 183. The alimentary canal is divided into tAvo portions, named, from their difference in diameter, the small and the What effect has the mind on digestion ? How do drinks affect digestion when taken in large quantities? How does the stomach get rid of the fluids? What course does the food take when it has become sufficiently digested to form chyme? Describe the pyloric orifice of the stomach. Into how many portions is the alimentary canal divided ? 7 90 CLASS-BOOK OF PHYSIOLOGY. large intestines. This distinction is much less marked in carnivorous animals than in those which feed on vegeta- bles, and the length of the canal differs greatly in the two orders. In the tiger, for instance, the intestines are about three times the length of the body. In the sheep, they are about twenty-eight times longer than the body, or seven times as long as those of the tiger in proportion to the size of the animal. In those animals Avhich live on a mixed diet, the intestines are of a medium length. In man, they are about six times the length of the body. 184. The small intestine, which commences at the pyloric orifice of the stomach and terminates in the large intes- tine, is about twenty-five feet in length in man, is a narrow tube with thin walls, and is coiled in various directions, as represented at d, fig. 1, Pl. VIII. 185. The large intestine, or colon, is about five feet in length, and resembles in appearance a long sac divided into numerous pouches. 186. The intestines, like the stomach, have three coats— the peritoneal or serous, the muscular, and the mucous. The mucous membrane of the small intestine is found in transverse folds, which greatly increase the extent of its surface. 187. In the substance of the mucous membrane, glands are imbedded, one of which is represented in fig. 36, and the surface is studded with minute process- es, termed villi. 188. Through these villi the nutritive Fig. 36.-sinole gland of portions of the digested food is absorbed, SMALL ;NTBSTINE- and conveyed by a set of vessels, called lacteals, into the general circulation. 189. After the chyme or digested food has passed out What is the length of the alimentary canal in the tiger?—in the sheep?__ in man ? Describe the small intestine—the large intestine. How many coats have the intestines? What glands are imbedded in the mucous membrane of the intestine ? DIGESTION. 91 of the stomach into the small intestine, it becomes mixed with the bile secreted by the liver, and with the juice se- creted by the pancreas or sweet-bread. 190. The pancreas is a long narroAV gland, situated partly behind the right side of the stomach, and within the first curve of the small intestine, as represented at m, fig. 1, Pl. VIII. The fluid which it secretes seems to be of a nearly similar nature with saliva, and is supposed to assist in digesting fatty substances, and rendering them fit for absorption. 191. The liver is the largest gland in the body. It is situated in the right side, beloAV and in contact with the diaphragm, and is divided into several lobes. At its loAver side is the gall-bladder, into Avhich the bile is poured, after being secreted. The gall-bladder thus serves as a reser- voir to the bile. Its duct opens into a duct leading direct from the liver, and forms Avith it the common bile-duct, through which the bile is poured into the small intestine at the same point with the duct from the pancreas. These ducts are represented in fig. 4, Pl. VIII. 192. The bile is a greenish-yelloAV fluid, having an ex- ) tremely bitter taste and a nauseous smell. It is constantly^ ecreted by the liver; but, during fasting, it accumulates ) in the gall-bladder, whence it is poured out on the intro-' duction of food into the stomach. 193. By the mingling of bile and pancreatic juice Avith the chyme in the small intestines, chyle is formed. The chyle is a whitish milk-like fluid, with a somewhat saltish taste. It is, in fact, imperfectly-formed blood, and nearly % resembles blood in its constituent parts. The chyle is ab- j sorbed as the digested food passes along through the small J intestine. The residue of the chyme, consisting of indi-*/ What changes does the food undergo after passing out of the stomach? Describe the pancreas. What is supposed to be the use of the fluid secreted by the pancreas? Describe the liver. How is the gall-bladder situated? What are the properties of the bile ? How is the chyle formed ? How is ii absorbed ? / 92 CLASS-BOOK OF PHYSIOLOGY. gestible substances, gradually becomes more and more dry and hard, till it is at length carried into the large intestine, and thence excreted from the system. 194. The food and the several secretions mixed with it are carried through the intestinal canal, and sloAAdy exposed to the action of the absorbent vessels by the peristaltic or vermicular movements of the intestines. This movement is effected by means of the successive contractions and dilations of the intestinal coats, which extend in a wave- like manner throughout the tube. In health, this move- ment is slow, and generally unperceiA^ed by the mind; but it becomes very perceptible in diarrhea, or when acceler- ated by the influence of purgatives. 195. The natural movements of the bowels are indis- pensable to the continuance of health, and they should not be restrained by tight waist-bands or ligatures around the body. The habit of wearing garments that are too tight is a frequent cause of inactiAre or constipated boAvels, and finally leads to impaired health and disease. What movements force the passage of the food along the alimentary canal? What effect does the wearing of tight garments have on the natural move- ments of the bowels and on health? ABSORPTION. 93 v" CHAPTER VII. ABSORPTION. 196. Absorption is that process by which all the various elements of the body are taken into the circulation. 197. All the tissues of the body, except the enamel of the teeth, and some of very Ioav organization, as the hair and nails, possess to some degree the power of absorption. 198. Fluids are absorbed most readily, and none are absorbed with so great facility as water. A large propor- tion of the weight of the body is due to the water which its tissues absorb. Muscle, in drying, loses three-fourths of its weight, and blood four-fifths. Mummies have been found of adult size with all the organs entire, but reduced, by the evaporation of the watery fluids, to a weight not exceeding eight pounds. 199. The process of absorption is performed throughout all the tissues by two sets of vessels, viz: the Igmphatics or lacteals and blood-vessels. 200. The lymphatics—a portion of which are also lac- teals—constitute a peculiar and distinct system of vessels, which are distributed to every part of the.body where there are blood-vessels. The lymphatics derive their name from the peculiar limpid fluid or lymph which they convey. 201. Those which have their origin in the mucous mem- brane of the alimentary canal have received the name of lacteals, from the milky appearance of the chyle Avhich they absorb. 202. The lymphatics are minute and delicate vessels, with walls so thin that they are nearly invisible unless What is absorption ? What tissues possess the power of absorption ? To what is a large portion of the weight owing? How much of its weight does muscle lose in drying?—and blood, how much? To how small a weight have mummies been found reduced ? By what vessels is the process of absorption performed? What are the lymphatics? From what do lymphatics derive their name ? What are the lymphatics called that have their origin in the mucous membrane ? Describe the lymphatics. 7* 91 CLASS-BOOK* OF PHYSIOLOGY. distended with some colored substance. They commence in closely meshed net-work, or in loops distributed upon the external surface of all the organs, and interspersed among the proper elements and blood-vessels of all the several tissues. Sm^mmk^^Mv 203. Externally the lymphatics present a |fg|||! constricted or knotted appearance, OAving to Fv.37.-LYMPHATics.the existence of numerous valves, formed by semi-circular folds of the lining membrane, arranged in pairs, (fig. 37,) and their edges coming together so as to prevent the flow of fluids, except in the direction towards the heart. By this arrangement, all muscular or external pressure accelerates the flow of lymph, as it does that of the blood in the veins. In reptiles and some birds the flow of the lymph is effected by muscular sacs, called lymph-hearts. 204. The lymphatic glands are small, oval, and some- what flattened or rounded bodies, composed of a plexus of minute lymphatic vessels, and a plexus of blood-ves« sels convoluted or twisted upon each other, and inclosed in a thin cellular capsule, in such a manner as to form small knots or kernels, as they are sometimes called, when they become inflamed and SAvollen. In scrofulous persons, these glands are exceedingly liable to become enlarged, particu- larly on the sides of the neck and in the armpits. 205. The contents of the lymphatics pass through the lymphatic glands, and undergo a process of digestion, by which they are renovated and fitted for further use in the animal economy. 206. The lacteals, or lymphatics of the alimentary canal, commence in the villi of the mucous membrane, as repre- How do they commence ? What is the appearance of the lymphatics externally ? How are the valves formed ? Of what use are the valves ? How is the flow of lymph effected in reptiles? What are lymphatic glands? What are they sometimes called when they become inflamed ? How are the contents of the lymphatics affected by passing through these glands ? How do the lacteals commence ? ABSORPTION, 95 Fig. 38.—Diagram of Mucous Membrane.—A, in the intervals of digestion; B, during digestion; a, a, absorb- ent vessels ; *, b, absorbent cells of a villus; c, c, pits or follicles between the villi. senled in fig. 38. Each villi resem- bles in appearance a minute papilla or point in the mu- cous membrane, and is formed of an extremely delicate membrane, termed epithelium Avhich contains a plexus of lacteals, all forming a fine net- work in the sub- mucous tissue, (fig. 39.) Each villus sends forth only p^r- one lacteal vessel, and this so small |j as to be imperceptible to the naked ; eye. || 207. As the lacteals issue from the | ■ ^ villi, they unite to form large trunks, t,' ' and, passing through the mesenteric 1,. glands, join in one common trunk the £_./, thoracic duct, and may thus be com- pared to the roots of a tree, which Fig. 39.— appearance oflym- l . , phatics in the mucous MEM- commence m numberless minute fibres brake of the stomach.—a, . ,, ., j n vt ■ • . n • + superficial layer;*, deep layer. in the soil, and finally join to form its main trunk, the body of the tree. 208. The thoracic duct is the main trunk, into which the contents of the lymphatics and lacteals are emptied. It commences in front of the second lumbar vertebra, and passes up in front of the back-bone to discharge its con- tents by a valvular opening into the large vein just beneath the clavicle or collar-bone. Describe a villus. What do the lacteals unite to form ? Through what gland* do the lacteals pass ? Describe the thoracic duct. Into what does the thoracic duct discharge its contents ? 98 CLASS-BOOK OF PHYSIOLOGY. 209. Absorption by lymphatics or lacteals takes place ■: hrough the Avails or coats of the vessels, and these are so onstituted that only those materials Avhich are suited to ibrm lymph and chyle can traverse them. This system ►f vessels therefore possesses the remarkable property of ■selecting those elements which are in a state of organiza- tion to fulfil certain specific purposes in the animal economy, md all others are rejected. Hence, the lymph and chyle invariably present nearly the same characteristics and contain the same elements. 210. But blood-vessels do not seem to possess any such power of selecting materials as has been attributed to lym- phatics and lacteals; for any substance, Avhether gaseous, Liquid, or a soluble, or a minutely divided solid, may be absorbed by a blood-vessel provided it is capable of per- meating its walls. 211. In the alimentary canal, the fluids are taken up by the veins, and the more solid materials by the lacteals. 212. From the skin, absorption is performed mainly by the veins, though it is a well-known fact that various poi- sonous and medicinal substances take effect through the lymphatics. When irritating substances are rubbed on the skin, red streaks soon appear in the course of the lym- phatics, and the lymphatic glands become swollen and in- flamed. In like manner, poisonous substances acquired through slight wounds, such as are received in dissecting dead bodies in a certain state of decomposition, or by the sting of insects or by the bite of reptiles, extend along the course of these, and produce fatal consequences. 213. There are peculiar conditions of the body in which How does absorption by lymphatics or lacteals take place ? What materials jan traverse the walls of the lymphatics? What remarkable property does this system- of vessels possess? Do blood-vessels possess any power of select- ing materials? What substances maybe absorbed by blood-vessels? How are substances taken up in the alimentary canal ? How is absorption per- formed from the skin? How are poisons known to take effect? Under what circumstances does absorption from the skin take place with great activity? ABSORPTION, 97 the absorption of fluids through the skin is performed with very great activity. Persons Avho are suffering from thirst, receive great relief by applying Avater to the skim When the natural amount of fluids in the body has been dimin- ished by excessive perspiration, or by long abstinence from fluids, the Aveight of the body may be increased several ounces or pounds by immersion in a Avarm bath. —-214. It is on account of the increased activity of the absorbents that contagious disease is acquired more easily after fasting than after a full meal. Fear, anxiety of mind, as well as every thing that depresses the tone and vigor of the system, renders it more susceptible to dis- ase through the increased activity of the absorbents. Hence, epidemics that are propagated by an impure state of the atmosphere are most fatal among those who have the most dread of disease, and thus a quiet mind and an easy conscience are the best safeguards against contagious disease. ^ 215. The necessity for a constant supply of food arises from the continual decomposition which is taking place in the living body. By this process, materials for nutrition are all the Avhile set free, Avhich it is the special office of the.lymphatics to take up and pass through a kind of second digestion, thereby fitting them for further use in forming the tissues of the body. By this process of inter- stitial absorption, animals may live for a long time on their own solids. Hibernating animals, for instance, which retire to their Avinter-quarters loaded Avith fat, gradually become, lean, and spend from four to five months of the year with no supply of nutriment, except such as is Why are contagious diseases acquired more easily after fasting than after a full meal? What causes render the system more susceptible to disease? Among what class of persons are epidemics most fatal? What are the best safeguards against contagious diseases? What creates a necessity for a con- stant supply of food ? What materials are taken up by the lymphatics? How may animals live for a long time on their own solids? What is said in regard to hybernating animals? 98 CLASS-BOOK OF PHYSIOLOGY. derived from the enormous quantities of fat which, they lay in store during the period of their activity. /"" 216. The mucous membrane is the most important i channel through which all the neAV materials for the nour- ishment and growth of the body are introduced. The (more nutritive portions are taken up chiefly by the lac- \teals, while liquids are absorbed by the veins. 217. In the lowest tribes of animals, and in the earliest condition of the higher groups, absorption by the skin seems equally important to the maintenance of life with that which takes place from the digestive cavity. CHAPTER VIII. NUTRITION. 218. Nutrition is that vital process by Avhich the aliment- ary materials are converted into organized tissue. It consists essentially in a deposition in each tissue of the body of those elements which maintain its life and growth. v"- 219. It is the function of the digestive organs to prepare ( from the food the elements of nutrition. By the process v. of absorption, they are taken from the digestive cavity, yand emptied through the thoracic duct into the large vein, t and thence into the right side of the heart, to be passed \ through the lungs into the left side, and from thence dis- tributed by the arteries to all parts of the body. 220. The peculiar process by which the nutritive ele- ments of arterial blood are formed into tissues, is called nutrition. But it is very evident that this process is dependent for its perfection upon other circumstances. There must not only be an abundant supply of nutritive Through what channel are the materials for the nourishment and growth of the body chiefly taken up? What portions are taken up by the lacteals, and what by the veins? What is said in regard to absorption by the skin? What is nutrition? In what does nutrition essentially consist? By what organs are the elements of nutrition prepared ? How are the elements of nutrition taken up? What circumstances may be regarded as essential to nutrition ? NUTRITION. 99 food, but the food must be Avell digested; the absorbents must transfer its nutritive portions, to the general circula- tion. The heart and lungs must also perform their functions properly. If there is a failure in any of these particulars, the body cannot be perfectly nourished. Hence, it not unfrequently happens that the causes of deficient nutrition are not easily comprehended. Appetite, digestion, absorp- tion, respiration, and circulation, may all be regarded as indispensable to a healthy nutrition. 221. The tissues of the body are constantly undergoing a series of rapid changes, in which the particles are decom- posed and removed, to be replaced by neAV ones, which in their turn die and pass aAvay. Yet every atom removed is supplied with a new one, so perfectly resembling the original, that the identity of an adult person is maintained through a series of years Avith the same form, size, and features, and in many instances the same weight, when perhaps not a particle of the original matter that consti- tuted his body is retained. Each part and organ exactly maintains its form and composition during the successive changes of many years. 222. Each individual particle or atom has its own period of life, which is long or short according to the character of the tissue in which it is found. In muscle, for instance, it is supposed the changes are much more frequent than in bone. The milk-teeth exist for a limited period, and are then partially absorbed, and fall out, to be replaced by a more permanent set. The hair of quad- rupeds, the feathers of birds, and the antlers of deer, are shed and reproduced annually with nearly the same form, color, &c. 223. It is supposed that every action of any of the parts of the body is attended with a change of particles. When What changes are the tissues constantly undergoing? How are the par- ticles removed from the body supplied? What is said of the life of each individual particle ? In what tissue are the changes the most frequent? What takes place with every action of any part of the body? 100 CLASS-BOOK OF PHYSIOLOGY. we move an arm, for instance, some of the atoms in the muscles which produce the motion die, and leave their places, being replaced by others. 224. The rapidity of the changes may also be supposed to be in proportion to the activity of the parts. In those who take much exercise, the atoms Avhich compose the body must be changed much oftener than in the indolent and sedentary. In the same manner, the exercise of par- ticular limbs will cause them to appropriate a much larger amount of nutritive matter. An individual Avho walks much, will therefore develop more fully the muscles of the lower limbs; Avhile the blacksmith will develop more fully the muscles of the arms. But a palsied limb, Avhich has fallen into disuse, will emaciate and groAV smaller. ,r'" 225. Physiologists have never been able to determine ( with certainty by what means each tissue selects from the \kblood just those elements which form its own composition. / We must therefore regard it as one of those ultimate facts in nature, Avhich lie beyond the limits of our present \ knowledge. 226. It is supposed, however, "that the process of nutri- tion consists in the groAvth of individual cells composing the fabric, and that these derive their support from the organic compounds Avith which they are supplied by the blood, and that the structure composing every separate portion of the body has what may be termed a special affinity for some particular constituents of the blood, caus- ing it to absorb from that fluid, and convert into its own substance, certain of its elements." /~227. When the amount of alimentary materials con- \ verted into tissue exceeds the waste of the decaying ele- How is this illustrated by a movement of the arm ? The rapidity of the changes are in proportion to what? What is the effect of exercising particu- lar limbs? Have physiologists been able to determine the means by which each tissue selects those elements which form its own composition? In what is the process of nutrition supposed to consist? When is the body said to increase in size? SECRETION. 101 / ments, the body is said to grow old, or increase in size. When the nutrition is less than the loss by decomposition, the body becomes emaciated. In the earlier periods of life, \ nutrition goes on rapidly, speedily producing the renewal \and groAvth of the various parts of the body. When the ,-nDody has attained its full size, the process of nutrition be- comes less active as life advances. In middle life, nutrition ! just equals the loss by decomposition; in old age, the body j \ becomes so imperfectly nourished, that it is literally worn -•'' \ out Avith years. CHAPTER IX. SECRETION. 228'. Secretion is the process in plants and animals by which various materials are separated from the circulating fluid. From the sap of different plants, in Avhich chemical analysis cannot detect the slightest difference, the most opposite and varying products are elaborated. Thus, the sap of the poppy produces the narcotic opium; that of the cherry-laurel, the deadly prussic acid; that of the olive, its oil. Acids are obtained from some; alkalies, from others; SAveet juices, nutritive principles, oils, and resins, from others; different secretions are obtained from different parts of the same plant. In animals, the nutritious milk is secreted in one organ; bile in another; mucus, saliva, urine, &c, in others; and yet we are wholly ignorant of the reasons why, by the same process and out of the same fluid, so many different secretions are formed. All that we know is in regard to the structure of the organs con- cerned in elaborating those various secretions. When is it said to become emaciated ? How does the activity of nutrition vary at different periods of life ? What is secretion ? What is said of the secretions obtained from the sap of plants? What are some of the secretions obtained from the sap of plants? What are some of the secretions obtained from animals? 8 102 CLASS-BOOK OF PHYSIOLOGY. 229. The process of secretion, both in the animal and in the plant, is performed by cells, which are arranged in different methods, according to the structure of the organ in Avhich they are found. Those cells seem to have the poAver of separating the peculiar secretion of each organ from the blood of the membrane with which they are con- nected. At parts where it is necessary that the secretion should be particularly abundant, the secreting surface is increased by great, numbers of little bags or follicles lined with cells. 230. There are tAVO kinds of materials which are secreted from the blood of man, namely, those which go to form such products as may serve some useful purpose in the animal economy, called secretions, and those which Avould be injurious, if retained in the system, called excretions. 231. The two principal divisions of the secreting ap- paratus are membranes and glands. The principal secret- ing membranes are the serous and mucous membranes and the skin. 232. The serous membrane forms closed sacs, which envelop all the organs of the body not exposed to the air, and also line the cavities in Avhich those organs are con- tained. The brain, heart, lungs, and the organs inclosed in the abdomen, are all protected by this membrane. A gaseous fluid is constantly secreted by it, which makes its surface smooth and moist, so as to render the movements of the organs free and easy, and prevent any friction where the surfaces come in contact with each other. 233. A peculiar variety of the same membrane is found at the ends of the bones, Avhich secretes a thick fluid to lubricate the joints, called synovial fluid, from its resem- blance to the white of an egg. How is the process of secretion performed in animals and in plants ? What power do the cells seem to possess? How is the secreting surface increased in particular parts ? What two kinds of materials are secreted from the blood of man? What are the principal divisions of the secreting apparatus? What organs are protected by the serous membrane ? What is secreted by the serous membrane I How are the ends of the bones protected ? SECRETION. 103 234. The mucous membrane lines all those passages which communicate with the air—'as the nose, the larynx, the bronchi and their minute ramifications in the lungs, and the whole digestive apparatus—from the mouth through the intestines. It secretes a peculiar viscid fluid, called mucus, which serves to moisten and defend this membrane, and aid in the exercise of smell and taste. When the mucous membrane becomes irritated or inflamed, as in common colds, causing catarrh, or in diarrhea and dysen- tery, it secretes a large amount of mucus, Avhich some- times becomes quite thick and hard, like the white of an egg when boiled or hardened by alcohol. 235. The most important of the secreting glands are the liver, the kidneys, the salivary and lachrymoJ glands. 236. The liver is found in some form in nearly all ani- mals which possess a digestive cavity. In man and the mammalia, it is made up of a vast number of minute lob- ules, of irregular form. Each lobule contains a mass of cells, and a plexus of biliary ducts, Avith three blood-ves- sels ; namely, the hepatic artery, the portal vein, and the hepatic vein. (Figs. 5 and 6, Pl. VIII.) 237. The hepatic artery furnishes nourishment to the substance of each lobule; the portal vein provides blood for the secreting operation; and the hepatic vein carries back the blood derived from both sources. The bile is taken up from each lobule by the plexus of biliary ducts which finally unite into a common duct. 238. The function of the liver in secreting bile is of \ very great importance to health. When the liver becomes'"') inactive, or ceases to perform its office perfectly from any' What passages are lined by the mucous membrane? What is the secretion of the mucous membrane called ? What is its use ? What are the most important of the secreting glands ? In what animals is the liver found ? Of . what is the liver made up in man and the mammalia ? What does each lobule contain ? Describe the use of tbe hepatic artery—the portal vein—the hepatic vein. How is the bile taken up from each lobule ? What is said of the function of the liver? How is the health affected by inaction of the liver ? I 104 CLASS-BOOK OF PHYSIOLOGY. other cause, the health very soon becomes impaired, caus- \ ing head-ache, languor, and nausea, if not more serious J disease. The object of the secretion of bile is to separate '.—from the blood those materials Avhich Avould be injurious if retained in the system, though in the form of bile they/ afford important aid to the process of digestion. 239. The kidneys also secrete or separate from the blood certain superfluous or Avaste materials. 240. The external or cortical substance of each kidney is composed of an infinite number of ramifications of blood- vessels. The central or medullary part is formed of nu- merous straight tubes, which are collected into conical bundles, all terminating in one common basin or pelvis, as it is called. (Fig. 40.) As the urine is secreted in the pelvis of the kidney it is conveyed by the tubes, named ureters, into the bladder. 241. The salivary glands, including the parotid, sub- maxillary, and lingual, and the lachrymal glands, are all What is the object of the secretion of bile? What is the function of the kidney ? Of what is the cortical substance of the kidney composed ? Of what the central or medullary portion ? What do the salivary glands include 1 SECRETION. 105 similar in structure. They are composed of an aggrega- tion of small lobules, the cells of which open by minute ducts, Avhich converge and unite into larger and larger ducts, and at last into a common trunk, through which their contents are carried to the point Avhere the peculiar fluid of each is needed. The structure of the parotid gland is represented in fig. 41. 242. The quantity and character of the secretion are influenced by variations in the quantity of blood, and of the peculiar materials for any secretion Avhich it contains, and by the condition of the nervous system. An increase in the quantity of blood which passes through a gland, generally increases its secretions. Through the nerves, various emotions of the mind influence the secretions: the thoughts of food increase the flow of saliva, and grief or pain produces a Aoav of tears. In the case of milk, not only the quantity, but the quality, of the secretion is influ enced by the state of the mind. 243. The office of excretion is especially performed by the lungs, the liver, the kidneys, and the skin. The lungs throw off carbon and hydrogen, in the form of carbonic acid and vapor. The liver separates the same elements from the blood, in the form of a peculiar fatty matter; the kid- neys, in the form of urine; and the skin, in that of sweat. If the excretions, or either of them, be checked, they speedily accumulate in the blood, and often lead to the most deleterious results. Of what are the salivary and lachrymal glaads composed ? What circum- stances influence the quantity and character of the secretions? How do emotions of the mind influence the secretions? By what organs is the func- tion of secretion performed? What substances are thrown off from the lungs ?—the liver ?—the kidneys ?—the skin ? 8* 106 CLAbS-BOOK OF PHYSIOLOGY. CHAPTER X. THE SKIN 244. The skin is the external covering of the body, and consists of tAVO principal layers. The epidermis, or scarf- skin, and the dermis, or true skin. 245. The epidermis is made up of cells in different stages of drying. Those nearest to the true skin are rounded or oval in form, and, as they approach the surface, lose by evaporation the fluid they contain, and are gradu- ally converted into scales or scarf-skin. The outer layers of the epidermis are constantly being Avorn off, and neAV layers as constantly formed from Avithin. Thus, each layer is gradually pushed from Avithin, outward. 246. The epidermis is raised from the true skin in the formation of a blister, and portions of a considerable extent peal off after some diseases, such as scarlet fever. It is also removed with the hair, in the process of dressing the hides of animals, leaving the true skin Avith a smooth surface. The minute scales of Avhich the surface of the scarf-skin is composed, are constantly cast off by ablution and fric- tion of the skin. On the scalp, these scales are removed in the form of "dandruff" or scarf. 247. The pigment cells, containing the coloring matter Avhich causes the different varieties of complexion, are de- posited in the soft layers of the cuticle next to the true skin. (Fig. 8, Pl. IX.) In Albinoes there is no Coloring matter: the skin presents a uniform and pearly whiteness, and the hair is also Avhite and colorless. 248. The epidermis is not provided with either nerves What are the principal layers of the skin ? How is the epidermis formed ? What becomes of the external layers of the epidermis ? How is the epider- mis raised from the true skin ? How are the minute,scales of which the surface of epidermis is composed, cast off? How are the different varieties of complexion caused ? Where are the pigment cells deposited ? Why is the epidermis a hard, insensible membrane ? PLATE IX. THE SKIN. Figure 1.----A Portion of the Epidermis of the Palm of the Hand, magnified by a simple lens, Showing the direction of the rugae or wrinkles, and the arrangement of the apertures of the sudoriferous Kiauus. Figure 2.---A portion of the same, magnified one hundred diameters. Figure 3.---A Transverse Section of the Ridges of the Epidermis of the Palm of Vie Hand, Showing a side view of the apertures of the sudoriferous glands, their spiral ducts, the thickness of the epidermis in that situation, its composition of layers of cells, and its mode of connexion with the true skin. Figure 4.----A Portion of the Epidermis from the back of the Hand, Showing tho dis- position of the folds in that situation, the arrangement of the papillae, the disposition of the hairs, and the apertures of the sudoriferous and sebaceous glands, magnified with a simple lens. Figure 5.----A piece of the same,.magnified one hundred diameters, Showing that each Line is a furrow or groove, a provision which allows of very great extension of the epidermis. Figure 6.----A Square of Cuticle, seen upon its internal surface.—a, The sulci, or depressions, which correspond with the ridges of the external surface, b, The ridges which correspond with the furrows of the external surface. Figure 7.----The Under Surface of the Epidermis, Showing the pigment-cells, which contain the coloring matter of the skin.—These cells are seen to be collected principally In the furrows between the papillas. Figure 8.----The Pigment Cells of the Negro, Showing that his darker complexion is owing to the darkness of the color of the pigment contained in these cells. THE SKIN. Ill or blood-vessels, and is therefore a hard, insensible mem- brane, investing the true skin, and protecting it from im- mediate contact with external objects. Where the friction or pressure to which it is exposed is very great, it becomes quite thick and horn-like. Thus, on the palms of the hands and the soles of the feet, it is much thicker than any Avhere else. 249. The epidermis is marked on the surface by a net- work of lines, Avhich are more numerous and larger near the joints, where they form deep Avrinkles. It is perforated by numerous openings or pores, for the passage of hairs and for the perspiratory ducts. (Figs. 1, 2, 3, Pl. IX.) 250. The dermis or true skin is a firm, elastic membrane, composed of innumerable fibres, interwoven in every di- rection by a vast number of blood-vessels, lymphatics, and nerves. 251. Within and beneath the true skin lie the perspira- tory glands, the hair follicles, and the sebaceous or oil glands. Its outer surface is studded with an infinite num- ber of minute conical elevations, called papilloe. (Fig. 5, Pl. X.) 252. Each pappilla is composed of a minute blood-vessel and a nerve, both of which form a loop, or are bent several times upon each other. The blood-vessel is usually the capillary termination of a vein in an artery. These pa- pillae are the seat of the sense of touch, and are so numer- ous over all the external parts of the body that the skin cannot be punctured by the finest needle without wounding some one of them. They are the most numerous in those parts of the body most exposed to the action of foreign agents, or where the sense of touch is most acute. They are most prominent and most densely set on the ends of How is the eperdermis affected by pressure ? How is it marked on the surface ? By what is the epidermis perforated ? Describe the dermis or true skin. What lies beneath the true skin ? Of what is each papilla composed? What is the blood-vessel ? What sense is located in the papillae ? Where are they the most densely set ? 112 CLASS-BOOK OF PHYSIOLOGY. the fingers and in the palms of the hands, Avhere each raised line is composed of a double toav of them. 253. The perspiratory glands are imbedded in the fatty layer beneath the skin, through which the ducts pass Avith a beautiful spiral coil, as represented 'v&fig. 3, Pl. IX., and fig. 1, Pl. X., and open at the surface obliquely, so as to form a kind of valvular opening. These glands are very numerous over all parts of the body, being on an average about 2800 to every square inch of surface, or about 7,000,000 in all. 254. Fluid is constantly exhaled from the perspiratory glands, and carried off by evaporation as fast as it reaches the surface, in the form of insensible perspiration. During actiAre exercise, or exposure to heat, it is poured off more rapidly, accumulating on the surface in drops, and is then called sensible perspiration. In warm, dry Aveather, Avhen evaporation takes place rapidly, the amount of insensible perspiration may be very great. It has been estimated that from two to three pounds are daily exhaled from the skin of an adult, though the amount of insensible perspira- tion may be very materially increased or diminished ac- cording to the quantity of fluid taken into the system. 255. This Avatery exhalation serves a very important purpose in the animal economy in maintaining a uniform standard of heat, and in carrying off substances that would be highly injurious if retained in the system. The most abundant matters thus carried off are carbonic acid and water. 256. The sebaceous or oil glands are also imbedded in the fatty layer beneath the skin: their ducts open either Where are the perspiratory glanas located ? How do the perspiratory ducts pass through the skin ? How numerous are the perspiratory ducts ? What is constantly exhaled from the perspiratory ducts? What is sensible and what insensible perspiration? How much is estimated to be exhaled from the skin of an adult daily ? How may the perspiration be increased or diminished ? Of what use in the animal economy is this watery exhalation ? What matters are carried off? Where are the oil glands located ? Where do their ducts open? THE SKIN. 113 on the surface of the skin or directly into the follicles of the hair. They are most numerous in those parts largely supplied with hair, as the scalp and face, and are thickly distributed in the nose, lips, and ears. The oily secretion of these glands serves to keep the skin supple and soft, and prevents it from drying and cracking. 257. Along the margin of the eyelids there is a row of these glands, which keep the lids smooth, and prevent the tears from trickling over their edges. Occasionally one of these glands becomes inflamed, and forms what is called a stye. In the passage of the ears, another set of these glands secretes the ear-Avax. 258. The hair and nails are both regarded as appendages to the skin, and contribute to its defence. Each hair originates in a small follicle or body in the fatty substance just beneath the skin. (Fig. 2, Pl. X.) Sebaceous glands open into these follicles, and furnish the oily material with which the hair is anointed and softened. The coloring matter of the hair is secreted by cells within the follicles, and through a deficiency of this secretion the hair becomes gray, and finally white. The shape of individual hairs is cylindrical for the smaller kinds, and oval for those which grow to any length. When left to their natural growth, the end or tip of the hair is always conical or pointed; and in most animals the portion of the hair next to the skin is smaller than the more distant portion. After the growth of the hair has reached a certain point, its nutrition ceases, and the hair falls off, to be replaced by another. In most animals, this occurs periodically, and is called moulting, or shedding of the coat. In the same manner In what parts are the oil glands most numerous ? Of what use is the oily secretion ? What purpose do these glands serve on the margin of the eyelids ? What are these glands called when they become inflamed? What are regarded as appendages of the skin ? How does each hair originate ? How are the hairs softened ? How is the coloring matter secreted ? What causes the hair to become gray? What is the form of each hair? What is the form of the end of hair? What part of the hair is the smallest in most animals ? How do most animals shed their hair ? 114 CLASS-BOOK OF PHYSIOLOGY. the antlers of deer are produced very rapidly till they have attained their growth, when they begin to harden at the base, gradually obstructing the vessels which nourish them, and then fall off, to be again replaced by a new and more beautiful production. 259. The nails are produced from the true skin by an organization of cells similar to the epidermis. Each nail is inserted between a fold of the skin, which is reflected backward to the root of the nail, and then passed forward beneath its under surface, to which it adheres. (Fig. 3, Pl. X.) In the process of groAvth, additions are made to its under surface and to the free edge of its root at the same time, so that growth in thickness and groAvth in length proceed together. 260. The use of the nails is to support and protect the ends of the fingers in grasping, and they are particularly useful in taking hold of minute objects. 261. The structure of the scales, feathers, hoofs, and horns of different animals, like that of the nails and hair, is similar to the epidermis, to which they belong. The feathers of birds are remarkable for the wonderful manner in which they combine strength and lightness with surpass- ing beauty. The stem or body is formed by an apparatus which may be likened to a hair follicle; but there are some additional parts, for the production of the laminse that form the vane of the feather, which are joined to the stem during its production. These laminas, when perfectly formed, are connected by minute barbs at their edges, which hook into one another, and thus give to the entire vane a very strong resisting surface. The substance of which feathers consist is a very bad conductor of heat; and when they are lying one upon the other, they form an admirable protection against cold and moisture. In what manner do deer shed their antlers ? How are the nails produced ? Describe the insertion of the nail and the process of its growth ? What is the use of the nail ? What do the feathers, hoofs, and horns of animals resemble in their structure ? For what are the feathers of birds remarkable ? How are they formed ? PLATE X. THE SKIN. Fiourb 1.----A Section of all the Layers of the Ski?i.—a, The epidermis, b, c, The two layers of the cutis vera, d, A sweat-gland, surrounded by cells of fat, and sending its spiral duct upward through all the layers of the skin to the surface. Figure 2.----A Section of the Skin, Showing the hair follicles and sebaceous glands. a, a, Sebaceous glands, opening into the hair follicle. 6, A hair, with its follicle, c, sur- rounded by fat-cells. Figure 3.----A Section of the Thumb, Showing the manner in which the nail is inserted between the folds of the skin.—a, The last bone of the thumb, b, The cuticle reflected upon the root of the nail, c, The nail, d, The cuticle of the point of the thumb, contin- uous with that at the inner surface of the nail. Figure 4.----Papilla of the Skin, magnified, containing expansions of the nerves and of the capillary blood-vessels. Figure 5.----The Spiral Arrangement of the Papilla at the ends of the Fingers, in which the sense of touch is especially seated. Fis.l PL.X Fie. 5 If 111 fife CHE NERVOUS SYSTEM. 119 CHAPTER XI THE JsERVOUS SYSTEM. 262. The nervous system of man and the higher animals consists of the brain, the spinal cord, the ganglia, and the nerves. 263. The brain is the centre of sensation. The spinal cord, the ganglia, and the nerves are conductors of nervous impressions. 261. The office of the nerves is to convey impressions made upon their extremities to the brain, the organ by which the mind becomes conscious of external objects, and to convey impressions' from the brain to the ultimate dis- tributions of the nerves. 265. The nerves are divided into tAVO classes—nerves of sensation and nerves of motion. Thus, if the hand be acci- dentally placed in contact with a hot iron, the feeling of pain * is communicated to the brain by the nerves of sensation, and the impulse to remove the hand is immediately return- ed from the brain by the nerves of motion. Nerves of sen- sation and nerves of motion have a separate origin from the nervous centre, though they are apparently the same in structure, and are usually in the same bundle of nerve fibres. v"" 266. Nerves have no power of originating impressions. V Those of sensation are stimulated by external agents, and" /those of motion by the will or some other force generated V. in the nervous centres. 267. Nervous force moves along a nerve, like the elec- tric fluid along a wire, without the lapse of an appreciable period of time in its passage. Each nerve-fibre can carry only one kind of an impression: a nerve of motion can Of what does the nervous system consist ? Of what is the brain the centre ? What is the office of the nerves ? How are the nerves divided ? What illus- tration can you give of the function of both ? Have the two classes of nerves • the same origin from the nervous centre ? Do nerves originate impressions ? How is each class of nerves stimulated ? How does nervous force move along a wire ? Can a single nerve-fibre convey more than one kind of impression? What doesra nerve of motion convey ? A\rhat a sensitive fibre ? 120 CLASS-BOOK OF PHYSIOLOGY, convey only motor impulses; a sensitive fibre can transmit only such as may produce sensation, as that of light or sound. ^"268. The fibres of motor nerves, Avhich are distributed f to the muscles, spread forth from their trunks into branches, \ which anastomose with each other, forming a kind of net- \work through the muscle; but the nerves of sensation, ^•hich start from the skin, and convey impressions to the I brain, originate in minute elevations or papillae on the \ surface of the true skin, and in their infinite ramifications \is seated the sense of touch. N269. In order that an individual may become conscious of what is passing around him, an impression must be made by external objects upon the organs of the sensitory nerves in the papillae, and this impression must be con- ducted by the nervous trunk to the brain, and then it \becomes a sensation. On the other hand, before the mind /"can cause any movement of the body to be here formed, > an emotion or act of the will must produce a change at [ the origin of the motor nerves in the brain, and this y change must be conducted along those nerves to the mus- \ cles, where it excites a contraction or muscular effort ^suited to the required purpose. 270. There is another kind of movement which does not seem to be excited by impressions made on the brain. They are called reflex movements, because impressions of this class are conducted only to certain nervous centres, and are then reflected back to the organ whence they originated. 271. Movements of this class are more particularly con- How are the motor fibres distributed to the muscles ? How do the nerves of sensation originate ? What sense is seated in the ramifications of the nerves of sensation ? How does an individual become conscious of what is passing around him ? What takes place before any movement of the body can be effected ? What are those movements called which do not seem to be excited by impressions made on the brain ? Why are they called reflex move- ments? In what ehanges arc the reflex movements particularly concerned? THE NERVOUS SYSTEM. 121 cerned in those changes which have for their object the maintenance of organic life, such as the movements of the digestive canal, the respiratory movements, or the contrac- tion of the eyelids and pupil, to exclude a portion of the rays of light when they are too intense for the retina. 272. The nervous centres of reflex action are the ganglia and the spinal cord. The exercise of this power by the ganglia and spinal cord is a remarkable arrangement, by which those processes that are in constant action for the maintenance of life are performed without any exercise of the will or any consciousness of the mind. It is to be feared that, if the brain was made sensitive to all the digestive and respiratory movements, the mind of many individuals Avould be constantly occupied Avith vigilant inspection of these hidden processes of nature. 273. The true nature of reflex action can be more easily comprehended by studying the comparative development of the nervous system in different groups of animals, and by noting the poAvers of its simplest, compared with its most perfect developments. 274. One of the simplest forms of the nervous system is found in the ascidia, (fig. 42,) one of the lowest of the class of mollusca. In this and similar animals, we find only a single ganglion or nervous centre. At a is seen the orifice by which water enters for supplying the stomach with food, and at b that through which it passes out again. Between these ori- fices is the single ganglion, c, which sends filaments to both orifices, and also over Fig 42-lNERVOUa SYg. the surface of the envelop or mantle, d. TE:*or abcim*. What are the centres of reflex action ? What particular advantage may be supposed to result from this arrangement ? How may the true nature of reflex action be more easily comprehended ? In what animal is one of the simplest forms of the nervous system found ? Describe the nervous system of the ascidia and the phenomena of life exhibited by that animal. 9* 122 CLASS-BOOK OF PHYSIOLOGY. These animals are mostly fixed to one spot during their Avhole existence. The continual entrance and exit of the currents of Avater constitute the only phenomena of life which they exhibit, except Avhen the current draws in an injurious substance. The mantle then contracts, causing a jet of water to issue from each orifice, and throw off the offending material. This little animal has no eyes or other organ of special sense. The small tentacula or feelers, at the upper orifice, are the only parts Avhich seem to be peculiarly sensitive; and the irritation-caused by the con- tact of a hard substance Avith these, or Avith the general surface of the body, produces an instinctive contraction of the mantle, for the purpose of getting rid of the irritating cause. This contraction can only be performed by the aid of a nervous system, which has the power of receiving impressions, and of exciting the most distant parts of the body to act in accordance with them. 275. Those animals which in- habit bivalve shells exhibit the powers of respiration, sensation, and voluntary motion, and pos- sess a corresponding develop- ment of the nervous system. In the nervous system of the pecten or scallop-shell, (fig. 43,) there is a large ganglion at B, which dis- tributes branches to the gills and mouth, and regulates the re- ng. 43.-nervous system of pec- spiratory movements. Another SS^gSffifftfi&5 ganglion, c, is connected with the the foot; e, oesophagus. thick fleshy part on which the animal crawls, and which is called its foot. Near the sides of the oesophagus, e, are situated two other ganglia, A, a, the nerves of which are distributed to the sensitive ten- tacula which guard the mouth. These two cephalic Describe the nervous system of the pecten. • THE NERVOUS SYSTEM. 123 ganglia, or ganglia of the head, evidently correspond to the brain of higher animals, being the instruments of sen- sation and voluntary power, and they exert a controlling direction over the movements of the animal; while the pedal and bronchial ganglia (those of the foot and of respiration) minister to the reflex actions of the organs supplied by them. 276. In animals of higher orders, the ganglia are more numerous as the variety of functions to be performed becomes greater; and in proportion as Ave ascend the scale, the cephalic ganglia become more and more developed, and more and more elevated above the oesophagus, until they finally meet on the central line above it, and, in the more perfect animals, take their place in the top of the head—overlooking, as it Avere, all the other" organs. 277. The nervous system of in- sects whose actions are generally energetic and rapid, and in Avhich the apparatus of motion (Avings and legs) is highly developed, presents a marked difference from that of the Mollusca, Avhich are usually inert and sluggish. It consists of a large ganglion in the head, analo- gous to the brain of vertebrated animals, and a chain of ganglia, one for each segment of the body, uni- ^ 44.-NERvoua svSTEM of a„ ted by a double cord, as in fig. 44. lNSECT- In the larva or caterpillar, before it is changed into the per- fect insect, the nerves arising from the ganglia are chiefly distributed to the muscles of the legs, and the ganglia are only repetitions of each other, being nearly of uniform size; but in the perfect insect, the wings and legs, which con- What is said of the ganglia in the higher orders of animals ? How does the nervous system of insects compare with that of the Molusca ? Of what does the nervous system of insects consist ? How are the ganglia united ? 124 CLASS-BOOK OF PHYSIOLOGY. stitute the apparatus of locomotion, are confined to the thorax, and the segments of the abdomen have no legs. Accordingly, the ganglia of the thorax, in the perfect insect, are found to be very much increased in size, and sometimes concentrated in one mass, while those of the abdomen are much smaller. 278. This difference is shoAvn in fig. 45, where A represents the nerv- ous system of the caterpillar, from which is produced a species of sphinx or hawk-moth, and B that of the moth itself. In B, the cephalic ganglion is much larger than that of A, since it has become connected with more per- fect eyes and other organs of sense. The ganglia which supply the wings and legs (those next below that of the head) are also greatly enlarged and concentrated, while the abdominal ganglia are diminished, and some of them are wanting. The whole chain is also considerably shortened, the body of the moth not being so long as that of the caterpillar. 279. The chain of ganglia in in- sects is found to consist of two dis- tinct tracts—one is composed of nerve-fibres only, and passes from the cephalic ganglion over the surface of all the other gang- lia, giving branches to the nerves which proceed from each other, while the other tract connects the ganglia themselves. Thus, each segment of the insect has a distinct nervous connection with its own ganglion, and a sympathetic con- nection with the others, extending to the cephalic ganglion, How do the ganglia of the perfect insect differ from that of the larva or caterpillar? Describe Fig. 45, with the difference between A and B. Ex- plain the arrangement of the ganglia of insects. Fig. 45.—Nervous System of Sphinx-ligustri.—A, that of the caterpillar; B, that of the perfect insect. THE NERVOUS SYSTEM. 125 which seems to have a controlling influence over all the rest, and alone to possess the faculty of sensation. Hence, the motions produced by the ganglia of the trunk, when the head of the insect is removed, although they may seem to indicate sensation, are found to be only reflex in their nature—a certain irritation producing a certain movement without choice, and probably without consciousness, on the part of the animal. 280. Thus, if the head of a centipede be cut off while it is in motion, the body will continue to move as before; or if the body be divided as many times as it has segments, each portion will still continue to move, but all conscious- ness seems to be lost: for as the decapitated body comes in contact with an obstacle equal to its own height, it remains fixed against it, the legs moving as before, but without change of direction or the ability of surmounting the ob- stacle—being no longer subject to the will of the animal, but performing reflex movements by the influence of their own ganglia. 281. In vertebrated animals, (those Avhich possess a back- bone or spinal column) the ganglia are no longer scattered through the body, but are united into one continuous mass; and this mass, constituting the brain and spinal cord, is inclosed within the skull and vertebral column, in such a manner as to be protected from the injuries to which it would otherwise be liable. The brain in the higher ver- tebrata consists of a principal mass, called the cerebrum, which occupies all the front and upper part of the skull, and is divided into two hemispheres or halves by a mem- branous partition, running from back to front, and of a smaller mass, called the cerebellum, situated beneath the cerebrum, at the back part of the skull. At the base of the cerebrum there are two ganglia, the olfactory and How is it shown in the case of the centipede that consciousness depends on the cephalic ganglia ? How are the ganglia arranged and protected in verte- bral animals ? Of what parts does the brain of the higher vertebrata consist ? What ganglia at the base of the cerebrum ? 126 CLASS-BOOK OF PHYSIOLOGY. the optic, Avhich belong to the nerves of smell and sight. All these parts in the human brain will be particularly described. 282. In the brain of a fish, (fig. 46,) the parts just men- tioned are beginning to ac- quire considerable develop- ment. The cerebral hemis- pheres (c, h), the optic ganglia (o, p), the cerebellum (c, e), are plainly to be distinguish- ed, and their relative sizes are in proportion to the intelli- gence of the animal. Thus, Fig. 46.-A, brain of a cod; b, brain of the the ganglia which control the shark; ol, olfactory ganglia; v, spinal cord. gense of gight are very large, since this sense is possessed in a very high degree by fishes; while the cerebral hemispheres, to which belong more especially the manifestations of will and thought, are correspondingly smaller. • The cerebellum also, which, as we shall see, is connected with the poAvers of motion, is large, as we should expect to find it in animals possessed of the power of rapid movement, and is larger in the ac tive and rapacious shark than in the less energetic cod. The spinal cord is large, and is divided at the top, so as to form an opening be- tAveen its tAvo halves. In man, as we shall see, this opening becomes entirely closed. 283. In birds, the brain has made a considerable advance towards Avhat it finally becomes in man. The cerebral ^.47.-BRi.iN of a bird, hemispheres (a> ^ 47^ are greatly in- creased in size, and cover in the olfactory ganglia entirely What is said of the brain of a fish ? Describe Fig. 46. What parts are large and what small in a fish ? What is the comparative size of the cerebral hemispheres in birds ? THE NERVOUS SYSTEM. 127 and the optic gangha (b) partly. The cerebellum (c) is also much more developed, as we should expect from the variety of movements performed by birds, but is as yet un- divided into lobes. The spinal cord (d) is still of consider- able size, and is much enlarged at the points from Avhich originate the nerves of the Avings and legs. As might be inferred from these improvements in the structure of the brain, the intelligence of birds is vastly superior to that of the animals previously considered, and they are the first in the ascending scale which are capable of education or training. Did the limits of this work permit, we might thus go on to trace the progressive development of the brain through individual species of birds and mammalia, and to show at each step a corresponding advance in intel- ligence as we approximate to man. 284. In man, the cerebral developments are greatly increased, while the spinal cord is diminished in size. The surface of the brain is not smooth, but divided by furrows into a series of convolutions, by which the surface over which the blood comes in relation Avith the nervous mat- ter is greatly increased, thus adding to the activity of the organ. The tAVO hemispheres are more closely connected Avith each other by fibres running across from each side. The cerebellum is divided into tAVO hemispheres. NERVOUS SYSTEM OF MAN. 285. The nervous system of man and of the higher animals consists of two portions—the cerebrospinal and the sympathetic or* ganglionic. The cerebro-spinal system is composed of the brain and spinal cord, with the nerves which proceed from them to the skin and muscles. It has What is said of the cerebellum ?—the spinal cord ? How do birds compare in intelligence with the animals previously considered ? How are the cere- bral developments in man ?—how the spinal cord? How is the surface of the brain in man ? Of what does the nervous system of man or the higher ani- mals consist ? Of what is the cerebro-spinal system composed ? What has it been called ? 128 CLASS-BOOK OF PHYSIOLOGY. been called the nervous system of animal life, because it is that by which sensations are received and voluntary motions executed, and with which the mind is more im- mediately connected. The sympathetic system is connected Avith the nutritive processes alone, and from its influence over the organs of the thorax and abdomen, has been called the nervous system of organic life. It consists of a chain of ganglia, communicating Avith each other by nerv- ous cords, extending from the head along each side of the oack-bone. Nerves proceed from it to the organs of di- gestion and secretion, and to the heart and blood-vessels. •/spinal cord. 286. The spinal cord is a long irregular cylindrical col- umn of nerve substance, surrounded by a membranous envelop, and inclosed within a long canal, formed by the vertebras (or pieces of the back-bone). It extends from the head to the first or second vertebra of the loins, and is composed of both white and gray nervous matter. The white matter constitutes its greater portion, and is situated externally, while the gray matter occupies its central part. The spinal cord consists of two symmetrical halves, united in the middle line by a set of converging fibres, called a commissure, but separated anteriorly and posteriorly by a vertical fissure—the posterior fissure being deeper, but not so wide and distinct as the anterior. Each lateral half is marked by two longitudinal furrows, which divide it into three columns or tracts, called anterior, lateral, and poste- rior. There is a considerable enlargement of the cord in the lower part of the neck, at the part from which arises the large nerves which supply the upper extremities; and a similar enlargement is also found in the loins, at the origin of nerves which go to the lower extremities. With what is the sympathetic system connected, and what has it been call- ed ? Of what does it consist ? What is the spinal cord ? How is the canal in which it is lodged formed? What is the extent of the spinal cord?—and of what does it consist ? How are the two halves united ? How is each half divided ? What enlargements in the spinal cord ? * THE NERVOUS SYSTEM. 129 )rtio:i of thb - 287. The number of nerves given off by the spinal cord is thirty-one on each side: eight pairs in the region of the neck; twelve in that of the back, corresponding to the twelve pairs of ribs; five in the loins, and six in the pelvis. Each of these nerves arises by two distinct roots, one springing from the anterior column, and the other from the posterior column of the cord, as in fig. 48—a pair of roots on each side, corresponding to each verte- bra. These roots (the posterior ones hav- ing first formed a ganglion) soon unite in a single trunk, which thus combines the properties possessed by both. The poste-^v^p^^ rior set of roots consists exclusively of tl^?n^k^l those fibres which convey impressions to "sler&rTootT^lanJ- the spinal cord and brain, and thus form Si^n&medt/both'i the nerves of sensation; Avhile the anterior /> branch. roots consist of fibres which convey nervous influence from the brain and spinal cord to the muscles, and form the nerves of motion. Thus, if the spinal cord of an animal be laid bare, and the posterior set of roots be touched, acute pain is obviously produced; while, if the anterior roots be irritated, violent motions of the muscles ^supplied by the injured nerves are occasioned. 288. The spinal cord is both a conductor of nervous impressions and a nervous centre of reflex action. "When an impression is made on the extremity of a spinal nerve, it is transmitted to the spinal cord, and thence conducted to the brain, where alone it can be perceived by the mind. In like manner, the stimulus of the will which originates in the brain is conducted along the cord, and thence to the How many nerves are given off by the spinal cord? How many pairs are given off in the neck—back—loins—pelvis? How does each nerve arise from the spinal cord? Of what fibres do the posterior roots consist?—the ante- rior ? What would be the effect of irritating the anterior or the posterior, if the spinal cord of an animal was laid bare? What two functions does the spinal cord perform ? Explain how it acts as a conductor of nervous im- pressions. 10 130 CLASS-BOOK OF PHYSIOLOGY. nerves. In case the spinal cord is lacerated or torn asunder at any point, as sometimes happens in fracture of the spine, this communication to and fro between the brain and the nerves is interrupted, so that the will has no longer any control over the parts below the injury, nor is the mind conscious of any injury that may be inflicted on them. Thus, if the cord be severed in the loins, sensa- tion and the power of voluntary motion in the loAver extremities Avill be lost, Avhile they will be in full exercise in the upper extremities. 289. The spinal cord is also a nervous centre of reflex action. During sleep, when the brain is inactive, respira- tion and all those movements necessary to life go on as usual: liquid poured into the mouth is swallowed—the body changes its position. An animal whose cerebrum has been removed, does the same things. A pigeon, for instance, has been kept alive for several months in that condition; running Avhen pushed, flying when thrown into the air, drinking when its beak Avas plunged into water, and swallowing food Avhen placed in its mouth, but at all other times appearing in a profound sleep. 290. From such facts, it is evident that the spinal cord must possess a great degree of independent poAver; but it differs from that of the brain in this, that it is exerted without the concurrence of the judgment and the will; and the movements produced by it are rather like those of an automaton, set in motion by pulling certain wires or touching certain springs. Thus, the motions of a decapi- tated animal are never spontaneous, but always excited by a stimulus of some kind. What are the effects of lacerating or dividing the cord? Explain how the spinal cord is a nervous centre of reflex actions? Does the spinal cord pos- sess any degree of independent power? How does it differ from that of the brain ? THE NERVOUS SYSTEM. 131 MEDULLA OBLONGATA. f 291. At its upper part the spinal cord becomes greatly \ / enlarged, and forms a bulb-shaped body, called the medulla \ j oblongata, which is inclosed Avithin the skull, and through which the connection between the spinal cord and the / , brain is preserved. Its columns are continuous Avith those . ^>i the spinal cord, but are more prominent,'and it contains \ : a larger quantity of gray matter. It is connected with the ; nerves of respiration, mastication, and deglutition, and / may be considered as the common centre of the cerebro^/ Nspinal system. 292. The medulla oblongata differs from the rest of the spinal cord in its functions mainly in the importance and extent of the actions governed by it. Like the cord, it is a conductor of nervous impressions; but it has a wider extent of function, since all impressions which pass to and from the brain and spinal cord must be transmitted through it. Motor impressions are transmitted through its anterior columns, and sensitive impressions through its posterior columns. Thus, if one of its anterior columns be divided, the animal will lose the poAver of motion in one half of the body, while its sensation will remain unimpaired. 293. The functions of the medulla oblongata, as a nerv- ous centre, are more immediately important to the main- tenance of life than that of any other part of the system. The nervous force necessary for deglutition and respira- tion is generated in this organ. It has been proved by repeated experiments that the entire brain may be cut away, in successive portions, and life yet continue for a considerable period, and the respiratory movements be Where is the medulla oblongata? How is it formed? What is said of its columns? With what functions are its nerves connected? How does the medulla oblongata differ from the rest of the spinal cord? Of what import- ance to the maintenance of life are the functions of the medulla oblongata? By what experiments has it been proved that life can be maintained so long as this portion remains uninjured? 132 CLASS-BOOK OF PHYSIOLOGY. uninterrupted. Life may also continue Avhen the spinal cord is in like manner cut away from below upAvard as high as the phrenic nerve, which commences near the throat. In some of the amphibia, (tortoises, frogs, &c.,) the brain and spinal cord have both been thus removed, and still respiration and life continued so long as the medulla oblongata Avas untouched. But a very slight wound at its central portion will produce suffocation and sudden death. 294. The power of reflex action is more apparent in this than in any other portion of the nervous centres. By this power, the respiratory movements are carried on. Thus, the application of stimuli to many parts of the body, the nerves of Avhich transmit impressions to the medulla, will cause respiratory movements by reflection through the nerves which proceed from the medulla to the muscles concerned in respiration. This accounts for the " catching for breath" produced by dashing cold water in the face, Convulsive movements are also produced through the agency of this part of the cord. In those convulsions which result from the teething of children, in lock-jaw, and other like diseases, death usually ensues from suffoca- tion—the muscles of respiration becoming so fixed that the air cannot be breathed. THE BRAIN. 295. The human brain consists of two principal portions —the cerebrum or large brain, and the cerebellum or small brain—protected by their membranes, and inclosed within the cranium or skull. The cerebrum, which constitutes nearly seven-eighths of the whole mass, occupies all the upper and anterior portions of the cranium, and is divided What has been 6aid of the power of reflex action in this portion? What effects are produced on the respiratory movements by stimuli applied to other portions of the body? How are convulsions often produced? What is usually the immediate cause of death in such cases? Of how many portions does the brain consist, and what are they called? What portion of the cranium is occupied by the cerebrum? THE NERVOUS SYSTEM. 133 into tAvo hemispheres or lateral halves. These are con- nected by transverse bands or commissures of nervous matter, and each hemisphere has three prominent masses or lobes—one at the forehead, one at the temples, and one at the back of the head. Its surface is intersected by deep fissures and eminences, which produce those Avinding ine- qualities called convolutions. 296. The nervous tissue of the cerebrum is composed of Avhite and gray substances, disposed in a peculiar man- ner: a layer of the gray substance occupies the surface, and folloAvs all the irregularities of the convolutions, while the Avhite substance is placed in the central parts. Thus, each convolution consists of a layer of gray substance at the outside and of white substance Avithin. The manner in which the two substances are thus arranged to form the convolutions, may be rudely illustrated by taking two pieces of cloth, laying one upon the other, and collecting them up into folds in a globular shape. By such arrange- ment, the surface over which the blood comes in contact with the nervous matter is greatly increased, and it is interesting to know that in man the convolutions are more numerous and extensive, and the depressions between them deeper, than in any other animal. The brain is most abundantly supplied with blood—the amount sent to it having been estimated as high as one-fifth of that con- tained in the whole body, though one-tenth Avould be probably nearer the truth. 297. ToAvards the base and centre of the cerebrum, the surface is inflected inward, so as to form an intricate inter- nal cavity, several compartments of which are called ven- How is the cerebrum divided ? How are the two halves connected to- gether? What is the external appearance of each hemisphere"? Ofwhatis the nervous tissue of the cerebrum composed ? How are the white and gray Bubstances disposed ? How may the manner in which the two substances are disposed be illustrated? What is the effect of this arrangement? What proportion of the whole amount of blood has been estimated to be sent to the brain ? What are the internal cavities railed ? 10* 134 CLASS-BOOK OF PHYSIOLOGY. tricles. They communicate with each other, and serve still further to increase the cerebral surfaces. It is in these ventricles that collections of serous fluid take place, in cases of what is called "dropsy of the brain," or Avater in the head. 298. The cerebellum is situated at the back of the head, below the cerebrum. It is divided into two lateral lobes, and, instead of waving convolutions, its surface presents a number of plaits with furrows between them. It is com- posed, like the cerebrum, of gray and white matter, and the Avhite matter presents, on a vertical section of one of the lobes, a tree-like appearance, called arbor vitce (tree of life). 299. The membranes Avhich surround and protect the brain are three in number, and are called the dura mater, the pia mater, and the arachnoid. These membranes are also prolonged downwards, so as to form a tubular sheath to the spinal cord. The dura mater (hard mother) was so called by the old anatomists, because they supposed it to be the origin or mother of all the hard, firm membranes of the body, and the pia mater (tender mother) Avas also thus named by them, as being the origin of the soft membranes. ' 300. The dura mater is a strong, dense, fibrous mem- brane, which forms the external envelop of the brain, and is in contact with the bones of the skull, to Avhich it strongly adheres. It is separated into two layers, the inter- nal of which is doubled on itself, so as to form two remark- able processes: the one, from its resemblance in shape to a sickle, is called falx cerebri, and is interposed between the two hemispheres of the brain, so that Avhen the head How do they communicate with each other? Where is the cerebellum situated?—how is it divided? Of what two substances is it composed? What is the appearance of the matter when divided vertically? How is the brain protected ? What was the origin of the name applied to the dura mater and pia mater ? Describe the dura mater. What is that portion of the dura mater called which divides the two hemispheres ? THE NERVOUS SYSTEM. 135 rests on one side, the uppermost hemisphere is prevented from pressing upon the lower. The other process is called tentorium cerebelli, and is extended over the cerebellum, so as to prevent the pressure of the cerebrum upon the latter when the head is in an erect position. There is also a smaller process, called falx cerebelli, which separates the two hemispheres of the cerebellum. 301. The arachnoid (spider's web) membrane, so called. from its extreme thinness, lies beneath the dura mater, and is spread over the entire surface of the brain. It corres- ponds in its use to the serous membranes of the heart and other organs, and serves to keep the opposite surfaces of the dura mater and the pia mater, between which it lies, moist and smooth. 302. The pia mater is a loose transparent web, in which a multitude of blood-vessels cross each other in every pos- sible direction. Minute branches of these vessels, in im- mense numbers, penetrate the brain, to which they afford nutriment, at the same time that they serve as a means of attachment between the brain and membrane. The pia mater follows all the convolutions, entering all the cavities of the brain, and is also prolonged over the spinal cord and •the nerves, constituting one of the most important membranes of the body. 303. Twelve pairs of nerves are given off from the brain, and are named as follows: 1st pair, Olfactory nerves; 2d, Optic Nerves; 3d, Motores Oculorum; 4th, Patheticus; 5th, Trifacial nerves; 6th, Abducentes; 7th, Portia dura, or facial nerves; 8th, Portio Mollis, or auditory nerves; 9th, (xlosso-pharyngeal nerve; 10th, Pneumogastric nerve; 11th, Hypo-glossal nerve; 12th, Spinal accessory nerve. ^X 304. The first pair is distributed to the inner membrane \pf the nose, and transmits to the brain the impression pro- "What other processes are there ? Describe the arachnoid membrane. De- scribe the pia mater. How many pairs of nerves are given off from the brain ? What are they called? Where is the first pair distributed, and what is its size ?—the second ? 186 CLASS-BOOK OF PHYSIOLOGY. /^duced by odors. The second pair is distributed to the / retina of the eyes, and in like manner conveys the im- ; pression of sight. The third, fourth, and sixth pairs are nerves of motion only, and are distributed to the muscles \ which move the eyes. The fifth pair is for the most part \ a nerve of sensation only; it is divided into three branches ^sr-fhe f[rst of which, called the ophthalmic nerve, passes / into the orbit or cavity in which the eye is lodged, and is then distributed to the forehead and temples. The second branch, or superior maxillary, supplies the cheeks, nose, and upper-lip with sensitive filaments. The third, or inferior maxillary, which, like the spinal nerves, has also a motor root, imparts the poAver of moving to the masti- cating muscles, and gives sensibility to the parts about \ the mouth. \ 305. The seventh pair is the general motor nerve of the muscles of the face; the eighth pair is the nerve of hear- ing, and is distributed to the internal ear; the ninth pair supplies the back of the mouth and pharynx, and is con- cerned in the act of swallowing; the tenth pair originates from the medulla oblongata, and supplies the lungs and air passages, as also the heart and stomach; the eleventh pair gives motion to the tongue; and the twelfth pair is concerned in respiration. All these nerves are supposed to contain two sets of filaments—one communicating with the cerebral hemispheres, and the other with the spinal cord. Plate XI., fig. 4, represents a section of the brain, showing the arrangements of these nerves. That of the nerves proceeding from the spinal cord is seen in Plate XL, Jig. 1. What are the third, fourth, and sixth pairs ? What is the fifth pair ? How is it divided ? Where is each branch of the fifth distributed ? What is the seventh pair?—the eighth? What is the origin of the ninth pair? What organs does it supply ? To what parts do the eleventh and twelfth give motion? What two sets of filaments are all these nerves supposed to contain ? XI / PLATE XI. THE BRAIN AND SPINAL CORD. Figure I.----Anterior View of the Brain and Spinal Cor.i, isolated from the Skeleton, —a, b, Right and left hemispheres of the brain, the left hemisphere covered by the arach- noid and pia-mater; the right hemisphere naked, displaying the convolutions, c, d, Right and left lobes of the cerebellum, e, The medulla oblongata. /,/, The spinal cord, cov- ered by the pia-mater on the right side, and showing the origins of the spinal nerves on the left. At/,/, are seen the two enlargements of the cord, in the neck and the loins, \ g1 to Si The eight cervical pairs of nerves; h to h, The twelve dorsal pairs; i to t, Tho five \ lumbar pairs; k, k, The six sacral pairs. I, Lateral column of the cord, m, Posterior column. Figure 2.----A Section of the Brain and Spinal Cord, inclosed in the Skull and Verte- bral. Column.—a, The cerebrum, b, cerebellum, c, c, The spinal cord.—The vertebra are cut through, so as to display a lateral view of the cord. Fioure 3.----The Cerebellum.—A, Anterior view. B, Posterior view. s"^ Figure 4.---A Vertical Section of the Brain, showing the Origins of its Nerves.— f a, a, The cerebrum, with its convolutions, b, The cerebellum, displaying the arbor-vitae / upon its section, c, The medulla oblongata, d, The corpus callosum, a band of fibres which connects the two hemispheres of the brain, e, The eye. | /, The first pair of nerves. I \ g, The second pair. 1 h, The third pair, n, The fourth pair. Ij, The fifth pair, k, The sixth \ pair. /, The seventh pair, m, The eighth pair, n, The ninth pair, o, Tho tenth pair. p, The eleventh pair, q, The twelfth pair, r, Spinal nerves. Figure 5.---A Horizontal Section of the Brain, showing its Interior.—a, a, The cineritious or gray substance at the outside of the brain, following the convolutions. ft, *, The white or medullary substance at the inside of the brain, c, c, The lateral and middle ventricles. Fi°. 2 THE nervous system. 141 FUNCTIONS OF THE CEREBELLUM. 306. Much discussion has taken place in regard to the functions of the cerebellum. It seems to be noAV agreed, however, by the most intelligent physiologists, that it is the organ Avhich is more especially connected Avith motion. Numerous experiments have been made on living animals, all of which go to shoAV that, after the removal of the cerebellum, the power of executing those movements necessary to locomotion is lost. The faculties of volition and sensation remain; but the power to walk, fly, or even stand, is uniformly lost, from the inability to combine the action of the muscles in groups. 307. From facts of this character, it seems to be most probable that the function of the cerebellum is to harmon- ize and regulate the actions of the voluntary muscles. In accordance with this theory, Ave find that those animals which possess variety of motions, or muscular action in the highest degree, are endowed with a cerebejlum cor- respondingly large. 308. In the animals of the cat tribe, which use their limbs for seizing their prey, and which are capable of great muscular exertion, the cerebellum is larger than in those whose limbs are subservient to locomotion only. In birds, the variety of whose movements is still greater, it is larger than in most of the mammalia. It acquires its highest development in man, as might be expected, from the muscular combinations necessary to maintain the erect position, and to perform the intricate and varied movements of the human hand. What is said in regard to the functions of the cerebellum? How is it regarded by the most intelligent physiologists? What do experiments on living animals show in regard to the formations of the cerebellum ? What is most probably the function then, of the cerebellum ? What is said in regard to the development of the cerebellum in the cat tribe and in birds? In what class does the cerebellum acquire its highest development ? 142 class-book of physiology. 309. The influence of each half of the cerebellum ia directed to muscles on the opposite sides of the body; and it would appear that, for the right ordering of movements, the action of its tAVO halves must be ahvays mutually bal- anced and adjusted. FUNCTIONS OF THE CEREBRUM. 810. The cerebrum is the organ through which the phenomena of thought and intelligence are manifested. By its means, we reason upon the ideas excited by sensa- tions, we judge and decide upon our course of action, and put that decision into practice by issuing a mandate Avhich is conveyed by the nervous trunks to the muscles. 311. It is a common, but erroneous idea, that reason is peculiar to man, and that the actions of the lower classes of animals are due to instinct alone. There can be no doubt that reasoning processes, exactly resembling those of man, are performed by many animals: such, for instance, as the dog, the horse, and the elephant. We must admit that an animal reasons when it profits by experience, and obviously adapts its actions to the end it desires to gain, especially when it departs from its natural instincts to do this. The presence of intelligence is also perceived in the differences of character found in various individuals of the same species. Thus, some dogs are stupid, others saga- cious, some ill-tempered, others good-tempered; just as there are stupid men and intelligent men, ill-tempered men and good-tempered men. But the actions of insects seem to be wholly instinctive, so that we observe no dif- ference of temper or capacity in them. 312. Birds, however, which resemble insects in many What is said in regard to the influence of each half of the cerebellum? What is the function of the cerebrum? Is reason peculiar to man? When may an animal be said to reason? How may we discover the presence of intelligence in animals ? What example is given of different degrees of intel- ligence in an animal ? What seems to be the nature of the actions of insects? What is said in regard to birds? THE NERVOUS SYSTEM. 143 of their instinctive tendencies, exhibit a remarkable dis- tinction in their actions. In escaping from danger, in obtaining food, and in constructing their habitations, the actions of birds, like those of insects, are instinctive. But in adapting their operations to peculiar circumstances, birds display a variety and fertility of resource not to be found in insects; and birds also learn by experience, and may be educated, while insects observe perfect uniformity in all their actions. 313. The relative amount of intelligence in different animals bears a pretty constant proportion to the size and development of the cerebral hemispheres. Size alone, hoAvever, does not produce all the difference. In ascend- ing from the lower to the higher animals, a marked ad- vance in the complexion of the brain is observed: the convolutions become more and more prominent, giving a proportionably increased surface for the entrance of the blood-vessels, and an equally increased amount of the gray matter which seems to be the real centre of all the opera- tions of the organ. Still, the size of the cerebrum, com- pared with that of the spinal cord and the ganglia at its top, usually affords a tolerably correct measure of the intelligence of the animal. The same rule holds good in comparing different men, if due allowance be made for the comparative activity of their general functions, or, in other words, for their differences in temperament. 314. Thus, two men whose brain is of the same size may differ widely in mental vigor, because the general system of one performs its functions more actively and energetic- ally than that of the other. For the same reason, a man In what actions do birds seem to be governed by instinct, and in what by intelligence? How does the amount of intelligence in different animals com- pare with the development of the cerebral hemispheres Is all the difference produced by size alone? What other circumstances are to be taken into the account? What is said of the size of the cerebellum, compared with that of the spinal cord ? How does this rule hold in regard to different men ? Why may two men whose brains are of the same sfee differ widely in mental vigor? Under what circumstances will a large brain surpass a small one in power ? 11 144 CLASS-BOOK OF PHYSIOLOGY. of small brain, but whose general habit is active, may have a more powerful intellect than another of much larger brain, but whose system is sluggish and inert. But of two men alike in temperament, arid having the same general configuration of head, it cannot be doubted that the one with the larger brain will surpass the other. It is a strik- ing fact, that almost all those persons who have been eminent for their acquirements, or for the influence of their talents over their fellow-men, have had large brains. This was the case, for example, with Newton, Byron, Cuvier, Cromwell, and Napoleon. The average weight of the brain is about three pounds two ounces. That of Cuvier weighed four pounds eleven and a half ounces, and those of Byron and Cromwell are said (though the fact is doubtful) to have weighed nearly six pounds. In idiots, on the contrary, the brain is usually very small—in some instances weighing only one and a half pounds. 315. The size of the brain may be pretty correctly estimated by the facial angle. This angle is obtained by drawing a horizontal line (c, d, Jigs. 49 and 50) from the Fig. 49.—Skull of European. Fig. 50.—Skull of Nbgro. entrance of the ear to the floor of the cavity of the nose, and a second line (a, b) from the most prominent part of the forehead to the front of the upper jaw, so as to inter- sect the other. This line will evidently be more inclined to the former, and the angle formed by the two will be What has been said in regard to the size of the brain in men who have been distinguished for their talents? What examples are given of large brains? How is the brain in idiots ? How may the size of the brain be estimated ? 5L PLATE XII. THE NERYES. Figure 1.----Posterior View of the Principal Nerves.—The general arrangement of the nervous centres and the distribution of the nervous trunks are shown in this figure.— The spinal column is laid open, so as to display the cord, with the nerves which pass from it. The muscles of the left side and limbs are dissected, to show the course of the princi- pal nerves. a, The hemispheres of the cerebrum, b, The lobes of the cerebellum, c, The spinal cord, d, The facial nerve, the principal motor nerve of the face, e, The brachial plexus, a net-work of nerves, originating by several roots from the spinal cord, nnd going to sup. ply the arm.—From this plexus proceed—/, the scapular nerve; g, the median nerve; A, the ulnar nerve; i, the musculo-spiral nerve ;j, the intercosto-humeral (nerve of Wrisberg). From the spinal cord proceed—the intercostal nerves, k, k, running between the ribs; the nerves forming the luobar plexus, I, from which the front of the leg is supplied; those forming the sacral plexus, m, which supplies the back of the leg. The chief branch of the sacral plexus is the great sciatic nerve, n, which divides into the tibial nerve, o, the peroneal or fibular nerve,p. and many other branches. The nerves seen on the right side of the figure, are the ramifications of the sub-cutaneous nerves, branching beneath the skin, in which they are finally lost. Some of the superficial veins are also represented. THE NERVOUS SYSTEM. 149 more acute in proportion as the face is more projecting and the forehead more retreating; while it will be nearer a right angle, if the forehead be prominent and the muzzle projecting but little. Hence, the facial angle Avill indicate the proportion which the brain bears to the face. 316. This angle is more open in man than in any other animal, and it varies greatly in the various races of men. The difference between the facial angle of the European and American head, and that of the negro, is seen in figs. 49 and 50. In the one, it is about eighty degrees, and in the other about seventy. In monkeys, it varies from about sixty-five to thirty degrees, and as we descend still lower, it becomes still more acute. CONNECTION OF THE MIND WITH THE BODY. 317. The brain, as appears from what has already been said, is the connecting link between the mind and the body. All the organs of the body may be said to be the agents of the mind, inasmuch as the mind manifests itself through them all. Thus, the hand, the eye, the muscles which control the features, are all made to express, more or less forcibly, the varied emotions of the mind. The mind, too, controls, to a certain extent, the operations of the various functions of the body, to which it gives a shape and form in correspondence with itself. 318. In this view, we may regard the entire body as an assemblage of organs for the manifestation of the mind. But the influence of the mind upon the body is recipro- cated by that of the body upon the mind. If the body can be made to suffer from the condition of the mind, the In what class is the facial angle more open than in all others ? How does it vary in different races? What is the facial angle of an European or an American—of a negro—of a monkey? What may we regard as the connect- ing link between- the mind and the body ? How may we regard all the organs of the body? What parts are made to express the emotion of the mind? What control does the mind exert over the body? What then may we regard the body ? How is the influence of the mind upon the body reciprocated ? 11* 150 CLASS-BOOK 'OF PHYSIOLOGY. mind itself is not less affected by the condition of the body. A single illustration will suffice to shoAV this fact: Melan- choly, or depression of spirits from any cause whatever, will often produce disease and derangement of the liver; and, on the other hand, a derangement of the liver will almost always insure melancholy, though no other cause exists. 319. But the intellect is not limited in its influence to the organic and animal functions: it also stamps itself upon every lineament and feature of the man. Thus, as we ascend in -the scale of being, intelligence becomes more highly developed, and is marked by a more perfect organ- ization of the brain and nervous system. The nervous power in man is at its highest point, as is shoAvn by the very liability to disease or derangement of the vital func- tions, arising from the sympathy between the physical and intellectual powers. The mere animal has no desires to gratify but those of appetite. These satiated, he has no anxiety for the future—no repinings for the past; he is not wasted by care, nor harassed by toil and disappointment. Man, on the contrary, is the constant subject of exciting and depressing influences—of functional and organic de- rangements, growing out of his complex nature. Ever restless, never satisfied, he is constantly on the rack of physical and mental torture. He consequently obtains the highest perfection and excellence when he obeys most perfectly the laws of his nature, and retains the physical and the intellectual in the most complete harmony of development. 320. Hence, in the education of the young, the powers of the body and mind should be cultivated together. The What illustration is given to show this fact? Upon what does the intellect Btamp itself? As we ascend the scale of being, how is intelligence marked ? How is it shown that the nervous power is at its highest point in man ? What is said in regard to the cares and anxieties of man, compared with other animals ? When does man obtain the highest perfection ? What should be observed in the education of the young ? THK NERVOUS SYSTEM. 151 mind should not be excited to such a degree as to overtask the brain, nor so neglected as to leave the latter without a proper degree of exercise. One extreme wastes the vital energies while yet immature; the other degrades heaven- born powers to a level with the brute. The injudicious course, so often pursued, of stimulating the intellect of what is called a "precocious child," too often results in misery in the sufferer itself, and in disappointment to its unwise friends. Such children rarely fulfill the promise of their earliest years, and the anxious parent usually sees the brilliancy of their youth fade into dullness or dis- ease in after-life. 321. During the first six or eight years, the most im- portant object in the education of the young is to develop the physical powers, and secure vigor and strength to the body. If there is a tendency to an early development of the intellectual poAvers, special care should be taken to restrain the exercise of the mind, till the physical powers shall become strong and healthy, rather than fan the flame already kindled till it consumes the citadel of life. 322. In the subsequent development of the intellectual faculties, gradual and uniform progress is greatly to be pre- ferred to rapid strides in knowledge. They who at first rush furiously up the rugged path of science, are sure very soon to tire or fall sick by the way. Nor should the mind ever be confined too long on any particular study; a suit- able variety not only refreshes the mind, but affords a more harmonious development of all the powers. The young especially require variety: their minds are con- stantly on the alert for something neAv—something to see, What is said in regard to neglecting or exciting the mind What is the result of either extreme? What is often the result of injudicious stimulating of the intellect of precocious children? What is the most important objec' during the first six or eight years? What should be done if there is a tendency to an early development of the intellectual powers ? What is to be preferred in the subsequent development of the intellectual faculties? What is said in regard to variety of study? 152 CLASS-BOOK OF PHYSIOLOGY. handle, and taste—or for some new thought or truth. Continued application, for a great length of time, is wholly incompatible with their natural impulses, and fatigue very soon ensues when it is attempted; for the mind seeks variety with the same irresistible tendency that the body seeks change of position. 323. In more mature years, the mind should be trained to earnest and patient thought, but not with unremitting toil. The best thinkers are those who possess the power of exerting their brains Avith the most energy for a limited period of time, and Avho then take liberal relaxation. The brain thereby exercises the poAver of prompt, ener- getic action; and when it acts, it is to some purpose. Those who keep the brain constantly at work, either be- come dull and lazy thinkers, or are prematurely worn out. The men Avho have done the most for the world and for the race, are men of vigorous thought and liberal exercise —men who take care of the body while they work the mind. The reason why genius so often finds an early grave, is because the intense exitement it creates in the brain soon exhausts the physical energies. How should the mind be trained in more mature years ? Who are usually the best thinkers? What is the effect of keeping the brain constantly at work ? Who are the men who have done the most for the world ? Why does genius so often find an early grave ? THE SENSES. 153 CHAPTER XII. THE SENSES. 824. There are five senses: touch, taste, smell, hearing, and sight. The office of the senses is to make us acquainted with the material world around us, and the states of our own bodies. Our knowledge of the properties of matter, and our ideas of form, taste, odor, and sound, are all obtained from impressions made on the mind through the senses. SENSE OF TOUCH. 325. The sense of touch is the most universally diffused of all the senses, existing in greater or less perfection in all animals. In some, as in man, nearly the whole exterior of the body is endoAved with it; but in others it is limited to certain parts in which it is specially seated. In insects, the surface of the body is covered with an insensible shell, and the organs of touch are the antennas, or feelers. These are pro- longations from the por- tion of the head near the mouth; they are often of very great length, as in fig. 51, usually contain- ing a great number of points, and are very flex- •ii ,i , ,i t_ Fig. 61.—Capricorn Beetle.—a, a, antennae. lble, so that they can be * ' ' turned toward any object the insect may wish to examine. In walking, the antennse are used almost incessantly in touching the surfaces over which they pass, and they seem How many senses are there ? What is the office of the senses ? How do we obtain our ideas of form, taste, odor, and sound ? Which of the senses is the most universally diffused ? What is the extent in man? How is it limited in other animals ? What are the organs of touch in insects 1 154 CLASS-BOOK OF PHYSIOLOGY. to be the medium of some kind of correspondence with each other; for ants or bees are seldom seen to meet with- out reconnoitering one another with their antennas. 326. In fishes, the body is so covered by scales as to be nearly insensible. In those animals Avhich are covered with hair, the sense of touch is most perfectly developed in those parts which are most uncovered, and are brought in contact with the substances on which they feed. The sensibility of the lips of the horse, for example, is very gTeat. The carnivorous animals are usually provided with long hairs or "whiskers" each of which at its root is in contact with a filament of nerve. (Fig. 6, Pl. X.) 327. In most of the mammalia, the lips and tongue are employed as the chief organs of touch. In the elephant, this sense is possessed very acutely by the little finger-like projection at the end of its trunk; and in the bat, it seems to be diffused over the whole membrane of which the wings are formed. 328. In man, the true skin is studded with minute papillas, and each papilla is provided with a termination or a loop of a sensitory nerve. The whole external surface of the body is thus more or less highly endowed with this faculty, according as the papillae are more or less numer- ous in any particular part. In the hand, and especially at the extremities of the fingers, these papillae are extremely numerous, and the sense of touch is here most acute. (Figs. 4 and 5, Pl. X.) The hand is capable of a great variety of movements, as well as a high degree of improve- ment by education. In the blind, who learn to read with What is the condition of the body in fishes ? Where is the sense of touch most perfectly developed in animals which are covered with hair? What is said of the lips of the horse and the whiskers of carnivorous animals? What are the chief organs of touch in most of the mammalia ? In what parts is it possessed very acutely by the elephant and the bat ? With what is the true skin in man studded ? With what is each papilla provided ? Where are the papilla the most numerous, and the sense of touch the most acute ? What is said of the improvement of the hand by education ? How do the blind learn to read ? THE SENSES. 155 facility by simply touching the raised letters, this sense acquires most wonderful acuteness and precision. 329. In animals, the sense of touch is almost wholly confined to the surfaces of the body, while the deeper parts are comparatively free from it, though not insensible to pain from injury or disease. In case of an ordinary cut or bruise, or a surgical operation, the most acute suffering arises from the violence inflicted on the sensitive nerves near the surface. Thus, a Avound that is only "skin deep," will sometimes cause quite as much immediate pain as" a more serious injury to the deep-seated parts. The sense of Touch may therefore be regarded as an ever-watchful sen- tinel, placed on the exterior of our bodies to guard Avith untiring vigilance the delicate textures within. SENSE OF TASTE. 830. Taste is that sense by which we judge of certain properties of matter—as SAveet, sour, bitter, &c. Its chief purpose is to direct animals in the choice of food. Like the sense of touch, it is not excited except by the direct contact of particular substances; and only those substances which are soluble in water, or in the fluids of the mouth, make any impression on this organ. Substances that are not soluble, are said to be either insipid or tasteless. 331. The sense of taste, in the higher animals, is con- fined principally to the mucous membrane of the tongue, Avhich is very thickly set, especially on its upper surface, Avith papillae. The papilla? of the tongue have some re- semblance in structure to those of the skin, though they are far more highly organized. (Fig. 1, Pl. XIII.) 332. The papilla? of the tongue are of various forms To what parts is the sense of touch almost wholly confined in all animals ? Where is the pain the most acute in cases of injury? What may we regard the sense of touch ? What is taste ? What is its chief purpose ? How is the sense of taste excited ? What is said of substances that are not soluble? To what parts is the sense of taste chiefly confined ? What is said of the papillae of the tongue ?—of their form and size 1 156 CLASS-BOOK OF PHYSIOLOGY. and sizes: some are conical; others are enlarged and flat- tened at the top; some are elevated above the surface of the tongue; and others are imbedded beneath it. The fungiform papillae, which are supposed to be the special instruments of the sense of taste, are composed of numer- ous loops of capillary vessels, Avith a bundle of nerve- fibres, as represented in fig. 5, Pl. XIII. 333. The tongue itself is made up of muscular substance, Avhich accomplishes the various movements required in the act of mastication, and in the production of sounds. It possesses nearly the same structure in the other mamma- lia as in man. In birds, it is usually cartilaginous or bony in its texture, and destitute of nervous papillae, so that their sense of taste cannot be very acute. Several of them use their tongues for other purposes. The wood- pecker, for instance, (fig. 6, Pl. XIII.) whose tongue is sharp and forked, transfixes insects with it; and the par- rot uses it to keep steady the nut or seed which is being crushed between its mandibles. In some reptiles, the tongue is large and fleshy; in others, as the serpent tribe, it is sharp and forked, and possessed of very great quick- ness of motion. In the frog and chameleon, it serves as an organ of prehension, and is darted out with great ra- pidity to catch the insects on which these animals feed. The tongue of the fish is generally in a more rudimentary state; it is fixed in the throat, and often covered with teeth. The tongue of the bee forms a little tube, through which it draws up the juices of flowers. 334. Most of the lower animals are instinctively directed in the choice of their food by the sense of taste, rejecting what is pernicious, and selecting whatever is adapted to Describe the fungiform papillae ? Of what is the tongue itself made up ? What is the structure of the tongue in birds ? For what other purposes is the tongue of some birds used ? What is said of the tongue in reptiles ?__in ser- pents ?—in the frog, and the chameleon ? What of the tongue of the fish and the bee ? How are most of the lower animals directed in the choice of their food ? PLATE XIII. ORGANS OF SENSE. Figure 1.----The Dorsum, or Upper Surface of the Tongue.—a, Tho epiglottis, b, The root of the tongue, c, c, Mucous glands, covering the root of the tongue, d, d, d, The large papillaj, arranged in two oblique lines, meeting at the middle of the root of the tongue. Spreading over the rest of the surface are seen the small papillae, in great num- bers and of different shapes and sizes. Figure 2.----The Inferior Surface of the Tongue.—This figure represents the lower side of the tongue laid Open, so as to show the distribution of the nerves, a, The hyoid- bone, to which the base of the tongue is attached, b, b, The stylo-glossus muscles, reaching from tho tongue to the styloid processes of the temporal-bone, e, c. Their action is to draw the tongue backward, d, The hypo-glossal nerve.—Minute filaments of these nerves are spread throughout the tongue and in its papillae. Figure 3.----Organ of Hearing: Vertical Section of the Organ of Hearing.—a, b, e, The external ear. d, Entrance to the auditory canal. /, Auditory canal, e, e, Petrous portion of the temporal-bone, in which the internal ear is excavated, g, Membrane of the tympanum, h, The tympanitic cavity or drum of the ear. (For the relative positions of these bones, see fig. 54, page 167.) i, Cells excavated in the temporal-bone, j, Opening from the ear-cavity into these cells.—On the side of the cavity opposite the membrana tympani, are seen the fenestra ovalis and rotunda, which open into the vestibule. /, The vestibule, k, The Eustachian tube, to, Semi-circular canals, n, Cochlea, o, Auditory nerve, p, Canal, by which the carotid artery enters the skull, q, Part of the glenoid fossa) which receives the head of the lower-jaw. r, Styloid process of the temporal-bone. Figure 4.----A magnified representation of the Labyrinth, laid open, so as to display its Cavity—a, The vestibule, b, The cochlea, e, The partition by which the cochlea is divided into two parts, c, The fenestra ovalis. d, The fenestra rotunda. /, /, /, The semi-circular canals, g, Portion of the temporal-bone. Figure 5.----Capillary Plexus of Papilla of Human Tongue. Fioure 6.---Tongue of a Woodpecker. Figure 7.---Tongue of a Giraffe, LE.C.ICe: -i.LiB THE SENSES. 161 their Avants. Man is guided more by reason than by in- stinct, and his taste is very much influenced by habit. Thus, many substances that are exceedingly disgusting at first, become by use highly grateful. Innumerable in- stances of those acquired tastes are observed in all ages, and in every state of society. Thus, epicures in various stages of civilization make use of meat, fish, and eggs, after decomposition has commenced, or after those articles of food are in what is called a "ripe condition;" and mul- titudes, from all classes of society, chew tobacco, the most disgusting of all Aveeds, as a SAveet morsel. 335. The susceptibility of the organs of taste to pleas- urable sensations, depends very much on the condition of the stomach, even in health. Accordingly, the relish or the pleasure Avith which we partake of any particular arti- cle of food, diminishes as hunger is appeased. If indul- gence of the appetite is persisted in beyond the actual demand for food, nausea or disgust will very soon super- vene, and compel even the glutton to desist. The sympathy ( subsisting between the organ of taste and the stomach, is an important and wise provision, informing us Avhen a sufficient amount of food has been taken. SENSE OF SMELL. 336. Smell is the sense by which Ave perceive odors. Certain bodies seem to possess the property of diffusing through the air in extreme minuteness the particles of which they are composed, and of exciting a peculiar sensa- tion in the organ of smell. Those particles which emanate from odorous bodies are so exceedingly minute, that many How is man guided ? How do substances naturally loathsome come to be agreeable ? What examples are given ? Upon what does the susceptibility of the organs of taste to pleasurable sensations depend ? What ensues if indulgence of the appetite is persisted in beyond the actual demand for food ? What is said in regard to the sympathy of the organ of taste with the stom- ach? What is smell? What properties do certain bodies seem to possess? What is said of the particles which emanate from odorous bodies ? 162 CLASS-BOOK OF PHYSIOLOGY. substances do not seem to lose Aveight by freely imparting their scent to an unlimited quantity of air. Musk, for instance, will impart its scent to a room that is constantly open for many years, without any appreciable diminution of Aveight. But there are other bodies, such as camphor, Avhich lose Aveight by loss of particles from their surface Avhen freely exposed to the air. 337. The special seat of the sense of smell is in the thin and delicate membrane which lines the internal surfaces of the nasal cavities. It is called Schneiderian membrane, from the man who first described its structure. 338. The acuteness of this sense seems very nearly in proportion to the amount of Schneiderian membrane, or to the extent of surface which the nasal cavities present. Hence, in all those animals re- markable for acuteness of scent —as the deer, sheep, and dog —the nose is very large, and its internal surface greatly in- creased by the curiously con- voluted form of the internal or turbinated bones. Fig. 52 is a vertical section of the R§As5IrcAEvTTv.-a, moCuTth°r b°,iosi™, nasal cavities in man, shoAving a&^^^.^E*^ now tnis membrane is increa* j£Zi;^^o«S?SLi sphen°idal ed in surface by the turbinated bones,/; g, h. 339. When air charged with odorous particles passes over the Schneiderian membrane, in the act of ordinary respiration, or in the voluntary efforts of snuffing air, some How long will musk scent a room without any apparent loss of its particles? What is said of bodies which lose weight by loss of particles? Where is the special seat of the sense of smell? What is the membrane called? The acuteness of this sense is in proportion to what ? What is said of the size of the nose in those animais which are remarkable for acuteness of smell? How is the interior surface of the nose greatly increased? Explain the manner in which the odorous particles are made to act on the Schneiderian membrane? THE SENSES. 163 of the particles come in contact with it, and act upon the delicate extremities of the olfactory nerves, with which the membrane is thickly set. If this membrane is either too moist or too dry, the odorous particles make no impression on it. Thus, in colds or in fevers, the sense of smell is almost entirely lost for the time. 340. All animals do not perceive the same odors equally well. Carnivorous animals excel in detecting the odors of animal substances, and of tracking other animals by the scent, but have no apparent sensibility to the odors of plants and flowers. Herbivorous animals are peculiarly sensitive to the latter, but their sensibility to animal odors is much less acute. Some herbivorous animals, however, possess very great acuteness of smell in regard to animal odors. Thus, the deer and the antelope frequently scent the approach of the hunter, and make their escape, unless he steals upon them in a direction contrary to that of the Avind. In this class of animals, the sense of smell seems to be an important means by Avhich they are warned of the presence of their enemies. 341. Herbivorous animals undoubtedly require an acute susceptibility to the odors of plants, to guide them in select- ing their food from a variety of plants, some of which are highly deleterious, if not destructive, to animal life. 342. The wonderful scent of the dog is Avell known. The certainty with which he detects the foot-steps of animals, long after they have passed—the facility with which he traces the progress of his master through crowded streets, recognizing the emanations which his foot has left among all the diversity and multitude of odorous parti- cles—is truly astonishing. In like manner, the deer-hound Under what circumstances do the odorous particles fail to make any impres- sion? Do all animals perceive the same odors equally well? In what do car- nivorous animals excel?—in what herbivorous? What is said of the deer and antelope? Of what special use does the sense of smell seem to be to this class of animals ? How do the herbivorous animals seem to be guided in eelecting their food ? What is said of the scent of the dog ?—the deer-hound ? 12* 164 CLASS-BOOK OF PHYSIOLOGY. pertinaciously pursues his victim, and follows its traces through the herd of its fellows, among which it vainly seeks for protection. 343. Birds and fishes do not in general possess the sense of smell in as high perfection as quadrupeds, though they do not appear to be entirely destitute of it. Many insects are able to distinguish odorous substances at considerable distance; but physiologists are not agreed as to the loca- tion of the organs of smell. 344. Like all the senses, that of smell is greatly improved in acuteness by education. In the blind, especially, it becomes next to that of touch, the sense on which they place the greatest reliance, and by which they distinguish individuals from each other. The Indians of Peru, accord- ing to Humboldt, can ascertain by the smell, in the middle of the night, whether a visitor be an European, an Amer- ican, an Indian, or a jSTegro. Sometimes the smell becomes morbidly sensitive: there are those who sicken at the scent of cheese, or faint at that of a cat. An eminent French physician was exceedingly annoyed during an illness, in which his smell became remarkably acute, by the odor of copper; and after a careful search, the source of his annoyance was found to be a brass pin, which had fallen among the bed-clothes. SENSE OF HEABING. 345. Hearing is the sense by which we perceive sounds. The ear is the special organ of hearing, though all the parts which are found in the higher orders of animals are not essential to constitute an organ of hearing. The essential part of this organ is a nerve, endowed with the peculiar property of receiving and transmitting to the What is said of the sense of smell in birds and fishes ?—what of insects ? How may the sense of smell be improved ? How is it in the blind ? What is said of the Indians of Peru? What examples are given in which this sense has become morbidly sensitive ? What is hearing ? What is the special organ of hearing? What is the essential part of this organ? THE SENSES. 165 brain the impressions derived from the vibrations of the air or Avater Avith which the animal is surrounded. 346. Thus, in some of the loAver animals—as the crab, lobster, &c.—the ear consists of a cavity in the side of the head, lined with a membrane on which the nerve is dis- tributed, and filled with a watery fluid. In some, this cavity is completely shut in by its bony walls, and in others there is a small aperture covered by a membrane, upon which the external medium can act. This cavity, Avhich corresponds with the vestibule in higher animals, is the Avhole organ in lower animals; it is the essential part in all, and is never omitted in the most perfect develop- ments of the organ. 347. All other parts may be regarded as superadded to the vestibule, to render this essential organ more perfectly adapted to the wants of each group of animals. Thus, in the lowest fishes, a single semi-circular canal is superadded to the vestibule; a little higher, there are two; and in all other fishes, three semi-circular canals. In animals which live in the air, there are added other parts, which adapt the organ more perfectly to this element. 348. In man, and most of the mammalia, we find the ear composed of three parts—the external^ the middle, and the internal ear. 349. The external ear is a fibro-cartilaginous appendage, placed on the outside of the head, to receive and conduct the sounds to the interior. In most of the quadrupeds, the external ear is very large, and capable of being turned in any direction at the will of the animal, as in the horse, Describe the ear, as found in the crab, lobster, &c. ? How is it inclosed in some species? With what in higher animals does this cavity correspond? Is this part ever omitted in the most perfect development of the organs? What may all other parts be regarded? What parts are added in fishes? What other pans are added in animals which live in the air? Of how many parts is the ear composed in man and most of the mammalia? What are these parts? Describe the external ear. What peculiarity has it in most of the quadrupeds ? Do men ever popses? the power of moving the external ear? 166 CLASS-BOOK OF PHYSIOLOGY deer, and hare. Some men also possess the power of moving the external ear, though to a very limited extent. 350. Persons who are partially deaf may make use of ear-trumpets, by which the vibrations of sound are col- lected from a greater extent of surface, and thereby act with greater intensity on the drum of the ear; or, in case this is destroyed, on the internal ear. 351. The canal (fig.S, Pl. XHI.) into which the external ear collects the sonorous vibrations, passes inwards until it is terminated by the tympanum or the membrane of the drum of the ear, which separates the external from the middle ear. This canal is about an inch in length, and is protected from the entrance of insects and dust by short hairs, which grow across the tube. 352. In reptiles and birds, there is no external ear, and the tympanum is found on the surface of the head, where it can be easily seen, just back of the eye, as in the frog. In birds, it is found in a slight depression in the head, sur- rounded by a tuft of feathers. In man, it is a rough, tense membrane, stretched across the auditory passage, in a manner similar to the parchment on a drum, with its edges set in a bony groove. 353. The middle ear, which is also called the tympanitic cavity or drum of the ear (h), is an irregular bony cavity, filled with air. It communicates with the external air by means of the Eustachian tube, which terminates in the back part of the throat. The use of this tube seems to be to produce an equilibrium between the air contained in the drum of the ear and the external ear. In severe colds, What may be made use of to collect the vibrations of sound by persons who are partially deaf? What effect does it have on the vibrations of sound? What is said of the canal into which the external ear collects the sonorous vibra- tions? How is it separated from the middle ear? What is its length? How is it protected from insects? Have reptiles and birds any external ear? Where is the tympanum formed in frogs?—in birds? Describe it in man. What other name is applied to the middle ear? With what is it filled ? How does it communicate with the external air ? Where does this tube terminate ? What is its use? What effect do colds sometimes have on this tube? THE SENSES. 167 this tube is frequently obstructed, causing "ringing in the ear," confusion of sound, and partial deafness. a vol Fig. 53.—Bones or Fig. 54.—Cavity or the Tympanum, with the Ear. the bones in their places. 354. Within the drum of the ear (fig. 3, Pl. XIII.) there is a very curious arrangement of small bones, which con- nect the tympanum to the membrane of the internal ear. These bones, which are represented in figs. 53 and 54, are so arranged as to form a continuous chain of bones. When the tympanum vibrates, they move backward and forward in such a manner that the slightest vibration is communi- cated to the membrane of the internal ear. In birds and reptiles, instead of a chain of bones, there is a single bone, Avith one extremity attached to the tympanum and the other to the membrane that incloses the internal ear. This bone'can easily be seen pressing against the tympanum in , the ear of a bird by removing the skin and feathers from the head. 355. The internal ear or labyrinth is composed of three cavities—the vestibule, the semi-circular canals, and the cochlea. What peculiar arrangement within the ear? How are these bones ar- ranged ? How do they move with the vibrations of the tympanum ? What substitute for a chain of bones do we find in birds and reptiles? How can this bone be seen in the ear of a bird ? How many cavities has the internal ear ? 168 CLASS-BOOK OF PHYSIOLOGY. 356. The vestibule (porch or entrance) corresponds with the simple sac that constitutes the whole organ 'of hearing in the lower animals, and is the essential part in all. It is separated from the middle ear by a bony partition, Avhich is perforated by two small holes, called, from their form, foramen ovale (oval opening), and foramen rotundum (round opening). The first is closed by a membrane, on which the stapes, one of the small bones, rests, and the other by a membrane similar to the tympanum. 357. The semi-circular canals are passages running off from the upper part of the vestibule, in the form of three arched tubes. These canals are found in fishes, and in all animals which live in the air. Their use is supposed to be to assist in producing an idea of the direction of sounds. 358. The cochlea, so called from its resemblance to a snail-shell, is a spiral canal, running off from the lower part of the vestibule. It is peculiar to those animals which live in the air, and has been supposed to be the organ by which we judge of the pitch of sounds. It also affords a greater extent of surface for the spreading out of the nervous fibres. 359. All the above cavities of the internal ear are lined with a delicate membrane, on which the extremities of the auditory nerve are minutely distributed, and are filled with a watery fluid, in which filaments of the nerve are also found floating. These terminations of the auditory nerve constitute the real organ by which impressions of sound are received and transmitted to the brain. The* With what does the vestibule correspond ? How is it separated from the middle ear? How is this partition perforated? How are these openings closed ? What are the semi-circular canals ? In what animals are those canals found ? What is their use ? From what does the cochlea derive its name ? What is the form and position of the cochlea ? To what class of animals is it peculiar ? What has been supposed to be the use of the cochlea ? How are the cavities of the internal ear lined ? What is distributed on this membrane ? With what are these cavities filled ? What constitutes the real organ by which impressions of sound are received and transmitted ? How are the vibrations of sound communicated to the nerve ? THE SENSES. 169 vibrations of sound are communicated to the nerve through the medium of a watery fluid. 360. Sound travels through the air at the rate of about twelve and a half miles in a minute; through water, its velocity is four times greater; and the intensity of the vibrations of sound in air and water are in about the same proportion* Hence, the impressions of sound on the nerves Avhich are the immediate seat of hearing are much more intense in the watery fluid, Avhich 'fills the cavities of the internal ear, than they could be if those cavities Avere filled with air. For the same reason, animals Avhich live in the air require an apparatus for hearing more com- plicated than those animals which live in the Avater. Ac- cordingly, we find in those animals which live in the air, the addition of just those parts that Avould communicate the vibrations of sound from the air to the fluid of the internal ear with the greatest intensity. Thus, a tense membrane, like that of the drum of the ear, Avith air on both sides, is better adapted than any other animal struc- ture to receive the vibrations of the air; and the arrange- ment of the bones Avhich connect this membrane with that of the internal ear, is equally well fitted for the office of conducting those vibrations to the fluid of the internal ear. 361. Still, the faculty of hearing will not be entirely lost if the drum of the ear is destroyed; so that the vibra- tions of sound act directly on the membrane of the internal At what rate does sound travel through the air? What is its velocity through water ? What advantage then is gained by the nerves receiving impressions through water instead of air ? Why do animals which live in air require an apparatus more complicated than those which live in water? What parts do we find added in those animals which live in air? What is said in regard to the adaptation of the tympanum and the bones which con- nect it to the internal ear to receive the vibrations of the air? Will the faculty of hearing be entirely lost if the dmm is destroyed ? How will the hearing be affected by its loss? * The difference in the intensity of the vibrations of sound in air and water may be easily perceived by striking two stones together with equal force—first in the air and then under water, with the head immersed. 170 CLASS-BOOK OF PHYSIOLOGY. ear, though it will be greatly impaired, rendering the individual partially deaf. 362. The faculty of hearing may be very much increased in acuteness by cultivation; but this increase depends rather upon the habit of attention to the faintest impres- sions made upon the organ, than upon any change of the organ itself. Thus, the Avatchful Indian, recognizes foot- steps, and can even distinguish the tread of a friend from that of a foe, while his Avhite companion, who lives among the busy hum of cities, is unconscious of such slight sounds. Yet the latter may be a musician, capable of dis- tinguishing the tones of all the different instruments in a large orchestra—of following any one of them through the part which it performs—and of detecting the least discord in the blended effects of the whole—effects which would be to the Indian only a confused mass of sound. SENSE OF SIGHT. 363. Sight is the sense by which we are enabled to perceive luminous impressions; and through these we be- come acquainted with the form, size, color, and position of objects that transmit or reflect light. 364. The eye is the organ of sight. It consists essen- tially in an instrument capable of making a distinct picture of surrounding objects on the expanded surface of the nerve of sight. 365. Some of the inferior animals, such as the leech, seem to be guided in their movements by certain dark spots on their surface, supposed to be rudimentary eyes, though no fully developed eyes have as yet been dis- covered. In some of the star-fishes those eye-spots are found at the extremity of the rays, as in fig. 1, Pl. XYI. How may the faculty of hearing be increased in acuteness? Upon what does this increase seem to depend? What examples are given of the habit of attention? What is sight? What is the organ of sight? In what does it consist essentially? How do some of the inferior animals seem to be guided in their movements ? PLATE XIY. ORGANS OF VISION-THE ETE. Figure 1.----The Left Eyeball, showing the Posterior Surface of the Retina.- a, A small transparent spot, situated near the optic nerve, called the foramen of Soemmering, surrounded by a yellow halo, the limbus luteus. b, The optic nerve, cut off at its entrance into the retina, c, The central artery of the retina, d, d, Ramifications of the artery on the inner wall of the retina, seen through the outer layers. Figure 2.----The Globe of the Eye, magnified, and seen in front.—This figure repre- Bonts the second or choroid coat, the sclerotica being removed, a, The pupil, b, The iris. c, c, The ohoroid membrane, d, d, The ciliary nerves, running from every part of the cir- cumference towards the iris, e, e, The ciliary arteries. Figure 3.----The Globe of the Eye, seen in the same view.—The iris is removed, in order to display the ciliary processes, and their position in regard to the lens. The ciliary nerves and arteries are the same as in fig. 7. Figure 4.----The Anterior Half of the Globe of the Eye, seen from behind.—The lens and the vitreous humor are removed, a The pupil, b, b, The posterior surface of the iris, called, from its dark grape-like color, uvea, c, c, The ciliary processes, d, d, The internal surface of the choroid and sclerotic, Figure 5.----The Posterior Half of the Globe of the Eye, showing its internal surf ace. —a, a, The cut edges of the sclerotic, choroid, and retina, b, b, The internal surface of the retina, e, The foramen of Soemmering, d, The optic nerve, e, Branches of the central artery of the retina. Figure 6.----The Fibres of the Iris, detached.—a, The pupil, b, The circular fibres at the central margin of the iris, by the action of which the pupil is diminished or enlarged. e, c, The radiating fibres, which proceed from the external border of the circular fibres. Figure 7.----A Portion of the Pigment Membrane of the Choroid Coat, highly magni- fied..—-It is seen to consist of regular six-sided plates, the tissue of which is filled with grains of coloring matter. Figure 8.----Compound Eyes of the Bee, highly magnified, showing the Division into Facets, and also the Conical Shape of each separate Portion.—A, Facets still more highly magnified. B, The same with hairs growing between them. Figure 9.----A, Front View of the Crystalline Lens. B, Side View of the Lens.—a. Its anterior and least convexity, b, Its posterior and greatest convexity. Figure 9.----Front View of the Crystalline Lens. Figure 10.----Side View of the Crystalline Lens.—a, Its anterior and least convexity, b, Its posterior and greatest convexity. Figure 11.----The Crystalline Lens, after being immersed in boiling water.—The lines on its surface show its division into three parts. Figure 12.----The Three Segments of the Crystalline Lens.—The faces of the segments show the concentric layers of which it is composed (like the coats of an onion). The nucleus, or central portion of the lens, is seen on one of the segments, and on the other two are corresponding depressions. Fi 2-1 Fi<5. 6 Fig. 7 Fl6.ll. E. AUooA.Lith. Ry.1'2. .:i.-ulford C. \ THE SENSES. 175 366. Most insects are furnished with compound eyes, AA'hich consist of several hexagonal facets, (fig. 8, Pl. XIV.) united together in such manner as to form a large, dark- colored protuberance on each side of the head. Each of these facets is an eye in itself. In some insects, these facets are exceedingly numerous. In the eye of a butter- fly, 17,000 have been counted; in some species of beetle, 25,000; in the common horse-fly, 4,000. These numer- ous eyes, no tAVO of which have the same direction, seem to compensate for the inability of the insects to move the head without moving the whole body, since they are pro- vided Avith an eye in every direction. 367. Spiders are furnished with from six to fourteen eyes, situated on the most prominent part of the back, instead of on the sides of the head. (Fig. 3, Pl. XYI.) 368. In the snail and some other mollusks there are only two eyes, situated on or near the tip of a flexible stalk—the tentacula, (fig. 2, Pl. XYI.) which can be extended in any direction at the pleasure of the animal. In lobsters and crabs the eye is placed at the extremity of a long tube, which can be moved in various directions, and extended or withdrawn. 369. In all the higher animals there are only tAvo eyes, Avhich are constructed on the same general plan, with some variation in the form of particular parts, adapting the eye to the medium with which different animals are surrounded. 370. The form of the eye is nearly globular in all, and placed in a bony socket (the orbit) in the head. The orbit is lined with a cushion of fat, on which the eye rests and rolls with very great ease. When a person becomes very With what are most insects provided ? Of what are those compound eyes made up? What is each facet? How numerous are these facets? How many in the eye of a butter-fly—in the beetle? What seems to be the use of so large a number? With how many eyes are spiders provided? How many eyes are there in the snail, and how are they situated ? How is the eye placed in crabs and lobsters ? How many eyes are there in all the higher animals ? What is the form of the eye, and how is it placed ? How is the orbit lined ? 176 CLASS-BOOK OF PHYSIOLOGY. much emaciated, this cushion of fat is gradually absorbed, alloAving the eye to recede into the orbit. 371. The eye is admirably protected against insects, dust, and other foreign substances, by the eye-brows, eye- lids, and eye-lashes. 371. In birds and reptiles there is a thin transparent membrane which is draAvn across the eye by a muscle that passes through a loop in the membrane, (fig. 4, Pl. XYI.) This nictating membrane is so transparent as not to prevent the power of vision, while it protects the eye from too strong light, and guards it against foreign bodies. 373. The horse is provided with a beautiful contrivance for protecting the eye, called the haw. It is a triangular- shaped cartilage, admirably adapted to the convexity of the eye, and so arranged that it is made to sweep across the eye at the will of the animal, and shovel up any dust or insect that may fall upon it. 374. The anterior surface of the eye is covered by the conjunctiva—a thin mucous membrane, which is reflected upon the lids, so as to form their internal surface. 375. The conjunctiva is constantly moistened by the mucus from its surface, and by the tears that are secreted in the lachrymal glands, and poured upon the inner sur- face of the upper lid by seven or eight tubes. The lach- rymal gland (fig. 2, Pl. XY.) is situated at the upper or outer angle of the orbit. It is continually pouring out a watery fluid, which moistens the globe of the eye, and keeps it free from impurities. This secretion is increased by mental emotion, and by irritation of the eye. The tears pass out from the eye by tAVO lachrymal ducts, Avhich What takes place when a person becomes very much emaciated ? How is the eye protected against insects, dust, &C. ? What membrane is there in birds and reptiles? How is it drawn across the eye ? How does it protect the eye without preventing sight? With what is the horse provided? De- scribe the haw ? How is the interior surface of the eye covered ? How is the conjunctiva moistened? Where is the lachrymal gland situated ? What is continually pouring out from this gland ? How is this secretion increased 1 How do the tears pass out of the eye ? PLATE XV. ORGAN OP VISION.-THE EYE. Fioure 1.----Both Eyes, with their Muscles, as they appear upon a horizontal section through the orbits, immediately above the eyes—as if the upper part of the head were removed, as far down as the top of the eyes, the observer looking at them from above. They are represented in this position, in order to show clearly the situation and action of the superior oblique muscle, and also the crossing or decussation of the optic nerve. The letters of reference indicate the same parts in figures 1 and 3. a, The optic nerve, s, figure 1, The chiasma or commissure of the optic nerve, whence each nerve extends forward and outward, passes into the eye at t, and becomes continuous with the retina, b, The common oculo-motory nerve, which is distributed to five of the muscles of the eye. k, Trunk of the fifth pair, a branch of which constitutes the ophthal- mic nerve, and gives sensibility to the different parts of the eye.—The distribution of this nerve is seen in the left eye in figure 1. /, Artery of the eye. c, The elevator muscle of the upper eyelid, d, The superior rectus, or elevator of the eye. «, The inferior rectus, or depressor of the eye. /, The internal rectus, g, The external rectus, h, figure 1, The superior oblique muscle. ?n, Its pulley, i, figure 3, The inferior oblique muscle. In figure 1, parts of several of these muscles are removed, in order to display the others distinctly. Figure 2.----The Lachrymal Apparatus.—a, The lachrymal gland, b, b. The lachry- mal ducts, which collect the tears, and transmit them to the lachrymal sac, c, whence they pass into the cavity of the nose. Figure 3.----The Right Eye, with its Muscles, displayed in the cavity of the orbit, on the vertical plane of a section corresponding to the middle of the arch of the eyebrow :— that is, as if the right side of the head were removed, as far as the middle of the right eye- brow, leaving the eye in its place, to be seen from the right. EL. TV. J! Harlfonl THE SENSES. 181 * commence in a small opening in the edge of each lid, near the corner of the eye, and unite to form the lachrymal sac that lies upon the side of the upper parts of the nose. From the lachrymal sac a canal passes down to the interior of the nose, Avhere the tears pass off in vapor Avith the breath. 376. The Avails of the eye are composed of three coats: the sclerotic, the cornea, and the choroid coat. Its interior is occupied by three humors: the aqueous, the crystalline, and the vitreous. 377. The sclerotic, (fig. 55,) so named from its firmness and density, constitutes about four-fifths of the globe of the eye. It is a tough fibrous structure, admirably adapted to protect and support this delicate organ. Fig. 55.—Interior or the Eye. Fig. 56.—Section of the Eye. c, cornea; s, sclerotic; s, portion of the sclerotic turned back to show the subjacent parts; ch, choroid ; r, retina; n, optic nerve; ca, anterior chamber; i, iris; p, pupil: cr, crys- talline lens; cp, ciliary processes; v, vitreous humor; b, b, conjunctiva. 378. The cornea (c, fig. 55) is a dense transparent struct- ure, convex in front and concave posteriorly. It resem- bles a Avatch-glass in form, and is received into a circular groove in the margin of the sclerotic, in the same manner that a watch-glass is inserted into the case. It occupies about one-fifth of the front part of the eye, and projects forward beyond the sclerotic. In some individuals and Where is the lachrymal sac situated, and how do the tears escape from it ? Of what are the walls of the eye composed ? How is its interior occupied? Describe the sclerotic. Describe the cornea. How is it received into the sclerotic? What portion of the eye does it occupy ? What is said in regard to its prominence at different periods of life and in different animals? 182 CLASS-BOOK OF PHYSIOLOGY. in some animals it is more prominent than in others, and it is also more prominent at some periods of life than at others. In fishes, and in some water-birds, it has only a slight projection forward, as represented in figs. 5 and 6, Pl. XYI. 379. The choroid coat, (ch, fig. 55,) which is a thin, delicate structure, consisting mostly of blood-vessels and nerves, lines the interior of the sclerotic. The internal surface of the choroid is covered by a layer of black pig- ment cells. (Fig. 7, Pl. XIY.) / """380. The aqueous humor, which occupies the anterior / chamber of the eye, (fig. 56,) is nearly pure water. It 1 gives prominence to the cornea, causing it to be more or 'sLess convex, according to the amount of this fluid. r 3,81. The vitreous humor, which resembles thin jelly or f melted glass in consistence, occupies the greater part of ! the globe of the eye behind the iris. V 382. The crystalline humor resembles thick jelly or soft gristle. It has the form of a double-convex lens, Avhence it has received the name of crystalline lens. In those animals which have only slight projections of the cornea, it is nearly spherical. (Figs. 5 and 6, Pl. XYI.) It is suspended in its place by the ciliary process, (fig. 3, Pl. XIY.) a set of little bands from the choroid coat. 383. The aqueous and vitreous humors are separated from each other by the iris, (fig. 2, Pl. XIY.) a kind of curtain, which divides the anterior from the posterior phamber of the eye. The iris receives its name from the great variety of its colors. It is perforated in the centre by an opening, called the pupil. (Fig. 2, Pl. XIY.) In man, this pupil is round, and dilates or contracts according to the amount of light and the sensibility of the optic nerve. What fluid occupies the anterior chamber of the eye ? How does it affect the form of the cornea ? Describe the vitreous humor—the crystalline humor. What is the form of it ? How is it suspended in its place ? How are the aqueous and vitreous humors separated from each other ? From what does the iris receive its name ? What opening has the iris in the centre? What is the form of the pupil in man ? THE SENSES. 183 When the light is very strong, it is but a speck, and enlarges to nearly half the size of the iris in the dark. In birds, especially in oavIs, its motions are more free and extensive than in man. 384. The blackness of the pupil is owing to the dark color of the internal surface of the choroid. In Albinoes, which are destitute of coloring matter, the pupil, as well as the iris, are a bright red, from the numerous capillary blood-vessels of the choroid. In many quadrupeds—as the ox, the lion, and the cat—a portion of the surface of the choroid is covered with a bluish layer of bright metallic lustre, by Avhich the light is brilliantly reflected, when the eye is seen in certain directions, causing the eyes to appear in an obscure light like two balls of fire. 385. In animals whose range of vision is required to extend widely in a horizontal direction—as in the deer, cow, &c.—it is lengthened horizontally, so as to give a wide side-view. In the carnivorous animals—as the lion, cat, &c.—which watch their prey in situations either above or below them, the pupil is elongated vertically. 386. Inside of the choroid coat, and at the back part of the eye, is the retina, which consists of a delicate film of nervous fibres, spread out from the optic nerve as soon as it has passed through the sclerotic and choroid coats. 387. The globe of the eye is moved by six muscles— four recti or straight, and two oblique—all of which are lodged in the orbit. The superior rectus or straight muscle (fig. 3, Pl. XY.) rolls the eye upward, and the inferior rectus turns it downward. The internal rectus rolls the globe inward, or toward the nose, and the external How does it vary in size ? To what is the blackness of the pupil owing? What is the color of the pupil and iris in albinoes? What peculiar appearance has the eye in many quadrupeds? What is the form of the pupil in grazing animals?—in carnivorous animals? What is situated inside of the choroid coat? Describe the retina. How is the globe of the eye moved? How does the superior rectus move the eye ?—the inferior rectus ?—the internal rectus?—the external rectus? 184 CLASS-BOOK OF PHYSIOLOGY. rectus turns it outward. The superior oblique, a very remarkable muscle, has its origin at the back of the orbit, passing forAvard through a little cartilaginous pulley, and then turning backward, to be inserted into the sclerotic coat. The direction of its action is thus changed, like that of a rope which is passed through a block in an ordinary pulley. The use of this muscle is to roll the eye downward and inward. The inferior oblique turns it upward and outward. 388. The coats and humors of the eye seem admirably adapted to modify the rays of light, which fall on the nerve of sight in such a manner as to make the impression clear and distinct. All the parts of the eye are constructed in such strict accordance with the laws of light, that it excels all other optical instruments in perfection and ac- curacy. Human skill has thus far effected only an imper- fect imitation of the instrument designed by Infinite Wisdom. 389. It is well knoAvn that Avhen the rays of light pass obliquely from the air through a dense medium, they are refracted or bent out of their course, as in fig. 7, Pl. XYI., a and that the refraction is in propor- tion as the rays fall more or less obliquely. Thus, if a piece of money be placed in a cup (fig. 57) in such a position that the side of the cup will just hide the money from sight, and then let the cup be filled with Fig- 57. water, the money will be distinctly visible from the refraction of the rays of light. For the Describe the superior oblique ? What is the use. of this muscle ? In what direction does the inferior oblique turn the eye ? To what do the coats and humors of the eye seem adapted? How does the eye compare with optical instruments ? How are the rays of light affected by passing obliquely from a rarer to a denser medium ? The refraction is in proportion to what ? What examples are given, illustrating the refraction of the rays of light in passing from air to water ? PLATE XVI. ORGANS OF VISION-THE EYE. Figure 1.---Eye-spots of a Star-fish, at the extremities of the radii. Figure 2.---Eyes of a Snail, on the extremities of the tentacular Figure 3.---Eyes of a Spider, on the anterior part of the back. Figure 4.---Nictating Membrane of a Bird.—A, Nictating membrane, covering one- half the suiface of the eye. B, The muscles bjt which the membrane is drawn across the eye. The muscle a forms a loop at b, through which the muscle c acts as through a pulley by its tendon d, which is inserted into the edge of the nictating membrane at c. Figure 5.---Eye of a Fish, with very slight convexity of the cornea, and the lens nearly spherical, so as to compensate for the want of refractive power in the anterior part of the eye. Figure 6.---Eye of a Duck, with a similar formation. Figure 7.---Refraction of Light.—a, The ray of light falling obliquely on a dense me- dium at b, is refracted to c, instead of pursuing its original course to d; when it passes into a rarer medium again at c, it is again refracted in a new course down to d, instead of/. Figure 8.---The Lenses.—a, Single convex lens, b, Single concave lens, c, Double convex lens, d, Double concave lens, e, Concavo convex lens. Figure 9.----Short-sightedness—The image formed in front of the retina. Figure 10.---Long-sightedness—Tho image formed back of the retina. Figure 11.---Eye of a Lynx.—a, Ciliary process, by which the lens, b, is moved backward and forward, to adapt it to different distances. Figure 12.---Eye of an Eagle.—a, Fan-like muscle, attached to the lens, to change its position. Figure 13.---Eye of an Owl, surrounded by a kind of bony case, which causes the anterior portion of the eye to become more, and the posterior portion less convex, when- ever the eye i* drawn back into the socket. By this means the focal distance is changed, and the eye adapted to different distances. PL. XVI THE SENSES. 189 same reason an oar or a stick, when partly immersed in water, will appear bent in proportion to its obliquity. 390. The rays of light which fall upon the cornea of the eye are refracted toAvards the pupil, both by the dens- ity of the cornea and aqueous humor, and by the con- vexity of the cornea, Avhich causes the rays to fall on it more obliquely. 391. After passing through the pupil, the rays of light continue to be refracted by the crystalline lens and vitre- ous humor, so that they meet in a focus on the retina, Avhere a complete inverted image or picture of the object is thus formed. This is shown in fig. 58, where, for the sake of conve- nience, two rays only are represented as issuing from each of the two extremities of an object, a, c. Those rays cross each other in the middle of the eye—those from a being brought to a focus at b, and those from c at d; and as all the other rays are refracted in the same man- ner, a complete inverted picture of the object is formed at the back of the eye. 392. The inverted image may be easily seen by remov- ing the fat and muscle from the eye of a white rabbit, and bringing a lighted candle in front of it, when an inverted image may be distinctly seen on the retina through the transparent coat of the sclerotic; or it may be seen by removing the sclerotic from the back part of the eye of a sheep or an ox. In either case, the eye should be fresh, as the cornea and humors soon lose their transparency, and the image becomes indistinct.* How are the rays of light refracted towards the pupil ? How are the rays of light affected by the crystalline lens and the vitreous humor? What is the position of the image on the retina ? Describe Figure 53. How may the inverted image be seen ? * To show the retina of the eye, clip off the muscular and adipose substance about it, then describe with the knife a circle about the 14 190 CLASS-BOOK OF PHYSIOLOGY. 393. The black pigment, which is situated in the inter- nal surface of the choroid, and immediately "behind the retina, absorbs the rays of light as soon as they have passed through the retina, and prevents them from being reflected from one part of the eye to another, causing confusion and indistinctness in the picture. The black pigment also diminishes the intensity of the impression of light on the retina. In Albinoes, (in whose eyes the pigment is defi- cient,) vision is not only extremely imperfect, but the strong sunlight is even painful to the eye. Hence, Albi- noes can see most clearly when twilight comes on, or during cloudy days, or by moonlight. 394. The ordinary forms of defective vision, known as short-sightedness and long-sightedness, are caused by defects in the refractive poAver of the eye. 395. In short-sightedness, the refractive power is too great, causing the rays to be brought to a focus forward of the retina, unless the object be held very near the eye, so as to increase the angle at which they fall on the cor- nea. (Fig. 9, Pl. XYI.) In most cases of. short-sighted- ness, the cornea is too convex from an excess of the aqueous humour. 396. Short-sightedness is frequently caused in students What is the use of the black pigment on the internal surface of the choroid ? How, does the light affect the eye in Albinoes ? How are short and long- sightedness caused? What is the defect in short-sightedness? What is the form of the cornea in short-sightedness? How is it frequently caused ? entrance of the optic nerve that shall include one-fourth of the ball of the eye; holding the eye in the thumb and fingers of the left hand, gradually and carefully cut through the sclerotic to the choroid coat, at a single point in the circle already described, and then complete the dis- section in water by carefully insinuating the sharp point of the scissors between the coats, and clipping round the circle. The slight attach- ment of the choroid to the sclerotic may now be separated with the back of the knife till the detached portion of the sclerotic can be clipped off, leaving the nerve entire. The dissection is now complete; and, if well performed, inverted images of objects can be seen on the retina as in life. THE SENSES. 191 and artisans by the habit of holding objects near the eyes, till they adapt themselves to a short focal distance, and it may be remedied by perseverance in the opposite practice. Thus, sailors, from the long habit of using their eyes in search of distant objects, acquire the ability of recognizing objects at a distance that would be unobserved by a landsman. 397. If short-sightedness has continued from birth, it can seldom be remedied, except by the use of concave glasses, the curvature of which compensates for the excess of the convexity of the eye. If the glasses' used be toe concave, they will increase the difficulty; while those that are not quite concave enough, will have a tendency to remedy it. If glasses can be dispensed with, or if they are* changed for those less concave every few years, the difficulty will gradually diminish, till, in advanced life, the eye assumes the natural form. 398. In long-sightedness, (fig. 10, Pl. XYI.) which is commonly met with in persons in advanced life, the re- fractive power of the eye is not sufficient to bring the rays to a focus on the retina, but back of it, unless the object be held at a long distance from the eye. At this period the eye is not sufficiently convex, from the decrease of its fluids. This difficulty may be remedied by perseverance in the habit of holding objects as near the eye as possible, or wearing spectacles with convex glasses, which shall compensate for the deficiency of the convexity of the eye. 399. The eye is the most delicate of the organs of sense, and is more liable to disease than any other. In many persons the conjunctiva is exceedingly sensitive, and How may short-sightedness be remedied ? What power do sailors acquire by the habit of viewing objects at a distance ? What kind of glasses may be used to remedy short-sightedness? What will be the effect of using glasses that are too concave ?—not quite concave enough to compensate for the con- vexity of the eye ? How does the eye gradually return to its natural form ? What is the cause of long-sightedness? What change in the eye diminishes its convexity? How may the difficulty be remedied ? What is said in regard to the liability of the eye to become diseased? 192 CLASS-BOOK OF PHYSIOLOGY. becomes inflamed from very slight causes. Exposure to dust, to Avind, or to a strong light, not unfrequently induces severe inflammation. The optic nerve is also liable to serious injury from exposure to a strong light, from too pro- tracted application to study, or from using the eyes with insufficient light. Disease of the eyes may be prevented by a careful regard to their strength, by refraining from their use in improper light, and by affording them rest as soon as a sense of fatigue begins to be experienced. Moderate ablution at bed-time, and liberal washing in cold water on rising in the morning, will be found of great service in promoting strength and vigor. CHAPTER XIII. ANIMAL MOTION. 400. The power of voluntary motion is characteristic of all animals. 401. The organs of voluntary motion are the bones and muscles. 402. The bones constitute the frame-work of the body, and give strength and firmness to the entire organization. 403. The muscles form the greatest part of the mass of the body, and constitute what is commonly known as flesh or lean meat. 404. The principal hard parts in animals are shells, crusts, and bones. Shells are almost destitute of animal matter, being nearly the same in composition as a piece of marble. Crusts, such as the covering of the lobster, con- tain a considerable quantity of animal matter, though less than is found in bones. What parts are liable to become inflamed or t"» be injured by improper use ? What causes induce disease ? How may disease be prevented ? What power is characteristic of all animals? What are the organs of voluntary motion? What do the bones constitute?—what the muscles? What are the principal lard parts in animals ? What is the composition of shells ?—of crusts ? PLATE XVII. ORGANS OF MOTION.-THE BONES. Figure 1.----Front View of the Human Skeleton.—At the right half of this figure, the bones are represented in their natural connexions, and divested of all covering. At the left side, the joints are covered by their ligaments. The outer lines show the form of the body when covered with flesh. The Head.—a, The frontal-bone, b, The parietal-bone, c, The temporal-bone. e,The superior maxillary-bone. /, The malar-bone, g, The nasal bones, h, The vomer, i, The inferior maxillary-bone, k, The orbits. /, /, Sutures. The Trunk—1, 1, The spinal column. 2, The sternum. 3, 3, The ribs. 4, The oa innominatum, or haunch-bone. 5, The sacrum. The Superior Extremities.—6, The clavicle. 7, Acromion process of the scapula, which articulates with the clavicle. 8, The humerus. 9, The elbow-joint. 10, The radius. 11, The ulna. 12, Bones of the carpus. 13, Bones of the metacarpus. 14, The phalanges. The Inferior Extremities.—15, The hip-joint. 16, The femur, or thigh-bone. 17, The patella. 18, The tibia. 19, The fibula. 20, Bones of the tarsus. 21, Bones of the metatarsus. 22, Bones of the toes. On the left side of the figure, I indicates the ligaments of the shoulder, m, Ligaments of the elbow, n, Ligaments of the wrist, s, Ligaments of the hip-joint, o, Ligaments of the knee, and tendon of the extensor muscle of the leg. p, Ligaments of the ankle, q, Large vein and artery of the arm. r, Large vein and artery of the leg. Figure 2.---A Bone deprived of its earthy portion by maceration in a dilute acid, and tied in a knot, to show its flexibility. Figure 3.---A Section of the Femur, showing its cancellous structure. Figure 4.----A Dorsal Vertebra.—a, The body of the vertebra, b, The spinal foramen. c, c, Articulating processes, d, d, Transverse processes, e, Spinous processes. Figure 5.----A Lumbar Vertebra, to show the greater thickness and strength of its body and processes. Figure 6.----The Lower Jaw.—a, The condyle, which articulates with the temporal- bone, b, The coronoid process, c, The ramus. Figure 7.----Vertebra of a Fish.—9, End view. B, Side view of the same. Figures 8 and 9.----These figures are designed to show the minute structure of bone. Figure 8 represents a cross-section of bone, highly magnified, a, a, Orifices of small tubes, called, from their discoverer, Haversian canals. They usually run in the direction of the length of the bone, the membrane lining the hollow of which is prolonged into their canals. This membrane contains innumerable small blood-vessels, and the interior of the bone is thus supplied with blood. Around each of the Haversian canals are seen concentric circles of some small dark spots, which are found to be flattened cavities or bone-cells, from which proceed numerous minute tubules. These open into the sides of the Haver- sian canals, and communicate from one bone-cell to another, thus transmitting the nourishment with which they are supplied by the blood-vessels throughout the substance of the bone. Figure 9, represents a longitudinal section of bone, highly magnified, showing the Haversian canals seen lengthwise, their connexion with each other, and the direction of the bone-cells. *These figures are from HassalCs Microscopic Anatomy. pl xvn ICeTlogS Iith Hartford, Conn. ANIMAL MOTION. 197 405. At first, bones exist in a state of cartilage, and are gradually converted into bone by a deposition of phos- phate and carbonate of lime. 406. The lime of the earthy portion of the bones is continually increasing till old age, while the animal por- tion is gradually diminishing. In children, the animal matter constitutes about one-half; in adults, one-fifth; and in old age, one-eighth of the whole composition. 407. In children, the bones are soft and flexible, and admirably adapted to sustain the numerous falls and acci- dents, to which they are liable, without injury. At this age, the bones are not easily broken, though they are bent and twisted from their natural form. Thus, infants are not unfrequently made "bow-legged," by efforts to bear their weight on their limbs before they have acquired the proper proportion of earthy matter to give them the requi- site strength. If not encouraged by parental ambition, nature does not incline the young infant to make the effort to stand or walk until the period when the bones have become quite stiff and hard. 408. The bones do not arrive at their perfect state until about the twentieth year. Previous to this period, the form of the bones may be easily changed by improper habits. Indeed, some change may be effected in many of the bones at a much later period of life. 409. As the animal matter of the bones diminishes in old age, they become hard and brittle. In an aged per- son, a very slight slip or miss-step is sufficient to produce a fracture. Fortunately, the failure and decline of all the other powers compel this class of persons to move with so In what state do bones exist at first ? How are they converted into bone ? Which portion of the bones continues to increase and which to diminish to old ase? What proportion of the bones is animal matter in children 7—in adults?-in old age? What is the condition of the bones in children ? To what injury are the bones most liable at this age ? How are infants made bow-legged ? At what age do the bones arrive to perfection ? What is the cTdittaf of the bones in old age ? To what injury are the bones of the aged liable ? 198 CLASS-BOOK OF PHYSIOLOGY. much care and caution, that the frailty of this part of the system is but seldom tested. 410. The earthy and animal portions of the bones can be easily separated from each other. To obtain the ani- mal portion, we have only to soak the bone a few days in a diluted acid. When the earthy matter is then dissolved, the animal matter which is left becomes so soft and plia- ble, that a bone of sufficient length can easily be tied into a knot, as represented in fig. 2, Pl. XYII. 411. By exposing a bone to the action of a hot fire, the animal matter will be consumed, leaving it so brittle that it may be easily crumbled into fragments. 412. The bones of nearly all the higher animals present a great variety of form and structure, according to the position they occupy. 413. The principal bones of the extremities are long and cylindrical, and consist of a shaft and two extremities. The shaft of a long bone is dense and hard in structure, and hollow in the centre, forming a cylinder or a double arch—a form which, it is well known, receives the greatest amount of strength. The extremities of the long bones are broad and expanded, so as to present a large surface to articulate with adjoining bones, and their internal struc- ture is cellular and cancellous, so as to secure as much bulk as possible in proportion to the quantity of matter. The internal structure of the long bones may be seen by a" sec- tion of the thigh-bone, fig. 3, Pl. XVII. 414. The different bones of the body are united to each other by articulations and joints. 415. The different bones of the cranium and skull, which surround and protect the brain, are very firmly united by How can we separate the animal from the earthy portion of the bones ? How the earthy portion? What is said of the bones of the higher animals? What is the form of the principal bones of the extremities? Describe the shaft, and the extremities of a long bone. How are the different bones of the body united to each other? How are the different bones of the cranium united together ? PLATE XVIII. ORGANS OF MOTION.-THE BONES. 1 Figure 1.----Posterior View of the Human Skeleton.—The Head.—a, The frontal-bone. b, b, The parietal bones, c, The left temporal-bone, d, The occipital-bone. The Trunk.—1, 1, 1, Spinous processes of the vertebrae, t, t, t, Transverse processes j of the vertebra;. 2, 2, The ribs. 3, The os innominatum, showing its three parts, viz: a, '] the illium ; b, the ischium ; c, the os pubis. 4, The sacrum. 5, The coccyx. -I The Superior Extremities.—0, The scapula. 7, Acromion process of the scapula. 8, The clavicle. 9, Head of the humerus placed in the glenoid cavity of the scapula. 10, | Shaft of the humem?. 11, 12, Internal and external condyles of the humerus. 13, The olecranon process of the ulna, which articulates with the pulley-like surface at the lower J end of the humerus, forming the elbow-joint. 14, Shaft of the ulna. 15, Shaft of the | radius. 16, Lower extremity of the radius, which articulates with the bones of the carpus. . j 17, Bones of the carpus. The first row consists of four boms, viz: a, the scaphoid-bone ; ; b, the semi-lunar-bone; c, the cuneiform-bone; d, the pisiform-hone. The second row I consists of four bones, viz: c, The trapezium. /, The trapezoid-bone. g, The os magnum. | ft. The unciform-bone.—18, 18, Bones of the metacarpus. 19, 19, 20, 20, 21, 21, First, sec- • Olid, and third ranges of finger bones. The Inferior Extremities.—22, Head of the femur, placed in the acetabulum or | cotyloid cavity of the haunch bone. 23,24, Projections called trochanter major and minor, to which the muscles of the hip are attached. 25, Shaft of the femur. 26, 27, External and internal condyles of the femur. 28, Upper extremity of the tibia, which articulates with the lemur. 29, Shaft of the tibia. 30, The internal malleolus, a projection of the tibia which forms the inner ankle. 31, The fibula. 32, External malleolus, a projection ,. of the fibula which forms the outer ankle. 33, Bones of the tarsus. 34, Bones of the i metatarsus. 3.3, Bones of the toes. The ligaments of the various joints are seen on the j left side of this figure. Figure 2.----The Knee-joint—A, The knee-joint, with the patella removed, a, a, The ' condyles of the lemur, covered with cartilage, b, b, The two semi-lunar cartilages, which form cup-shaped depressions for the reception of the condyles, c, The anterior crucial ligament which passes from the tibia to the femur. B, Section of the knee-joint, showing ! the reflections of the synovial membrane, a, The cancellous structure of the lower part ] of the femur, b, The tendon of the extensor muscle of the leg. c, The patella, d, The , ] ligaments which attach the patella to the head of the tibia, e, The cancellous structure | of the head of the tibia. /, The anterior ligament, g, The posterior ligament. The synovial ! membrane may be traced along the under surface of the patella and its ligaments, and then over the head of the tibia, over the posterior ligaments, and then over the lower part of the femur. Figure 3.----A Section of the Hip-joint.—a, The head of the femur, b, b, The capsular • ligament, embracing the cavity of the hip-bone and the head of the femur, and keeping both bones firmly together, c, A round ligament attached to the inside of the cavity and to the head of the femur. Figure 4.---The Bones of the foot, seen upon the upper surface.—a, The OS calcis, or heel-bone, b, The astragalus, which articulates with the lower end of the tibia, c, The cuboid-bone, d, The scaphoid-bomi. c, e. f, Cuneiform bones. ANIMAL MOTION. 203 sutures (seams), which are formed by the interlocking of the ragged edges of one bone into corresponding notches in the adjoining one. These bones are thus so firmly united together in man at adult age, that it is difficult to separate them, except by breaking away some of the pro- jecting parts. 416. The bones forming joints are firmly bound together by muscles and ligaments, and the end of each bone cov- . ered over by a thin layer of cartilage, which has a smooth glassy surface. Each joint is inclosed by a sac or capsule of serous membrane, which secretes a peculiar fluid, called synovial fluid. The office of the synovial fluid is to keep the joints constantly moist and supple. 417. After a severe injury to the joint, the synovial fluid is sometimes secreted in excess, causing " dropsy of the joint." 418. The beautiful smoothness of the surfaces of the cartilages, and the manner in which the bones are bound ! together by ligaments and muscles, may be seen by ex- amining the knee-joint of any of our domestic animals. | In fig. 2, Pl. XVIII., is a section of the knee-joint in man, showing the reflection of its synovial membrane. 419. There are several kinds of joints in the body, the most important of which are the hinge-joint at the elbow and knee, and the ball-and-socket joint at the hip and shoulder. 420. In the hip-joint, the socket is much deeper than at the shoulder—an arrangement evidently designed to give greater security against dislocation in a part on which the weight of the whole trunk must press in walking, run- How are the sutures formed? What is said of their firmness at adult age? How are the bones forming joints bound together? How is the end of each bone covered? How is each joint inclosed? What is the office of the syno- vial fluid? How is this fluid sometimes affected by an injury? How may the structure of a joint be seen? How many kinds of joints are there? What is said of the hip-joint ? What circumstance seems designed to pre- vent dislocation at the hip-joint ? 204 CLASS-BOOK OF PHYSIOLOGY. ning, leaping, &c. There is also a strong cord, fastened by one end to the top of the thigh-bone, and by the other to the socket in which it moves. 421. The skeleton of man is formed by the union of about two hundred and fifty bones, and is divided into head, trunk, and extremities. 422. The head includes the bones of the face and the cranium; the trunk includes all the bones immediately attached to the spine, except the head; the extremities embrace all the bones of the shoulders, arms, and legs, which are called upper and lower extremities in man, or the anterior and posterior extremities in quadrupeds. 423. The face is formed by the union of fourteen bones, which serve for the lodgment and protection of the organs of sight, smell, and taste. 424. The cranium or skull is a bony case, of an oval form, occupying the upper and back part of the head, and serving for the protection of the brain, which is lodged in its cavity. Its walls are made up of eight bones, which are firmly united to each other in such a manner that every part is admirably adapted to resist external force. 425. The cranium is constructed on the principle of the arch, the bones at the base of the skull overlapping those above so firmly, that a separation is rarely, if ever, effected during life, and only with great difficulty after death. 426. The spinal column or back-bone, as it is commonly called, consists in man of thirty-three different bones, called vertebrae, which are divided as follows, in man: seven cervical vertebrae, (vertebrae of the neck,) twelve dorsal vertebrae, (vertebrae of the back,) five lumbar vertebrae, (vertebrae of the loins,) five sacral vertebrae, four cocygeal Of how many bones is the skeleton of man formed ? How is it divided ? What bones are included in the head ?—the trunk ?—the extremities ? Of how many bones is the face formed? Describe the cranium. Its walls are made up of how many bones ? On what principle is the cranium constructed ? How is a separation of the bones prevented? Of how many bones does the spinal column consist? How are they divided? ANIMAL MOTION. 205 vertebrae. All these vertebrae are separate at the time of birth, but the five sacral vertebrae are soon after united into one piece—the sacrum-—and the four coccygeal ver- tebrae are also united into one piece, the coccyx. 427. The number of the cervical vertebrae is the same in all the mammalia. Birds have a much larger number: the swan, which is remarkable for the gracefulness and beauty of its neck, has twenty-three vertebrae, the crane has seventeen, and the swallow thirteen. 428. The number of the dorsal vertebrae is the same as the number of the ribs on each side. In man, there are twelve dorsal vertebrae, in the lion thirteen, and in the elephant twenty. 429. The vertebrae are perforated by an aperture, so that when they are all united, they form a continuous tube or canal for the lodgment of the spinal cord. 430. By the division of the spinal column into so large a number of separate bones, very great freedom of motion is allowed, with only a slight bend at any particular point. 431. Each vertebrae consists of a body, which is situated in front of the spinal canal in man, and below it in quad- rupeds ; and of seven processes or projections, which serve to form the spinal canal, and unite the vertebrae to each other by affording attachments for the muscles. 432. In man and the other mammalia, the two surfaces of the body of the vertebrae are nearly flat, and are sepa- rated from each other by a disc of fibro-cartilage. 433. In reptiles and fishes, a different plan is adopted. In serpents, one surface of each vertebra is concave and the other convex, and thus the convex surface of each What change takes place with the five sacral vertebrae ? What is said of the number of the cervical vertebrae in the mammalia ?—in birds ? How many cervical vertebrae has the crane and the swallow? What do the vertebra form when united together? What is gained by the division of the spinal column into so many separate pieces? Of what does each vertebrae consist? What is the form of the two surfaces of the. body of the vertebrae ? What is the form in fishes and reptiles ? What kind of a joint do they form ? 15 206 CLASS-BOOK OF PHYSIOLOGY. vertebra fits into the concave surface of the next, in such a manner that the whole spine becomes a series of ball- and-socket joints—an arrangement which is remarkably adapted to the peculiar movements of those animals. 434. In fishes, both surfaces are concave, and between each vertebra there is interposed a bag, containing fluid, and having two convex surfaces, over which those of the vertebrae can freely play. Very great freedom of motion is thus acquired, though the strength is proportionably diminished. But great strength is not required by ani- mals whose bodies are supported by a medium which is # nearly of the same density with themselves. 435. The extreme flexibility of fishes enables them to propel their bodies by the movements of the tail and hinder parts from side to side, their fins being used prin- cipally for influencing their direction. 436. Man is the only animal whose spine seems adapted to the erect attitude. Its form has some resemblance to the letter S, being a # double curve, and it thus forms a kind of spring that is admirably adapted to diminish the shock produced by a sudden jar, as in the act of jumping upon the feet from a height. The bodies of the vertebrae are also broader, in proportion to their size, than in other animals, while the spinal processes, which form the ridge' along the back, are considerably shorter In quadrupeds which maintain a horizontal position of the spinal column, the spinal processes are very long, for the attachments of muscles to support the head. In figs. 9 and 10, Pl. XIX., may be seen a comparative view of the vertebrae in the lion and ox. 437. The ribs, which are twelve on each side in man, are attached by one extremity to the transverse processes How are the two surfaces separated in fishes ? How do fishes propel their bodies? For what purpose are fins used ? What attitude is peculiar to man ? What is the form of the spine in man ? What advantage is obtained by this form ? How do the vertebrae in man differ from those of all other animals ? What is said of the spinal processes in quadrupeds ? How many ribs arc there in man ? What are the attachments of the ribs? PLATE XIX. __---- £ THE EXTREMITIES. Figure 1.----Anterior Extremity of Man.—a, Scapula, or shoulder blade, b, The humerus, or principal bone of the arm. e, The radius, d, The ulna, e, The carpus, or wrist. /, The metacarpus, or palm, g, The phalanges, or fingers. In figures 2, 3,4,5,6,7, and 8, the corresponding bones are indicated by the same letters as in figure 1. Figure 2.---Anterior Extremity of the Deer. Figure 3.---Anterior Extremity of the Lion. Figure 4.---Anterior Extremity of the Whale. Figure 5.---Anterior Extremity of the Bat. Figure 6.---Anterior Extremity of the Bird, Figure 7.---Anterior Extremity of the Sloth. Figure 8.---Anterior Extremity of the Monkey. Figure 9.----Skeleton of the Lion.—a, The skull, b, The cervical vertebra, c, The dorsal vertebrae, d, The lumbar vertebras, e, The sacrum. /, The caudal vertebra, which compose the tail, g, The ribs, h, The scapula, i, The humerus, j, The fore-arm.—In this part, corresponding to the fore-arm of man, the two bones are united into a sintle one. k, Bones of the wrist, or carpus. /, Bones of the metacarpus, or hand, m, The phalanges, n, The femur, or thigh-bone, united to the bones of the pelvis, o, The patella, or knee-pan. p, The tibia; the two bones of the leg being united into one. q, Bones of the tarsus, or ankle, r. Bones of the foot and toes.—In the lion, the bones of the arm are stout and long, and the fingers short and compact, collectively combining freedom of motion and strength, and admirably adapting this class of animals to the sudden springs with which they pounce upon their prey. Figure 10.----Head and Shoulders of an Ox.—a, The skull, b, The cervical vertebra?. c, The dorsal vetebrae, which are very long, for the attachment of the muscles which sup- port the neck and head, d, The scapula, or shoulder-blade. E.C.lCelloM.Liih 145 ANIMAL MOTION. 211 of the dorsal vertebrae, and by the other to cartilages which' are continuations of the ribs. The cartilages of the first seven, or true ribs, (aa,fig. 1, Pl. XVII,) are united to the Bternum or breast-bone; the cartilages of the five lower ribs (bb,fig. 1, Pl. XVII) are not directly connected with the sternum, and are hence called false ribs. 438. The sternum, or breast-bone, (fig. 1, Pl. XVII) is flat in man, but in those animals which have need of great strength in the upper limbs it is increased in breadth, and furnished with a projecting keel'or ridge for the attach- ment of powerful muscles. In the turtle tribe, it is so far extended as to afford a complete protection to the under side of the animal. 439. The extremities, which have been regarded as appendages to the trunk, are four in number in all the vertebrated animals, and are constructed on the same general plan, though they are widely different in general appearance. The fin of a fish, the wing of a bird, the leg . of a lion or a deer, and the arm of man, are evidently adapted to very different uses, and are apparently destitute of any very striking resemblance to each other; yet, when those same limbs are stripped of the skin and muscles, they will be found to possess the same essential parts, slightly modified, according to their several uses. 440. One of the superior extremities of man, and one of the anterior extremities of a deer, a lion, a whale, a bat, and a bird, are represented by the same letters in figs. 1, 2, 3, 4, 5, and 6, Pl. XIX. 441. The scapula, or shoulder-blade, is a large flat bone, attached to the back part of the trunk by muscles. In How many of the cartilages are attached directly to the breast-bone? What are the five lower ribs called ? What is the form of the breast-bone in man ?—in those animals which have need of great strength in the upper limbs ? What is said of it in the turtle ? How many extremities are there in all the vertebrated animals? What is said of their construction? What illustrations are given ? Describe the scapula or shoulder-blade. How is it attached to the trunk? What advantages are derived from this kind of attachment ? 15* 212 CLASS-BOOK OF PHYSIOLOGY. animals which walk on all-fours, this kind of attachment secures much greater elasticity of movement, and also diminishes the force of the shock in the acts of leaping or running. In man, it gives greater freedom of motion to the arm. 442. The clavicle, or collar-bone, is attached at one ex- tremity to the shoulder-blade, and at the other to the sternum or breast-bone. It acts as a brace to separate the shoulders, and it is accordingly strongest in those animals, the action of whose superior extremities tend to draw them together; while it is comparatively weak, or altogether deficient, in those animals, the action of whose limbs nat- urally tends to keep them asunder. Thus, in birds, there is not only a strong clavicle, but a second bone also, tend- ing to keep the shoulders apart. In the lion, deer, &c, it is entirely wanting. 443. The arm is supported by a single long and cylin- drical bone, c, the humerus. In the fore-arm, there are two long bones, the radius and ulna (d, e), which lie par- allel to each other. The radius is on the outer or thumb side of the fore-arm, and the ulna on the inside. The radius and ulna are connected with one another by liga- ments at their extremities, and by a strong fibrous mem- brane, that passes between their adjacent edges along their entire length. Those bones rotate freely on each other, in such a manner that either the palm or back of the hand may be turned. In most quadrupeds, the radius and ulna are so firmly united together as to be incapable of rotation. 444. The hand is anatomically divided into three por- tions—the carpus, metacarpus, and phalanges. The carpus, What are the attachments of the clavicle ? What is the use of this bone ? In what animals is it strongest? In what animals weakest? How is it in birds ?—in the lion, deer, &c. ? How is the arm supported ? How many bones are there in the fore-arm, and what are they called ? On which side is the radius?—the ulna? How are the radius and ulna connected to each other? How do these bones rotate on each other? In what condition are the radius and ulna in most quadrupeds? How is the hand anatomically divided ? Describe the carpus—the metacarpus—the phalanges. ANIMAL MOTION. 213 which is nearest the wrist-joint, is composed of eight small bones, which are firmly united to each other by ligaments ■—the metacarpus, which consists of five long cylindrical bones, resembling the bones of the fingers so much, that in the skeleton they might be easily mistaken for their joints. The fingers are formed by a series of small bones, called phalanges, of which there are only two in the thumb, and three in each of the fingers. 445. As the foot and ankle, which correspond to the wrist and hand in man, are designed for solidity in quad- rupeds, we find in different parts that there is only one solid piece for two, three, or more bones in corresponding parts in man, though those solid pieces are found originally to have been sev- eral distinct bones that have after- wards united. Thus, in the rumi- nating animals, as the deer and horse (figs. 59 and 60), the number of phalanges is reduced to two in each foot, and in the horse there is but one, which is enveloped by the hoof. 446. The structure of the lower extremities has a very great analogy to that of the upper, the principal differences being such as are necessary to make the lower extremities more solid, and better adapted to serve as organs of locomotion, instead of organs of prehension. _ 447. The lower extremities are connected with the spine by a bony case or basin, called the pelvis. The thigh, How many phalanges are there in the thumb ?-in the fingers ? How are the foot and ankle in quadrupeds made more solid than the wrist and hand in man» Are these parts found solid originally ? How many phalanges are there in the deer ?-in the horse? How do the upper and lower extremities differ from each other? How are the lower extremities connected with the Fig. 59- Fig. 60. spine ? How is the thigh supported ? 214 CLASS-BOOK OF PHYSIOLOGY. like the arm, is supported by a single long cylindrical bone, called the femur, which is bent at the upper extremity at an angle, and its rounded head separated from the main bone by a narrow portion, called its neck. The lower end of the thigh-bone spreads into long condyles, on which the large bone of the leg moves backward and forward. 448. The leg has two bones, but they do not possess the power of rotating on each other like the fore-arm. The main bone, or tibia, is much larger than the fibula, which is a long slender bone, running parallel with the tibia, and apparently serving no other purpose than to give attachment to the muscles. The upper end of the tibia is broad, and has two shallow depressions, in which the con- dyles of the femur or thigh-bone are received. 449. In front of the knee-joint is a small bone, called the patella, which serves the double purpose of protecting the joint and changing the direction of the tendon which comes down from the thigh. 450. The foot is composed, like the hand, of three dis- tinct portions—the tarsus, the metatarsus, and the phalanges or toes. There are seven bones in the tarsus, all of which are larger than those of the carpus. The metatarsus is composed of five long bones in man. The toes, like the fingers, have three phalanges, each, except the great-toe, which has only two. The tarsus and metatarsus form a kind of arch on the inside of the foot, which serves to lodge and protect the vessels and nerves that descend from the legs to the toes. This arch also serves to deaden the shock that would be experienced every time the foot was put to the ground; for by the elasticity of the ligaments which bind these bones together, a kind of spring is What is the form of the upper extremity ?—of the lower extremity ? How many bones form the leg ? What is their comparative size ? Describe the tibia. What bone in front of the knee-joint ? What is its use ? Of how many portions is the foot composed, and what are they called ? How many bones in the tarsus?—the metatarsus? How many phalanges are there in the toes? What do the tarsus and metatarsus form? What advantages are derived from this form of the foot ? PLATE XX. ORGANS OF MOTION.-THE BONES. Figure 1.----Skeleton of the Camel.—-The black ground in this and the two following figures shows the outline of the form when clothed with flesh, a, The skull, b, The cer- vical vertebrae, c, The dorsal vertebra?.—The spinous processes of these vertebra are much longer and larger than those of man, for the purpose of giving attachment to the strong muscles and ligaments by which the heavy neck and head are supported, d, The lumbar vertebrae, e, The sacrum. /, The caudal vertebrae, which compose the tail, g, The ribs, h, The scapula, i, The humerus, j, The fore-arm.—In this part, corresponding to the fore-arm of man, the two bones are united into a single one. k, Bones of the wrist or carpus. I, Bones of the metacarpus or hand, m, The phalanges, n, The femur, or thigh-bone, united to the bones of the pelvis, o, The patella, or knee-pan. p, The tibia; the two bones of the leg being united into one. q, Bones of the tarsus or ankle, r, Bones of the foot and toes.—In animals which do not possess fingers, the bones of the fore-arm, wrist, and hand are always few in number. Thus, in the camel, and in all herbivorous quadrupeds, the fore-arm has only one bone; the wrist, about four; the hand, only one, with sometimes the rudiment of another; while some species have two toes, and others only a single one. t Figure 2.---Skeleton of the Vulture.—vc, Cervical vertebrae, fifteen in number, b, Dorsal and lumbar vertebrae, vs, The sacrum, vq, Vertebrae of the tail, st, The breast- bone, or sternum, cl, The clavicle, h, The humerus, o, The two bones of the fore-arm. ca, Bones of the wrist, imperfectly developed, ph, Bones of the hand and fingers. /, The thigh-bone, t, The two bones of the leg. ta, The shank or ankle bones. Figure 3.---Skeleton of Turtle.—a, Cervical vertebrae, b, Scapula, c, Clavicle, d, Coraooid-bone. e, Dorsal vertebrae. /, Ribs incorporated with the dermal plates which form the shell, g, Marginal plates, h, Pubic bones, t, Femur, k, Tibia. /, Fibula. Figure 4.----Skeleton of the Perch.—a, b, First and second dorsal fins, c, The caudal or tail-fin. d, The anal-fin. e, One of the ventral fins, which correspond to the legs. /, One of the pectoral fins, which are analogous to the an#s. The spinous processes of the vertebrae are long, and are connected with another set of boues, by which they are con- tinued upward, so as to form the frame-work of the fins which arise from the back. The whole number of vertebrae is forty-two, of which twenty-one are dorsal and twenty-one caudal or coccygeal. The number of pairs of ribs is the same as that of the dorsal vertebras. PI. XX ANIMAL MOTION. 219 formed, which yields for a moment to the shock, and then recovers itself. 451. In animals which walk on all-fours, the difference of direction in which the bones of the legs are connected with the spine, prevents a jar from extending to the body. In those animals which obtain their food by sudden and extensive leaps—such as the cat, tiger, and lion—there is an arrangement of the bones admirably adapted to dimin- ish the shock produced by a sudden descent of the body upon the ground. THE MUSCLES. 452. The muscles are the moving power by which the parts of the skeleton are set in motion. 453. Each muscle, if examined carefully, is found to be made of a number of bundles of fibres, and each fibre is formed of numerous smaller fibres or fibrils. The primi- tive fibrils are only about To.^ooth.of an inch in diameter. Each fibre, which extends from one end of the muscle to the other, is supplied with one or more loops of nervous filaments. Through the influence of the nervous system, the muscular tissue is capable of being excited to sudden and forcible contraction. In contracting, the two ends of a muscle approach each other, and swell out in the middle to a corresponding degree. • Thus, the whole muscle, when shortened by the drawing together of its two ends, is greatly enlarged in diameter, especially towards the middle. 354. The energy of muscular contraction depends, in a great degree, upon the power of the stimulus which is transmitted to the muscles from the brain. In animals which walk on all-fours, how is a jar prevented from extending to the body? How are the parts of the skeleton set in motion ? Of what is each muscle made up ? What is the size of the primitive fibrils ? With what is each fibre supplied? How is the muscular tissue excited to contraction ? What change takes place in a muscle when it contracts ? When is the muscle enlarged most in diameter? Upon what does the energy of the muscular contraction depend? 220 CLASS-BOOK OF PHYSIOLOGY. 455. This is frequently observed in instances of great nervous excitement, as when a person is under the influ- ence of violent passion or insanity. A delicate female is thus often a match for three or four strong men, and can even break cords and bands that would hold the most powerful man. 456. In the production of animal motion, the bones not only serve as points of attachment for the muscles, but they constitute a series of levers for the application of muscular force. 457. Most of the muscles act on the bones at great mechanical disadvantage, though it is a law in mechanics that what is lost in power is gained in time. Thus, in fig. 76, the muscle (a) which arises from the top of the Fig. 61. shoulder, and is inserted at e, a short distance from the elbow, acts at a great mechanical loss of power in raising the fore-arm^ but its contraction to a very slight extent will raise the hand through a considerable space. Thus, since the muscle is inserted about one-sixth of the distance from the elbow to the wrist, it will require a force of con- traction in the muscle equal to six pounds to raise one pound at the wrist, while its contraction of one inch will In what cases of great nervous excitement is this frequently observed ? What example is given ? What other purpose do the bones serve besides affording attachment for the muscles? How do most of the muscles act on the bones ? What law of mechanics in regard to this ? Explain Figure 65 What force in the muscle will be required to raise one pound at the wrist, ? P1.21. I C K"elli>'i».Lith Hirttnti PLATE XXI. ORGANS OF MOTION--THE MUSCLES. LATERAL VIEW OF THE PRINCIPAL MUSCLES OF THE HUMAN BODY. Muscles of the Head and Neck.—1, The occipito-frontalis muscle, which elevates the eyebrows, and wrinkles the forehead, expressing astonishment or attention. 2, The orbic- ularis palpebrarum, which closes the eyelids. 3, Triangularis nasi, which compresses the nostrils. 4, The levator labii superioris, which raises the upper lip and expands the nos- trils, expressing disdain. 5, Zygomaticus major. 6, Zygomaticus minor: These muscles raise the upper lip and draw it outward. 7, Orbicularis oris.—Only half of this muscle is seen on the plate. It extends completely around the mouth, which its action is to close. It also assists the lips in whistling, blowing and sucking. 8, The buccinator, or trumpet er's muscle.—It elongates the mouth transversely, and is greatly used in blowing wind- instruments. 9, Depressor labii inferioris, which draws down the under lip, expressing chagrin, disgust. 10, Depressor anguli oris, which depresses the angle of the lips. 11, The masseter.—This powerful muscle is the essential agent in mastication. It raises the lower-jaw, and brings its teeth strongly against those of the upper-jaw. 12, Omo-hyoideus, extending from the edge of the scapula to the hyoid-bone, which it depresses. 13, Sterno- hyoideus, a depressor of the larynx. 14, The digastricus, an elevator of the larynx and a depressor of the lower jaw. 15, The Sterno-cleido-mastoideus, extending from the tem- poral-bone to the sternum and clavicle.—When one of these contracts, it draws the head to its own side; when both contract, the bead is carried forward. 16, The upper portion of the trapezius.—(See Plate XXII.) 17, 18, Scalenus posticus and anticus, which ele- vate the ribs and assist in respiration. Muscles of the Trunk.—19, The pectoralis major, a large muscle, attached to the clavi- cle, sternum and ribs, and to the humerus. It draws forward the shoulder, and also ele- vates the ribs, acting as a muscle of inspiration. 20, The serratus magnus, a powerful muscle, lying beneath the pectoral muscle.—It is divided into slips or digitations, which extend from the ribs to the scapula. It is a muscle of inspiration, the contraction of its slips tending to separate the ribs and thus dilate the chest. 21, 21, &c. External inter- costal muscles, which elevate the ribs. 22, The transversalis abdominis. This muscle draws the front of the abdomen inward, thereby pushing up its contents against the dia- phragm, in the act of expiration. 23, Rectus abdominis, which draws the chest forward toward the pelvis, and assists in straightening the trunk when it has been thrown backward. Muscles of the Superior Extremities.—24, The deltoid muscle. This thick and powerful muscle is attached to the scapula and to the humerus. Ordinarily it elevates the arm; but when the body is raised by the arms, as in climbing, it draws up the trunk. 25, The biceps brachialis.—This Is the large muscle whose projection is felt in front of tho arm. It is a flexor of the fore-arm. 26,26,26, Portions of the triceps brachialis.—(See Pl. XXII). 27, The pronator teres.—This muscle, being attached to the radius, and having its fixed insertion on the humerus, rolls the radius inward, and thus turns the palm of the hand downward. 28, The supinator longus.—This muscle acts as an antagonist to the last, and 222 CLASS-BOOK OF PHYSIOLOGY. turns the hand upward. 29, The flexor carpi radialis. 30, The palmaris longus. 31 Flexor carpi ulnaris — These three muscles are flexors of the wrist upon the fore-arm. Their white and slender tendons are attached to the carpal and metacarpal bones. 32, Tendons of the flexor sublimis digitorum, which bend the second range of the finger-bones upon the first. 33, Extensor carpi radialis longior.—This muscle extends the wrist upon the fore-arm. 34, Extensor communis digitorum.—(See Pl. XXII.) 35, Lower portion of the abductor pollicis longus. 36, Lower portion of the extensor pollicis brevis.—These two muscles draw the thumb outward, and bend it on the carpus. 37, Extensor carpi ulnaris.—(See Pl. XXII.) Muscles of the Inferior Extremities.—-38, The rectus internus.—This muscle is a flexor of the leg. 39, The sartorius or tailor's muscle, the longest muscle in the body.—By its action the legs are bent inward, so as to cross' each other. 40, The rectus femoris. 41, The vastus internus. 42, The vastus externus.—These three muscles extend the leg upon the thigh. 43, Portion of the adductor magnus.—This muscle draws backward the thigh at the hip. 44, A portion of the gluteus raaximus.—(See Pl. XXII.) 45, The gluteus medius. 46, A tendinous expansion called the fascia-lata, forming part of the femoral aponeurosis. 47, The tendon of the vastus femoris, which contains the patella. 48, The triceps surae, or extensor of the foot, composed of three large fasciculi.—(See Pl. XXII.) 49,The peroneus longus; its tendon passes behind and beneath the outer ankle,in a pul- ley-like groove, and is inserted into the first metatarsal-bone. By its contraction, this muscle extends and rotates the foot. 50, The tibialis anticus— This muscle is attached by its tendon to a bone of the instep, and bends the foot upward. 51, The extensor digi- torum communis.—Its four tendons are seen proceeding under the annular ligament of the tarsus, and diverging to the four smaller toes. This muscle extends the joints of the toes, and bends the foot upon the leg. 52, Annular ligament of the tarsus. 53, Adductor pollicis pedis.—(See Pl. XXII.) ANIMAL MOTION". 225 move the wrist through six inches of space. In the same manner the muscle b, fig. 62, which lies on the back of the arm, and is attached to a short projection of the ulna be- hind the elbow, acts with a similar i disadvantage in regard to power, Fig. 62. though it has a corresponding advantage in point of time. 458. Under ordinary circumstances, the energy of mus- cular action depends on the condition of. the muscles themselves. Those muscles which are firm, plump, and high-colored, act with greater force than those which are pale and flabby. In the sedentary and inactive, the mus- cles are smaller in size and less vigorous than in those whose habits are more active. "Within certain limits, the muscles increase, both in size and power, by use. In the smith, who makes constant use of his arms, the muscles are proverbially large and powerful; in the pedestrian, the muscles of the legs become developed. 459. The free exercise of the muscles not only increases the general strength, but greatly improves the vigor and health of all the physical powers. 460. The energy of muscular contraction appears to be greater in insects, in proportion to their size, than in any other animals. Thus, a flea will leap sixty times its own length, and move as many times its own weight. The same muscular power, in a man of six feet, would enable him to leap a distance of over three hundred feet, and to If this muscle contracts one inch, through what space will it move the wrist? Under ordinary circumstances, upon what does the energy of muscular con- traction depend? What kind of muscles act with greatest energy? What is said of the muscles in the sedentary and inactive? How are the muscles affected by use ? What is said of the arms of the smith, and the legs of the pedestrian ? What is said of the influence of exercise on the muscle?, and on the health of all the powers ? In what animals is the energy of muscular contraction greatest in proportion to size ? How far will a flea leap ? How far would the same muscular power in man enable him to leap ? How much would it enable him to lift ? 226 CLASS-BOOK OF PHYSIOLOGY. lift a weight of over ten thousand pounds. A species of beetle can support a weight equal to at least five hundred times that of its own body; and another, by the power of its jaws, has been known to gnaw a hole of an inch in diameter in the side of an iron canister in which it had been confined. The rapidity of the movements in insects is also so great, that the vibrations of the wings in some species have been calculated at several hundred, and, in some of the smaller insects, at several thousand in a second of time. 461. The various kinds of progression in different ani- mals are called swimming, crawling, flying, walking, running, jumping, &c. 462. In fishes, the body is propelled through the water by the tail, on the same principle as a boat in impelled by sculling. The tail being vertical, in most cases its stroke is horizontal, and the body is propelled forward by the resist- ance of the water to its broad expansion", as it strikes from Fig. 63.—Tail-Fin of Whale. side to side, by the alternate contraction of the muscles which extend from the long process of the vertebrse to the tail. In the whale, the tail (fig. 63) is expanded hori- What is said of some species of beetle ? What is said of the movements of insects? What are the various kinds of progression in animals called? How is the body in fishes propelled through the water? How is the stroke made against the water in most cases ? What propels the fish ? How is the tail expanded in the whale, and how is its stroke made ? What is said of the structure of the tail ?—of the power of the whale ? ANIMAL MOTION. 227 zontally, so that its stroke is vertical. The texture of the tail is such, that it is insensible to pain, and it is so tough that it is rarely torn or injured. The whale is therefore capable of inflicting a tremendous blow without any pain to itself, and of cutting a strong boat asunder, or of driv- ing it Avith the swiftness of an arrow to the depth of many fathoms. 463. The fins of fishes are used principally to give direction to the body as it is propelled through the water. 464. In serpents, progression is effected by undulations of the spine, or by bringing the two extremities of the body near together, and then, the tail being fixed, the head is projected forward the length of the body: again the tail is brought forward, and the same process repeated. Sometimes, instead of this alternate contracting, the whole body is brought into a spiral coil, and by the contraction of all the muscles on one side, and then by a sudden con- traction of the muscles of the opposite side, the whole body is propelled, as by the unwinding of a powerful spring, with an impulse which raises it to some height from the ground. 465. Flying has some resemblance to swimming, both being executed in a fluid medium, which, to a certain ex- tent, buoys up the body and offers resistance to its progress. Water, however, affords more support to the body and a greater resistance to the propelling organs. Birds are especially adapted to flying by their formation. Their bones excel those of all other animals in combining strength and lightness. All the long bones are hollow cylinders filled with air. The breast-bone, which resembles in form the keel of a ship, presents a very large surface for the attachment of the muscles, and it is also connected very For what are the fins of fishes principally used 1 How is progression effected in serpents ? What other mode of progression is there in some ser- pents? What does flying resemble ? How are birds adapted to flying? In what respect do the bones of birds excel all other animals ? What are the long bones ? What does the breast-bone resemble ? 16* 228 CLASS-BOOK OF PHYS10LOOV. firmly by the ribs to the vertebrae of the back. The whole bony apparatus of the body is thus very strongly knit together, and yet the entire skeleton is exceedingly light— the body being suspended in the air by the resistance it affords to the wings. The muscles which are attached to the breast-bone are much larger than in other animals; while the muscles on the back, that raise the wings, are correspondingly smaller. The greatest power of the wings is consequently in 4;he downward direction. In the effort to fly, the wings are expanded, brought down with con- siderable force, and then brought up to the body, being raised again in such a form as to offer the least possible resistance to the air. 466. In many birds the rapidity of flight is very great, much greater than by any other mode of progression. The Eider-duck is said to fly ninety miles an hour, and some species of hawk one hundred and fifty. 467. Walking is produced by the alternate contraction and extension of the limbs. In man, one foot is placed in front upon the ground while the other is extended or car- ried backward, and its length increased, carrying forward the whole body. In quadrupeds, two feet are placed on the ground at once. 468. In running, the body momentarily quits its sup- port on the ground at intervals, the foot in advance not being planted on the ground at the time, the hinder one quits it. In this action, the ostrich probably surpasses all other animals, as it is said to outstrip the swiftest grey- hound at its greatest speed. 469. In trotting, the fore-foot of one side and the hind- What is gained by this peculiar form ? How is the bony apparatus united together ? What is said of the skeleton ? How is the body suspended in the air ? What is said of the muscles which are attached to the breast-bone ?__to the back ? In what direction is the greatest power of the wings ? Describe the movement of the wings in the effort to fly ? What is said of the rapidity of flight in birds? What examples? How is walking produced? How is it produced in man?—in quadrupeds? Describe running. What animal surpasses all others in running? Describe trotting. PLATE XXII. ORGANS OF MOTION.-THE MUSCLES. POSTERIOR VIEW OF THE MUSCLES. The right side of the figure displays the outer or superficial layer of muscles. On the left side, they are represented clothed with their fasciae or aponeuroses, which form a tendinous membrane. Muscles of the Head and Neck.—l, The occipitalis.—This muscle draws the scalp back- ward and downward, giving the expressions of joy or surprise. 2, Muscles of the ear, 3, The masseter. 4, The sterno-cleido-mastoideus. 5, The splenius.—When the two splenii act together, they draw the head backward, elevating the chin; when one acts, it bends the head and neck to one side. Muscles of the Trunk.—6, The trapezius.—This muscle is attached to the spinous pro- cesses of the dorsal vertebra? and to the scapula. It carries the shoulder backward. 7i The latissimus dorsi.—a, a, Its tendinous or aponeurotic portion.—This broad muscle draws down the arm, or, if the arm be fixed, it draws the trunk towards the shoulder. 8, Part of the obliquus externus.—This muscle compresses and raises up the contents of the abdomen. Muscles of the Superior Extremities.—9, Posterior portion of the deltoid muscle. 10, The infra-spinatus.—This muscle turns the shoulder outward, and assists in keeping the head of the humerus in its socket. 11, 11, 11, The three portions of the triceps extensor cubiti.—This muscle is fixed by its tendon to the olecranon process of the ulna, and by its contraction it extends the fore-arm. 12, The extensor communis digitorum.—Its four tendons pass under the annular ligament, and go to the four fingers ; so that it becomes an extensor of the whole hand, as well as of each of the phalanges. 13, The extensor proprius of the little-finger.—By means of this slender muscle, the little-finger can be sep- arately extended. 14, The extensor carpi ulnaris.—This muscle extends the hand on the wrist. 15, Flexor carpi ulnaris. 16, Tendon of the long extensor of the thumb. /, An- nular ligament of the carpus. Muscles of the Inferior Extremities.—17, The gluteus maximus.—This large, thick muscle is of greater volume than any other in the body, and is the principal agent in pre- serving its equilibrium. It extends from the iliac bones to the femur, and acts as an ex- tensor of the thigh. 18, The biceps femoris.—This strong muscle, extending from the pelvis to the tibia, bends the leg, and also extends the pelvis upon the leg, keeping it erect. 19, The semi-tendinosus.—This muscle bends the leg upon the thigh, turns the point of the foot downward and inward, and keeps the pelvis erect when standing. 20, The vastus externus. 21,21, The triceps surae, consisting of three portions—the gastrocne- mii, a, b, and the soleus, c; it terminates in the strong tendon of Achilles, d. It forms the extensor muscle of the foot. $h m '■. »« . -, E..C.Kelto««.Lrfli. ANIMAL MOTIOxX. 233 foot of the other one are raised and carried forward to- gether; and when these are set down, the other fore and hind foot are raised and advanced. 470. In leaping, the body is projected into the air by the sudden extension of the joints, especially those of the hinder parts of the body which have been previ- ously bent, and having traversed a greater or less distance, the body comes again to the ground. The hare, rab- bit, squirrel, kangaroo, &c, are especially adap- ted to this kind of pro- gression by the length of the posterior extrem- ities, which are nearly double that of the anterior, as may be seen in fig. 64. Fig. 64.—Kanoaroos. Describe leaping. 284 CLASS-BOOK OF PHYSIOLOGY. CHAPTER XIV. THE VOICE. 471. The voice in animals and in man consists essen- tially in the production of sounds expressive of ideas, feelings, passions, and desires. 472. Insects and all water-breathing animals may be said to be mute, since they have no voice. Insects, how- ever, possess the power of producing sounds by certain movements which to some extent are characteristics of the different species. The shrill chirp of the cricket is pro- duced by rubbing the wing-cases against each other. The harsh shriek of the grasshopper is caused by friction of the legs against the wings. The shrill trumpet sound of the mosquito and the busy hum of bees and flies are pro- duced by the rapid motion of the wings while flying. Other sounds are caused by the jaws in the act of masti- cating, as the remarkable "death-watch" so called.* 473. In the air-breathing vertebrata, the production of sounds depends upon the passage of air through a certain In what does the voice consist? What animals may be said to be mute? Why may they be said to be mute ? How do insects produce sounds ? How is the chirp of the cricket produced ? How is the sound of a grasshopper caused?—the mosquito?—bees and flies? How are other sounds caused? How is sound produced in air-breathing vertebrata ? * "-----—---------------------A wood-worm, That lies in old wood, like a hare in her form! With teeth or with claws, it will bite or will scratch, And chamber-maids christen this worm a death-watch; Because, like a watch, it always cries click! Then woe be to those in the house who are sick! For sure as a gun they will give up the ghost, If the maggot cries click when it scratches the post: But a kettle of scalding hot water ejected, Infallibly cures the timber affected: The omen is broken, the danger is over, The maggot will die, and the sick will recover." THE VOICE. 235 portion of the respiratory tube. In reptiles, it is at the point where the trachea or windpipe opens into the pharynx that the vibratory apparatus is situated. The sounds pro- duced by this class is very simple, being little else than a hiss. 474. In birds, the situation of the vocal organ is very different. The trachea opens into the pharynx, as in rep- tiles, by a mere slit, but the most essential organ is at the lower extremity of the trachea, near its division into the bronchi. This apparatus, which seems to be a kind of drum, is variously formed in different species. In fig. 65 Fig. 65. -Larynx of a spe- Fig. 66.—Larynx of Rook. Fig. 67.—Vertical cies of Duck. section of the same. a, trachea; b, a kind of beny drum; c, bronchial tpbes. is a representation of the larynx of a species of duck; fig. 66 represents the larynx of a rook; and fig. 67 a sec- tion of the same. At a, in each of the above figures, is seen the larynx; at b, a sort of bony drum; and at c, the bronchial tubes. In the most esteemed singing-birds these parts are much more complicated than in birds not distin- guished for their vocal powers. 475. In man and the mammalia, the vocal organ is situ- ated at the upper part of the windpipe, and is called the Where is it produced in reptiles ? How are the vocal organs situated in birds ? What does the apparatus resemble ? In what birds is it most com- plicated ? Where is the vocal organ situated in man, and what is it called^ From what does it receive its peculiar form? 236 CLASS-BOOK OF PHYSIOLOGY. larynx. The larynx receives its peculiar form from four cartilaginous pieces, called the thyroid, the cricoid and the arytenoid cartilages. Fig. 68.—Human Larynx, viewed sideways.—A, hy- oid-bone; /, point of origin of muscles of the tongue; t, thyroid cartilage; a, pro- jection in front, commonly called Adam's Apple; c, cricoid cartilage; tr, tra- chea; o, posterior side of the larynx, in contact with the oesophagus. Fig. 69.—Vertical Sec- tion of the Larynx.— ar, arytenoid cartilages; v, veDtricle of the glottis; e, epiglottis. The other references as before. Fig. 70.—Front View of the Larynx.— The interior wall is marked by the lines a, a, b, b,; li, inferior ligaments of nce in colour to a ripe grape. U'vula tx>. dim. of uva, a grape). The pendulous body which hangs down from t? < middle of the soft palate. V Vaccina tion (L. vacca, a cow). The act of inserting vaccine matter; inocuKtion for the cow-pox. Vac'cut-i (L. vaccus, empty). Literally, an empty place. This term gen- erally denotes the interior of a close vessel, from which the atmospheric air ard every other gas has been extracted. Valve - u. valva, folding-doors). A close lid affixed to a tube or opening in some lessel, by means of a hinge, or other movable joint, and which can be or-ned only in one direction. Hence it signifies a little membrane whic'i prevents the return of fluid in the blood-vessels and absorbents. VAL'vr.LA (L. dim. of valve). A little valve. Vas, Vasis. Plural Vasa. A vessel, or any utensil to hold liquor. Vas'cu* iR System. That part of the animal economy which relates to the vesseH Ve'nou . Belonging to a vein. [75.] Ventb c'ulus (L. dim. of venter, the belly). The term ventricle is also applied to two cavities of the heart, and to several cavities in other parts of the body. Ver'miform (L. vermis, a worm, forma, likeness). Worm-like. 19* 270 CLASS-BOOK OF PHYSIOLOGY. Vermic'ula. Having a worm-like motion. Ver'tebra (L. verto, to turn). A bone of the spine, so named from its turn- ing upon the adjoining one. [286.] Ver'tebral. Connected with the vertebra. Vertebra'ta. Animals which have an internal skeleton, supported by a ver- tebral column. [281.] Ves'sicle (L. dim. of vesica, a bladder). A little bladder. Ves'tibule (L. vestibulum, a threshhold). A small oval cavity of the inter- nal ear, so named from its forming an entry to the cochlea and semi-circu- lar canals. [46.] Vill'us. Literally, the shaggy hair of beasts. Some of the membranes of the body, as the mucous membrane of the intestinal canal, present a sur- face of minute papillae, termed villi, villosities, resembling a downy tissue, continually covered with fluid. Vit'reous Body (L. vitrum, glass). Vitreous humour. A transparent mass, resembling melted glass, occupying the globe of the eye, and inclosed in the hyaloid membrane. [376.] Voice. [471.] W Warm-blooded. A term applied to the mammalia and birds which have a two-fold circulation. [128.] X Xy'ph'oid (Gr. xiphos, a sword, eidos, likeness). Sword-like; a term applied to the cartilage of the sternum. Z Zool'ogy (Gr. zobn, an animal, logos, a description). That branch of Nat- ural History which treats of animals. Zygomat'icus (Gr. zugbs, a yoke). A name given to two muscles of the face, which are attached to the zygoma or arch formed by the processes of the bones. D. Appleton & Coi's Educational Publications. Prof. Mandeville's Beading Books. *** The annexed series of Reading Books are very extensively introduced and cob* mended by hundreds of the most experienced teachers in the country. " Prof. Mande- ville's system is eminently original, scientific and practical, and destined wherever It it tetrc-duced to supersede at once all others. I. P3IM ART, or FIRST HEADER. Price 10 cents. II. SECOND READER. Price 16 cents. These two Readers are formed substantially on the same plan; and the second is a continuation of the first. The design of both is, to combine a knowledge of the meaning and pronunciation of words, with a knowledge of their grammatical functions. The parts of speech are introduced successively, beginning with the articles, these are fol- lowed by the demonstrative pronouns; arid these again hy others, class after class, until all that are requisite to form a sentence have been separately conside' id; when the common reading lessons begin. The Second Reader reviews the ground passed over in the Primary, but add* largely to the amount of information. The child is here also taught to read writing as w ell aa printed matter; and in the reading lessons, attention is constantly directed to the differ- ent ways in which sentences are formed and connected, and of the peculiar manner in which each of them is delivered. All who have examined these books, have pro- nounced them a decided and important advance on every other of the same class in use. III. THIRD READER. Price 25 cents. IV. FOURTH READER. Price 33 cents. In the f\v< two Readers, the main object is to make the pupil acquainted with the moaning and t'unrrwns of words, and to impart facility in pronouncing them in senten- tial cnniHTtwn : the leading desien of these is to form a natural, flexible, and varied de- livery. Aceiu-diiiLriy, the Third Header opens with a series of exercises on articulation ' and modulation, c>n!aining numerous examples for practice on the elementary sounds (including en-ors to lie corrected) and on the different movements of the voice, produced by sentential structure, by emphasis, and hy the passions. The habits formed by these exercises, which should be tlioronghly, as they can be easily mastered, under intelligent instruction, find scope for improvement and confirmation in the reading lessons which follow, in the same book and that which succeeds. These lessons have been selected with special reference to the following peculiari- ties: 1st Colloquial character; 2d. Variety of sentential structure; 3d. Variety of sub- ject matter; 4th. Adaptation to the Progressive development of the pupil's mind; and, as far as possible, 5th. Tendency to excite moral and religious emotions. Great pains have been taken to make the books in these respects, which are, in fact, characteristic of the whole series, superior to any others in use; with what success, a brief comparison will readily show. V. THE FIFTH READER; or, COURSE OF READING. Price 75 cents. VL THE ELEMENTS OF READING AND ORATORY. Price $1. These books are designed to cultivate the literary taste, as well as the understanding and vocal powers of the pupil. The Cotjese op Reading comprises three parts ; the first part containing a more elaborate description of elementary sounds and the parts of speech grammatically con- sidered, than was deemed necessary in the preceding works; here indispensable: pari second, a complete classification and description of every sentence to be found in the Eno-lish, or any other language; examples of which in every degree of expansion, from a few words to the half of an octavo page in length, arc adduced, and arranged to be read; and as each species has its peculiar deliverv as well as structure, both are learned at the san.e time; partthi.-d, paragraphs; or sentences in their connection unfolding ztncral thoughts, as in the common reading books. Ii may be observed that the selec- tions of sentences in part second, and of paragraphs in part third, comprise some of the finest gems in the language: distinguished alike for beauty of thought and facility of diction! If not found in a school book, they might be appropriately called "elegant extracts." Tire Elements op Reading and Okatory closes the series with an exhibition of Lie whole theory and art of Elocution exclusive of gesture. It contains, besides the alassification of sentences already referred to, but here presented with fuller statement and illustration, the laws of punctuation and delivery deduced from it.-.the whole fol« towed by carefully selected pieces for sentential analysis and vocal practice. Q D. Appleton t compilation for that purpose we have ever seen. The selections are very judi- ciously made, and are calculated to interest the scholar, as well as to furnish him with models after which to form his style."—Evening Post. "This compilation for the use of students in elocution has been prepared with great judgment by the author, who has had much experience in this branch of education, having been for many years a professor of elocutioi^it West Point A principal feature in this work is that the selections have buen mostly made from the speeches of great orators of the present century, and many of them are specimens of tts eloquence ot dis- tinguished statesmen of our nnion."—Phil. Advertiser. " This work is a complete cyclopedia of specimens of English and American oratory and poetry, for the use of colleges and schools, selected and arranged with skill and good taste. The plan of the work, which is quite original, and will commend itself to every mind, is to present the best specimens which can bo obtained of the oratory and poetry of all parts of the Union, and of living authors. This has not been attempted so thor oughly in any similar work, and as this is decidedly the Book of Oratory of the Union wo doubt not that it will be eminently successful."—Home Journal. PROP. EDWARD C. MARSHALL. The First Book of Oratory. One volume 12mo. Price 62 cents. D. Applcton & Co?% Educational Publications. PROF. LATHAM. A Hand-Book of the English Language ; For the Use of Students of the Universities and Higher Classes ol Schools. 12mo. $1 25. TlHs is an abridgment, or a compendium, by the author, of his large and celebrated! work on the English language; and must prove an invaluable aid to every intellicent teacher of English Grammar. It is quite unlike any work that has heretofore been pub- lished among us, except it may be some that have borrowed largely from this treasure- bouse of information on the English language. It is divided into seven Parts. The first and second parts, occupying nearly SO pages, are devoted to the history and analysis of the English language, designed to show the various elements of which this language Is composed, and the manner in which these elements were introduced gradually into the language. These parts contain the results of very extended and profound study, and furnish in- formation of great value to the student. After this follows the grammatical parts of the work, strictly so called, in which the learned grammarian treats in Part III. of sounds, letters, pronunciation, and spelling, concluding with an historical sketch of the English Alphabet: in Part IV., of Etymology, under its various usual heads; in Part V., of Syntax; in Part VI., of Prosody; and in Part VII., of dialects of the English language. Biich is the general plan of a work which cannot fail to receive a cordial welcome among the scholars of America, coming to them, as it does, with the seal of English approval on it, as the work of the age on English Grammar, and presenting, as it does, on its very face, the evidence of profound and patient scholarship and philosophical discrimiuation and analysis. " We should be glad to see this excellent work introduced into all our colleges and higher schools of learning. The ethnology of the English language is too rarely under- stood. Most men of education seem to think their native tongue a vulgar study, and confine their researches to the dead languages, or to modern dialects, which are far in- ferior in variety and power to the noble idiom in which Milton sung and Addison wrote. Dr. Latham's profound and acute investigations have enabled him to throw a flood of light on the derivations of words, and the mutations of orthography. His larger trea- tises have given him a name among the most learned ethnologists, and this summary of the results of his labors will make him more widely known among general readers."— Commercial Advertiser. "A work of great research, much learning, and to every thinking scholar, it will ba a book of study. The Germanic origin of the English language, the* affinities of the English with other languages, a sketch of the Alphabet, a minute investigation of tho Etymology of the language, &c, of great value to every philologist—O&serper. G. F. GRAHAM English Synonymes, Classified and Explained; With Practical Exercises. Designed fcr Schools and Private Tuition. With an Introduction and Illustrative Authorities, by Henry Eked, LL.D. 12mo. §1. "This is one of the best books recently published in the department of language, and rMl do much to arrest the evil of making too common use of inappropriate words. The work is well arranged for classes, and can be made a branch of common school study. The excellent and elaborate work of Crabb is adapted to the private study, and has beea used by many scholars and professional men with great profit, but never could find us olace in the school-room; consequently, this important department of study has been left Kch meanrrcommon conversation and miscellaneous reading might afford. »TlnTwork is Xirably arranged. The Synonymes are treated with reference to their character, as generic and specific; as active and passive; as positive and negative; IBd«'l clasfinXPs boSoykHdbe organized in every school"- Teachers' AOvocaU U. Applcton <& Uo.\$ Educational Publications. PROP. G. P. QUACKENBOS. First Lessons in Composition, In which the Principles of the Art are Developed, in connection with the Principles of Grammar; embracing full Directions on the Subject of Punctuation: with Copious Exercises. 12mo. 45 cents. %* This work has received tho universal approbation of Practical Teachers and tba Ififcss of the United States. From the Principal, and assistant Teachers, of the Mechanics'1 Institute, 2T. Y. "With feelings of unfeigned pleasure, I have examined the 'First Lessons In Conn position,' by G. P. Quackenbos. " This work possesses the rare merit of combining what is new with what is already familiar in publications of the science upon which it treats. The matter which is com- prised in the first fifty pages is invaluable. If the pupil be faithfully trained in these preliminary exercises, he will be amply prepared to appreciate the principles detailed i» the latttr part of the volume. It is, without question, the best treatise that has ap. E eared on the subject which it professes to illustrate, as every part can be made availa- le to pupils by the judicious teacher; and it appears to be admirably adapted to mew* the present requirements of schools. J. T. BERGEN, Professor of Belles-lettres. M. C. TRACY, Principal of Mec. Inst. School J. OVERACRE. " This book presents an exceedingly simple method of learning the principles ot Grammar, and it is so completely adapted even to the understanding of the youngest pupil, that it cannot fail to be a great assistant to the teachers of schools. It is pre- pared with much skill and judgment, and from the suggestions of long experience. It will be found to possess & more than ordinary intrinsic value."—N. Y. Courier <& Enquirer. PROF. JOHN W. S. HOWS. The Shakspearian Reader; A Collection of the most approved Plays of Shakspeare, carefully ra vised ; with Introductory and Explanatory Notes, and a Memoir of the Author. Prepared expressly for the use of Classes and the Family Reading Circle. 12mo. $1 25. "At a period when the fame of Shakspeare Is "striding the wortd like a colossus," nd editions of his works are multiplied with a profusion that testifies the desire awak« ened in all classes of society to read and study his imperishable compositions,—there needs, perhaps, but little apology for the following selection of his works, prepared ex- pressly to render them unexceptionable for the use of Schools, and acceptable for Family reading. Apart from tho fact, that Shakspeare is the " well spring" from which may be traced the origin of the purest poetry in our language,—a long course of professional ex- perience has satisfied me that a necessity exists lor the addition of a work like the present, to our stoek of Educational Literature. His writings are peculiarly adapted for the purpose of Elocutionary exercise, when the system of instruction pursued by the Teacher is based upon the true principle of the art, viz.—a careful analysis of the struo- ture and meaning of language, rather than a servile adherence to the arbitrary and me ehanical rules of Elocution. "To impress upon the mind of the pupil that words are the exposition of thonei f, and that in reading, or speaking, every shade of thought and feeling has iis appropriate shade of modulated tone, ought to be the especial aim of every Teacher: and an atitho? like Shakspeare, whose every line embodies a volume of meaning, should surely form on« of our Elocutionary Text Books. * * * Still, in preparing a selection'of hia Works for the express purpose contemplated in my design, I have not hesitated to exer- cise a severe revision of his language, beyond that adopted in any similar undertaking «" Bowdler's Family Shakspeare" not even excepted;—and simply, because I practi- cally know the impossibility of introducing Shakspeare as a Class Book, or as a satisfac- tory Reading Book for Families without this precautionary revision."— Extract from the Preface. D. Appleton & Co. s Educational Publications. THOMAS ARNOLD, D. D. The History of Borne. Beprinted entire from the last London Edition. Three volumes in on* 6vo. $3. "Arnold's History of Rome Is a well known standard work. Full and accurate M Hiebuhr, but much more readable and attractive; more copious and exact than Knight- ley or Schmitz, and more reliable than Michelet, it has assumed a rank second to none In value and importance. Its style is admirable, and it is every where imbued with the truth-loving spirit for which Dr. Arnold was pre-eminent. For Colleges and School! this History is invaluable; and for private, as well as public libraries, it is indispensable." THOMAS ARNOLD, D. D. Lectures on Modern History. Edited, with a Preface and Notes, by Henry Eeed, LL. D. 12mo, $1 25. Extract from the American Editor's Preface :—"In preparing this edition, I have bad in view its use, not only for the general reader, but also as a text-book in education, especially in our college course of study. * * * * The introduction of this work gs a text-book I regard as important, because, as far as my information entitles me to ■peak, there is no book better calculated to inspire an interest in historical study. That It has this power over the minds of students I can say from experience, which enables me al»o to add, that I have found it excellently suited to a course of college instruction. By intelligent and enterprising members of a class especially, it is studied as a text- book with zeal and animation. " These Lectures, eight in number, furnish the best possible introduction to s philo- sophical stndy of modern history. Prof. Reed has added greatly to the worth and inter- est of the volume, by appending to each lecture such extracts from Dr. Arnold's othei writings as would more fully illustrate its prominent points. The Notes and AppendU which he has thus furnished are exceedingly valuable."—Evening Post. PROP. FREDERICK KOHLRAUSCH. A History of Germany ; From the Earliest Period to the Present lime. Translated from the last German Edition, by James D. Haas. With a Complete Index, prepared expressly for this Edition. 8vo. $1 75. "Messes. Appixton: " Gentlemen,—Having adopted Kolrausch's History of Germany, as a text-book rot •n advanced class in history, I take great pleasure in stating that I have found no work, In a wide range of historical instruction, both ancient and modern, devonred with more avidity by my pupils, or resulting in their greater profit Next to the history of our »wn country and that of England, I know of none so important to be familiarly under- stood by our American youth, as the History of Germany; in its bearings on modern eiviliwrtion, the Protestant Reformation, the progress of literature, the advancement, o! the Arts and Sciences, and high classical scholarship, as well as also our own very origin and language. , ... ., "The history of a nation with whose past and present we especially, not to say the whole civilized world, have such vital connections, though unknown perchance to a ereat extent to our educated men of a preceding generation, ought now to be introduced every where at once into all our high schools, as an essential part of a course of liberal MtaMtinn " YOUTS, &C, " Bsookmtn, Jan. 24i/t, 1852." 7 D. Appleton & Goh Educational Publications. PROP. GEORGE W. GREENE. History of the Middle Ages. For Colleges and Schools (chiefly from the French). 12mo. $1 25. "No portion of history has been less studied, either by old or young, than that of the middle ages. This is owing in a great degree, we believe, to the defective text-books which have hitherto been in use, for the period in question is itself one of the most in- teresting and important in the annals of mankind. It was the birth-time of modern society—the source and fountain of modern civilization—the period in which a larg« portion of the civil and religious institutions which we now meet highly prize had thei/ origin. ~" The work before us, compiled principally from the French, by Professor Qreene, of Brown University, is the fruit of much learning and research. It furnishes a brief, though clear and well digested, exposition of the leading revolutions of the middle ages, and is designed to introduce the student to an acquaintance with those various and complicated agencies which, out of barbarism and decay, slowly built up the nations of Modern Europe. The plan is judicious, and the execution is in the admirable literary- taste which always characterizes the writings of Mr. Greene. The period embraced in the work reaches from the first general irruption of .the barbarians at the beginning of the fifth century, to the fall of Constantinople, near the middle of the fifteenth—a period crowded with momentous changes in both the civil and ecclesiastical affairs of the world—marked by the rise and fall of numerous dynasties, and by the utter extinction of the ancient civilization and the formation of another entirely new. " We hope to see this work generally adopted as a text-book in schools and colleges where History is made a part of the course of instruction, for wo feel assured that both instructors and pupils will find it admirably suited to explain the interesting and impor> tant period to which it relates."—Providence Journal. PROP. GEORGE W. GREENE. Atlas of Mediceval Geography ; Designed to accompany the above. One volume Svo. (In press.) Contents :—Map 1. The Roman Empire and Northern Barbarians in the Fourth Century. Map 2. Europe in the Sixth Century. Map 3. Europe in the times of Char- lemagne. Map 4. Europe in the second half of the Tenth Century. Map 5. Europe in the time of the Crusades. Map 6. Europe at the end of the Fourteenth Century. E. M. SEWELL. The First History of Rome ; With Questions. 16mo. 50 cents. " Norfolk Academy, Norfolk, Va. "I must thank you for a copy of ' Miss Sewell's Roman History.' Classical teachers bave long needed just such a work: for it is admitted by all how essential to a propel comprehension of the classics is a knowledge of collateral history. Yet most pupils are construing authors before reaching an age to put into their hands the elaborate works we have heretofore had upon Ancient History. Miss Sewell, while she gives the most important facts, has clothed them in a style at once pleasing and comprehensible to th« most youthful mind. " R. B. Tschttdi, Prof, of Ancient Languages" E. M. SEWELL. The First History of Greece; With Questions, on the Plan of the First History of Rome. 16mo {In press.) H V