A COMPENDIOUS SYSTEM OF ANATOMY. IN SIX PARTS. PART I. OSTEOLOGY. II. OF THE MUSCLES, ETC. III. OF THE ABDOMEN. PART. IV. OF THE THORAX. V. OF THE BRAIN AND NERVES. VI. OF THE SENSES. ILLUSTRATED WITH TWELVE LARGE COPPERPLATES. EXTRACTED FROM THE AMERICAN EDITION OF THE ENCYCLOPÆDIA. NOW PUBLISHING, BY THOMAS DOBSON, AT THE STONE HOUSE, N° 41, SECOND STREET, PHILADELPHIA. MDCCXII.  A SYSTEM OF ANATOMY. ANATOMY is the art of dissecting, or arti- ficially separating and taking to pieces, the different parts of the human body, in order to an exact discovery of their situation, structure and œconomy. The word is Greek, ανατομη; derived from αγατεμγω, to dissect, or separate by cutting. INTRODUCTION. § I. History of Anatomy. This art seems to have been very ancient; though, for a long time, known only in an im- perfect manner.—The first men who lived must A2 have 4 INTRODUCTION. have soon acquired some notions of the structure of their own bodies, particularly of the external parts, and of some even of the internal, such as bones, joints, and sinews, which are exposed to the examination of the senses in living bodies. This rude knowledge must have been gradu- ally improved, by the accidents to which the body is exposed, by the necessities of life, and by the various customs, ceremonies, and super- stitions, of different nations. Thus, the observ- ance of bodies killed by violence, attention to wounded men, and to many diseases, the vari- ous ways of putting criminals to death, the fune- ral ceremonies, and a variety of such things, must have shown men every day more and more of themselves; especially as curiosity and self-love would here urge them powerfully to observation and reflection. The brute-creation having such an affinity to man in outward form, motions, senses, and ways of life; the generation of the species, and the effect of death upon the body, being observed to be so nearly the same in both; the conclusion was not only obvious, but unavoidable, that their bodies were formed nearly upon the same model. And the opportunities of examining the bodies of brutes were so easily procured, indeed so ne- cessarily occurred in the common business of life, that the huntsman in making use of his prey, the priest in sacrificing, the augur in di- vination, and, above all, the butcher, or those who 5 INTRODUCTION. who might out of curiosity attend upon his ope- rations, must have been daily adding to the little stock of anatomical knowledge. Accordingly we find, in fact, that the South-sea-islanders, who have been left to their own observation and rea- soning, without the assistance of letters, have yet a considerable share of rude or wild anatomi- cal and physiological knowledge. Dr Hunter informs us, that when Omai was in his museum with Mr Banks, though he could not explain himself intelligibly, they plainly saw that he knew the principal parts of the body, and some- thing likewise of their uses; and manifested a great curiosity or desire of having the functions of the internal parts of the body explained to him; particularly the relative functions of the two sexes, which with him seemed to be the most interesting object of the human mind. We may further imagine, that the Philoso- phers of the most early ages, that is, the men of curiosity, observation, experience and reflec- tion, could not overlook an instance of natural organization, which was so interesting, and at the same time so wonderful, more especially such of them as applied to the study and cure of diseases. We know that physic was a branch of philoso- phy till the age of Hippocrates. Thus the art must have been circumstanced in its beginning. We shall next see from the tes- timony of historians and other writers, how it actually appeared as an art, from the time that writing 6 INTRODUCTION. writing was introduced among men; how it was improved and conveyed down to us through a long series of ages. Civilization, and improvements of every kind, would naturally begin in fertile countries and healthful climates, where there would be leisure for reflection, and an appetite for amusement. Accordingly, writing, and many other useful and ornamental inventions and arts, appear to have been cultivated in the eastern parts of Asia long before the earliest times that are treated of by the Greek or other European writers; and that the arts and learning of those eastern people were in subsequent times gradually communi- cated to adjacent countries, especially by the me- dium of traffic. The customs, superstitions, and climate of eastern countries, however, appear to have been as unfavorable to practical anatomy, as they were inviting to the study of astronomy, geometry, poetry, and all the softer arts of peace. Animal bodies there, run so quickly into nau- seous putrefaction, that the early inhabitants must have avoided such offensive employments, as ana- tomical inquiries, like their posterity at this day. And, in fact, it does not appear, by the writ- ings of the Grecians, or Jews, or Phœnicians, or of other eastern countries, that anatomy was particularly cultivated by any of those eastern na- tions. In tracing it backwards to its infancy, we cannot go farther into antiquity than the times of the Grecian philosophers. As an art in the state 7 INTRODUCTION. state of some cultivation, it may be said to have been brought forth and bred up among them as a branch of natural knowledge. The æra of philosophy, as it was called, be- gan with Thales the Milesian being declared by a very general consent of the people, the most wise of all the Grecians, 480 years before Christ. The philosophers of his school, which was called the Ionian, cultivated principally natural know- ledge. Socrates, the seventh in succession of their great teachers, introduced the study of morals, and was thence said to bring down philosophy from heaven, to make men truly wise and hap- py. In the writings of his scholar and successor Plato, we see that the philosophers had carefully considered the human body, both in its organi- zation and functions; and though they had not arrived at the knowledge of the more minute and intricate parts, which required the successive la- bour and attention of many ages, they had made up very noble and comprehensive ideas of the subject in general. The anatomical descriptions of Xenophon and Plato have had the honour of being quoted by Longinus (§ xxxii.) as speci- mens of sublime writing: and the extract from Plato is still more remarkable for its containing the rudiments of the circulation of the blood. "The heart (says Plato) is the centre or knot of the blood-vessels; the spring or fountain of the blood which is carried impetuously round; the blood 8 INTRODUCTION. blood is the pabulum or food of the flesh; and, for the purpose of nourishment, the body is laid out into canals, like those which are drawn through gardens, that the blood maybe conveyed, as from a fountain, to every part of the pervious body." Hippocrates was nearly contemporary with the great philosophers of whom we have been speak- ing, about 400 years before the Christian æra. He is said to have separated the profession of philosophy and physic, and to have been the first who applied to physic alone as the business of his life. He is likewise generally supposed to be the first who wrote upon anatomy. We know of nothing that was written expressly upon the subject before; and the first anatomical dissection which has been recorded, was made by his friend Democritus of Abdera. If, however, we read the works of Hippo- crates with impartiality, and apply his accounts of the parts to what we now know of the human body, we must allow his descriptions to be im- perfect, incorrect, sometimes extravagant, and often unintelligible, that of the bones only ex- cepted. He seems to have studied these with more success than the other parts, and tells us that he had an opportunity of seeing a human skeleton. From Hippocrates to Galen, who flourished towards the end of the second century, in the decline of the Roman empire, that is, in the space 9 INTRODUCTION. space of 600 years, anatomy was greatly im- proved; the philosophers still considering it as a most curious and interesting branch of natural knowledge, and the physicians, as a principal foundation of their art. Both of them, in that interval of time, contributed daily to the com- mon stock, by more accurate and extended ob- servations, and by the lights of improving philo- sophy. As these two great men had applied very par- ticularly to the study of animal bodies, they not only made great improvements, especially in physiology, but raised the credit of natural knowledge, and spread it as wide as Alexander's empire. Few of Aristotle's writings were made public in his lifetime. He affected to say that they would be unintelligible to those who had not heard them explained at his lectures: and, ex- cept the use which Theophrastus made of them, they were lost to the public for above 130 years after the death of Theophrastus; and last came out defective from bad preservation, and corrupt- ed by men, who, without proper qualifications, presumed to correct and supply what was lost. From the time of Theophrastus, the study of natural knowledge at Athens was forever on the decline; and the reputation of the Lycæum and Academy was almost confined to the studies which are subservient to oratory and public speaking. The 10 INTRODUCTION. The other great institution for Grecian edu- cation, was at Alexandria in Egypt. The first Ptolemies, both from their love of literature, and to give true and permanent dignity to their empire, and to Alexander's favorite city, set up a grand school in the palace itself, with a museum and library, which, we may say, has been the most famed in the world. Anatomy, among other sciences, was publicly taught; and the two distinguished anatomists were Erasistratus the pu- pil and friend of Theophrastus, and Herophilus. Their voluminous works are all lost; but they are quoted by Galen almost in every page. These professors were probably the first who were au- thorized to dissect human bodies; a peculiarity which marks strongly the philosophical magnani- mity of the first Ptolemy, and fixes a great æra in the history of anatomy. And it was, no doubt, from this particular advantage which the Alex- andrians had above all others, that their school not only gained, but for many centuries pre- served, the first reputation for medical education. Ammianus Marcellinus, who lived about 650 years after the schools were set up, says, they were so famous in his time, that it was enough to se- cure credit to any physician, if he could say he had studied at Alexandria. Herophilus has been said to have anatomized 700 bodies. We must allow for exaggeration. Nay, it was said, that both he and Erasistratus made it a common practice to open living bodies, that 11 INTRODUCTION. that they might discover the more secret springs of life. But this, no doubt, was only a vulgar opinion, rising from the prejudices of mankind; and accordingly, without any good reason, such tales have been told of modern anatomists, and have been believed by the vulgar. Among the Romans, though it is probable they had physicians and surgeons from the founda- tion of the city, yet we have no account of any of these applying themselves to anatomy for a very long time. Archagathus was the first Greek physician established in Rome, and he was banished the city on account of the severity of his operations.—Asclepiades, who flourished in Rome 101 years after Archagathus, in the time of Pompey, attained such a high reputation as to be ranked in the same class with Hippocrates. He seemed to have some notion of the air in res- piration acting by its weight; and in accounting for digestion, he supposed the food to be no far- ther changed than by a comminution into ex- tremely small parts, which being distributed to the several parts of the body, is assimilated to the nature of each. One Cassius, commonly thought to be a disciple of Asclepiades, accounted for the right side of the body becoming paralytic on hurting the left side of the brain, in the same manner as has been done by the moderns, viz. by the crossing of the nerves from the right to the left side of the brain. From the time of Asclepiades to the second century, 12 INTRODUCTION. century, physicians seem to have been greatly encouraged at Rome; and, in the writings of Celsus, Rufus, Pliny, Cœlius, Aurelianus, and Aræteus, we find several anatomical observations, but mostly very superficial and inaccurate. To- wards the end of the second century lived Clau- dius Gallenus Pergamus, whose name is so well known in the medical world. He applied him- self particularly to the study of anatomy, and did more in that way than all that went before him. He seems, however, to have been at a great loss for human subjects to operate upon; and therefore his description of the parts are mostly taken from brute animals. His works contain the fullest history of anatomists, and the most complete system of the science, to be met with any where before him, or for several cen- turies after; so that a number of passages in them were reckoned absolutely unintelligible for many ages, until explained by the discoveries of suc- ceeding anatomists. About the end of the fourth century, Nimesi- us bishop of Emissa wrote a treatise on the na- ture of man, in which it is said were contained two celebrated modern discoveries; the one, the uses of the bile, boasted of by Sylvius de la Boe; and the other, the circulation of the blood. This last, however, is proved by Dr Friend, in his History of physic, p. 229. to be falsely ascribed to this author. The Roman empire beginning now to be op- pressed 13 INTRODUCTION. pressed by the barbarians, and sunk in gross su- perstition, learning of all kinds decreased; and when the empire was totally overwhelmed by those barbarous nations, every appearance of science was almost extinguished in Europe. The only remains of it were among the Arabians in Spain and in Asia.—The Saracens who came into Spain, destroyed at first all the Greek books which the Vandals had spared: but though their government was in a constant struggle and fluc- tuation during 800 years before they were driven out, they received a taste for learning from their countrymen of the east; several of their princes encouraged liberal studies; public schools were set up at Cordova, Toledo, and other towns, and translations of the Greeks into the Arabic were universally in the hands of their teachers. Thus was the learning of the Grecians trans- ferred to the Arabians. But though they had so good a foundation to build upon, this art was ne- ver improved while they were masters of the world; for they were satisfied with commenting upon Galen; and seem to have made no dissec- tions of human bodies. Abdollaliph, who was himself a teacher of anatomy, a man eminent in his time (at and before 1203) for his learning and curiosity; a great traveller, who had been bred at Bagdad, and had seen many of the great cities and prin- cipal places for study in the Saracen empire; who had a favourable opinion of original obser- vation, 14 INTRODUCTION. vation, in opposition to book-learning; who boldly corrected some of Galen's errors, and was persuaded that many more might be detect- ed; this man, we say, never made or saw, or seemed to think of a human dissection. He dis- covered Galen's errors in the osteology, by going to burying-grounds, with his students and others, where he examined and demonstrated the bones; he earnestly recommended that method of study, in preference even to the reading of Galen, and thought that many further improvements might be made; yet he seemed not to have an idea that a fresh subject might be dissected with that view. Perhaps the Jewish tenets, which the Maho- metans adopted, about uncleanliness and pollu- tion, might prevent their handling dead bodies; or their opinion of what was supposed to pass between an angel and the dead person, might make them think disturbing the dead highly sa- crilegious. Such, however, as Arabian learning was, for many ages together there was hardly any other in all the western countries of Europe. It was introduced by the establishment of the Saracens in Spain in 711, and kept its ground till the restoration of learning in the end of the 15th century. The state of anatomy in Europe, in the times of Arabian influence, may be seen by reading a very short system of anatomy drawn up by Mundinus, in the year 1315. It was ex- tracted principally from what the Arabians had preserved of Galen's doctrine; and, rude as it is, in 15 INTRODUCTION. in that age, it was judged to be so masterly a performance, that it was ordered by a public decree, that it should be read in all the schools of Italy; and it actually continued to be almost the only book which was read upon the subject for above 200 years. Cortesius gives him the credit of being the great restorer of anatomy, and the first who dissected human bodies among the moderns. A general prejudice against dissection, howe- ver, prevailed till the 16th century. The em- peror Charles V. ordered a consultation to be held by the divines of Salamanca, in order to determine whether or not it was lawful in point of conscience to dissect a dead body. In Mus- covy, till very lately, both anatomy and the use of skeletons were forbidden, the first as inhu- man, and the latter as subservient to with- craft. In the beginning of the 15th century, learn- ing revived considerably in Europe, and parti- cularly physic, by means of copies of the Greek authors brought from the sack of Constantinople; after which the number of anatomists and ana- tomical books increased to a prodigious degree. —The Europeans becoming thus possessed of the antient Greek fathers of medicine, were for a long time so much occupied in correcting the copies they could obtain, studying the meaning, and commenting upon them, that they at- tempted 16 INTRODUCTION. tempted nothing of their own, especially in ana- tomy. And here the late Dr. Hunter introduces in- to the annals of this art, a genius of the first rate, Leonardo da Vinci, who had been formerly overlooked, because he was of another profession, and because he published nothing upon the sub- ject. He is considered by the Doctor as by far the best anatomist and physiologist of his time; and was certainly the first man we know of who introduced the practice of making anatomical drawings. Vassare, in his lives of the painters, speaks of Leonardo thus, after telling us that he had com- posed a book of the anatomy of a horse, for his own study: "He afterwards applied him- self with more diligence to the human anatomy; in which study he reciprocally received and communicated assistance to Marc Antonio della Torre, an excellent philosopher, who then read lectures in Pavia, and wrote upon this subject; and who was the first, as I have heard, who began to illustrate medicine from the doctrine of Galen, and to give true light to anatomy, which till that time had been involved in clouds of darkness and ignorance. In this he availed himself exceedingly of the genius and labour of Leonardo, who made a book of studies, drawn with red chalk, and touched with a pen, with great diligence, of such subjects as he had him- self dissected; where he made all the bones, and 3 to 17 INTRODUCTION. to those he joined, in their order, all the nerves, and covered them with the muscles. And con- cerning those, from part to part, he wrote re- marks in letters of an ugly form, which are written by the left hand, backwards, and not to be understood but by those who know the method of reading them; for they are not to be read without a looking-glass. Of these pa- pers of the human anatomy, there is a great part in the possession of M. Francesco da Melzo, a Milanese gentleman, who, in the time of Leo- nardo, was a most beautiful boy, and much be- loved by him, as he is now a beautiful and gen- teel old man, who reads those writings, and care- fully preserves them, as precious relicts, toge- ther with the portrait of Leonardo, of happy memory. It appears impossible that that divine spirit should reason so well upon the arteries, and muscles, and nerves, and veins; and with such diligence of every thing, &c. &c." Those very drawings and the writings are happily found to be preserved in his Britannic Majesty's great collection of original drawings, where the Doctor was permitted to examine them; and his sentiments upon the occasion he thus expresses: "I expected to see little more than such designs in anatomy, as might be use- ful to a painter in his own profession; but I saw, and indeed with astonishment, that Leonardo had been a general and a deep student. When I consider what pains he has taken upon every B part 18 INTRODUCTION. part of the body, the superiority of his univer- sal genius, his particular excellence in mecha- nics and hydraulics, and the attention with which such a man would examine and see objects which he was to draw, I am fully persuaded that Leo- nardo was the best anatomist at that time in the world. We must give the 15th century the cre- dit of Leonardo's anatomical studies, as he was 55 years of age at the close of that century." In the beginning of the 16th century, Achil- linus and Benedictus, but particularly Beren- garius and Massa, followed out the improvement of anatomy in Italy, where they taught it, and published upon the subject. These first im- provers made some discoveries from their own dissections: but it is not surprising that they should have been diffident of themselves, and have followed Galen almost blindly, when his authority had been so long established, and when the enthusiasm for Greek authors was rising to such a pitch. Soon after this, we may say about the year 1540, the great Vesalius appeared. He was studious, laborious, and ambitious. From Brus- sels, the place of his birth, he went to Louvain, and thence to Paris, where anatomy was not yet making considerable figure, and then to Louvain to teach; from which place, very for- tunately for his reputation, he was called to Italy, where he met with every opportunity that such a such genius for anatomy desire, that is, books, subjects, 19 INTRODUCTION. subjects, and excellent draughtsmen. He was equally laborious in reading the ancients, and in dissecting bodies. And in making the compa- rison, he could not but see, that there was great room for improvement, and that many of Ga- len's descriptions were erroneous. When he was but a young man, he published a noble sys- tem of anatomy, illustrated with a great num- ber of elegant figures.—In this work he found so many occasions of correcting Galen, that his contemporaries, partial to antiquity, and jealous of his reputation, complained that he carried his turn for improvement and criticisms to licentious- ness. The spirit of opposition and emulation was presently roused; and Silvius in France, Colum- bus, Fallopius, and Eustachius in Italy, who were all in high anatomical reputation about the middle of the 16th century, endeavoured to de- fend Galen at the expence of Vesalius. In their disputes they made their appeals to the hu- man body: and thus in a few years the art was greatly improved. And Vesalius being detect- ed in the very fault which he condemned in Ga- len, to wit, describing from the dissections of brutes, and not of the human body, it exposed so fully that blunder of the older anatomists, that in succeeding times there has been little reason for such complaint.—Besides the above, he pub- lished several other anatomical treatises. He has been particularly serviceable by imposing names on the muscles, most of which are retain- B2 ed 20 INTRODUCTION. ed to this day. Formerly they were distinguish- ed by numbers, which were differently applied by almost every author. In 1561, Gabriel Fallopius, professor of ana- tomy at Padua, published a treatise of anatomy under the title of Observationes Anatomicæ. This was designed as a supplement to Vesalius; many of whose descriptions he corrects, though he always makes mention of him in an honourable manner. Fallopius made many great discoveries, and his book is well worth the perusal of every anatomist. In 1563, Bartholomæus Eustachius publish- ed his Opufcula Anatomica at Venice, which have ever since been justly admired for the ex- actness of the descriptions, and the discoveries contained in them. He published afterwards some other pieces, in which there is little of ana- tomy; but never published the great work he had promised, which was to be adorned with copperplates representing all the parts of the human body. These plates, after lying buried in an old cabinet for upwards of 150 years, were at last discovered and published in the year 1714, by Lancisi the pope's physician; who added a short explicatory text, because Eustachius's own writing could not be found. From this time the study of anatomy gradu- ally diffused itself over Europe; insomuch that for the last hundred years it has been daily im- proving by the labour of a number of pro- fessed 21 INTRODUCTION. fessed anatomists almost in every country of Eu- rope. We may form a judgment about the state of anatomy even in Italy, in the beginning of the 17th century, from the information of Cortesius. He had been professor of anatomy at Bologna, and was then professor of medicine at Massana; where, though he had a great desire to improve himself in the art, and to finish a treatise which he had begun on practical anatomy, in 24 years he could twice only procure an opportunity of dissecting a human body, and then it was with difficulties and in hurry; whereas he had ex- pected to have done so, he says, once every year, according to the custom in the famous academies of Italy. In the very end of the 16th centry, the great Harvey, as was the custom of the times, went to Italy to study medicine; for Italy was still the favourite seat of the arts: And in the very be- ginning of the 17th century, soon after Har- vey's return to England, his master in anatomy, Fabricius ab Aquapendente, published an ac- count of the valves in the veins, which he had discovered many years before, and no doubt taught in his lectures when Harvey attended him. This discovery evidently affected the establish- ed doctrine of all ages, that the veins carried the blood from the liver to all parts of the body for nourishment. It set Harvey to work upon the use of the heart and vascular systems in ani- mals; 22 INTRODUCTION. mals; and in the course of some years he was so happy as to discover, and to prove beyond all possibility of doubt, the circulation of the blood. He taught his new doctrine in his lectures about the year 1616, and printed it in 1628. It was by far the most important step that had been made in the knowledge of animal bodies in any age. It not only reflected useful lights upon what had been already found out in ana- tomy, but also pointed out the means of further investigation. And accordingly we see, that from Harvey to the present time, anatomy has been so much improved, that We may reasona- bly question if the ancients have been further outdone by the moderns in any other branch of knowledge. From one day to another there has been a constant succession of discoveries, relating either to the structure or functions of our body; and new anatomical processes, both of investiga- tion and demonstration, have been daily Invent- ed. Many parts of the body, which were not known in Harvey's time, have since then been brought to light: and of those which were known, the internal composition and functions remained unexplained; and indeed must have remained unexplicable without the knowledge of the circulation. Harvey's doctrine at first met with consider- able opposition; but in the space of about 20 years it was so generally and so warmly embrac- ed, that it was imagined every thing in physic would be explained. But time and experience have 23 INTRODUCTION. have taught us, that we still are, and probably must long continue to be, very ignorant; and that in the study of the human body, and of its diseases, there will always be an extensive field for the exercise of sagacity. After the discovery and knowledge of the circulation of the blood, the next question would naturally have been about the passage and route of nutritious part of the food or chyle from the bowels to the blood-vessels: And, by good for- tune, in a few years after Harvey had made his discovery, Asellius, an Italian physician, found out the lacteals, or vessels which carry the chyle from the intestines; and printed his account of them, with coloured prints, in the year 1627, the very year before Harvey's book came out. For a number of years after these two publi- cations, the anatomists in all parts of Europe were daily opening living dogs, either to see the lacteals or to observe the phenomena of the cir- culation. In making an experiment of this kind, Pecquet in France was fortunate enough to discover the thoracic duct, or common trunk of all the lacteals, which conveys the chyle into the subclavian vein. He printed his discovery in the year 1651. And now the lacteals having been traced from the intestines to the thoracic duct, and that duct having been traced to its termina- tion in a blood-vessel, the passage of the chyle was completely made out. The same practice of opening living animals furnish- 24 INTRODUCTION. furnished occasions of discovering the lymphatic vessels. This good fortune fell to the lot of Rudbec first, a young Swedish anatomist; and then to Thomas Bartholine, a Danish anato- mist, who was the first who appeared in print upon the lymphatics. His book came out in the year 1653, that is two years after that of Pecquet. And then it was very evident that they had been seen before by Dr. Higmore and others, who had mistaken them for lacteals. But none of the anatomists of those times could make out the origin of the lymphatics, and none of of the physiologists could give a satisfactory ac- count of their use. The circulation of the blood and the passage of the chyle having been satisfactory traced out in full grown animals, the anatomists were na- turally led next to consider how these animal processes were carried on in the child while in the womb of the mother. Accordingly the male and female organs, the appearances and contents of the pregnant uterus, the incubated egg, and every phenomenon which could illus- trate generation, became the favourite subject, for about 30 years, with the principal anatomists of Europe. Thus it would appear to have been in theory: but Dr. Hunter believes, that in fact, as Har- vey's master Fabricius laid the foundation for the discovery of the circulation of the blood by teaching him the valves of the veins, and there- by 25 INTRODUCTION. by inviting him to consider that subject; so Fa- bricius by his lectures, and by his elegant work De formato fœtu, et de formatione ovi et pulli, probably made that likewise a favourite subject with Dr. Harvey. But whether he took up the subject of generation in consequence of his discovery of the circulation, or was led to it by his honoured master Fabricius, he spent a great deal of his time in the enquiry; and published his observations in a book De generatione anama- lium, in the year 1651, that is six years before his death. In a few years after this, Swammerdam, Van Horn, Steno, and De Graaf, excited great atten- tion to the subject of generation, by their sup- posed discovery that the females of vivaparous animals have ovaria, that is, clusters of eggs in their loins, like oviparous animals; which when impregnated by the male, are conveyed into the uterus: so that a child is produced from an egg as well as a chick; with this difference, that one is hatched within, and the other without, the body of the mother. Malpighi, a great Italian genius, some time after, made considerable advances upon the sub- ject of generation. He had the good fortune to be the first who used magnifying glasses with address in tracing the first appearances in the for- mation of animals. He likewise made many other observations and improvements in the mi- nutiæ 26 INTRODUCTION. nutiæ of anatomy by his microscopical labours, and by cultivating comparative anatomy. This distinguished anatomist gave the first public specimen of his abilities, by printing a dis- sertation on the lungs anno 1661; a period so remarkable for the study of nature, that it would be injustice to pass it without particular notice. At the same time flourished Laurentius Belli- nus at Florence, and was the first who intro- duced mathematical reasoning in physic. In 1662, Simon Pauli published a treatise De Al- bandis ossibus. He had long been admired for the white skeletons he prepared; and at last discove- red his method, which was by exposing the bones all winter to the weather. Johannes Swammerdam of Amsterdam also published some anatomical treatises; but was most remarkable for his knowledge of preserv- ing the parts of bodies entire for many years, by injecting their vessels. He also published a treatise on inspiration; wherein he mentioned his having figures of all the parts of the body at big as the life, cut in copper, which he de- signed to publish, with a complete system of anatomy. These, however, were never made public by Swammerdam; but, in 1683, Go- thofridus Bidloo, professor of anatomy at Ley- den, published a work intitled Anatomia corporis humani, where all the parts were delineated in very large plates as almost as big as the life. Mr. Cowper, 27 INTRODUCTION. Cowper, an English surgeon, bought 300 co- pies of these figures; and in 1698, published them with an English text, quite different from Bidloo's Latin one; to which were added letters in Bidioo's figures, and some few figures, of Mr. Cowper's own. To this work Cowper's name was prefixed, without the least mention of Bid- loo, except on purpose to confute him. Bid- loo immediately published a very ill-natured pamphlet, called Gulielmus Cowperus citatus coram tribunali; appealing to the Royal Society, how far Cowper ought to be punished as a plagiary of the worst kind, and endeavouring to prove him an ignorant deceitful fellow. Cowper an- swered him in his own style, in a pamphlet call- ed his Vindiciæ; endeavouring to prove, either that Bidloo did not understand his own ta- bles, or that they were none of his. It was even alledged that those were the tables promis- ed by Swammerdam, and which Bidloo had got from his widow. This, however, appears to have been only an invidious surmise, there being unquestionable evidence that they were really the performance of Bidloo. Soon after, Isbrandus Diembroeck, professor of anatomy at Utrecht, began to appear as an author. His work contained very little ori- ginal; but he was at great pains to collect from others what was valuable in their writings, and his system was the common standard among ana- tomical students for many years. About the same time, Antonius Liewen- hoeck 28 INTRODUCTION. hoeck of Delft, improved considerably on Mal- pighi's use of microscopes. These two authors took up anatomy where others had dropt it; and, by this new art, they brought a number of amazing things to light. They discovered the red globules of the blood; they were ena- bled to see the actual circulation of the blood in the transparent parts of living animals, and could measure the velocity of its motion; they disco- vered that the arteries and veins had no interme- diate cells or spungy substance, as Harvey and all the preceding anatomists had supposed, but communicated one with the other by a continu- ation of the same tube. Liewenhoeck was in great same likewise for his discovery of the animalcula in the semen. Indeed there was scarcely a part of the body, solid or fluid, which escaped his examination; and he almost every where found, that what appeared to the naked eye to be rude undigest- ed matter, was in reality a beautiful and regular compound. After this period, Nuck added to our know- ledge of the absorbent system already mentioned, by his injections of the lymphatic glands; Ruysch, by his description of the valves of the lymphatic vessels; and Dr. Meckel, by his accurate ac- count of the whole system, and by tracing those vessels in many parts where they had not before been described. Besides these authors, Drs. Hunter and Monro have 29 INTRODUCTION. have called the attention of the public to this part of anatomy, in their controversy concern- ing the discovery of the office of the lympha- tics. When the lymphatic vessels were first seen and traced into the thoracic duct, it was natural for anatomists to suspect, that as the lacteals absorb- ed from the cavity of the intestines, the lym- phatics, which are similar in figure and struc- ture, might possibly do the same office with respect to other parts of the body: and accord- ingly, Dr. Glisson, who wrote in 1654, sup- poses these vessels arose from cavities, and that their use was to absorb; and Frederic Hoffman has very explicitly laid down the doctrine of the lymphatic vessels being a system of absorbents. But anatomists in general have been of a con- trary opinion; for, from experiments, particu- larly such as were made by injections, they have been persuaded that the lymphatic vessels did not arise from cavities, and did not absorb, but were merely continuations from small arteries. The doctrine, therefore, that the lymphatics, like the lacteals, were absorbents, as had been sug- gested by Glisson and by Hoffman, has been revived by Dr. Hunter and Dr. Monro, who have controverted the experiments of their pre- decessors in anatomy, and have endeavoured to prove that the lymphatic vessels are not continu- ed from arteries, but are absorbents. To this doctrine, however, several objections have 30 INTRODUCTION. have been started, particularly by Haller (Elm. Phys. 1. 24. § 2, 3.); and it has been found, that before the doctrine of the lymphatics being a system of absorbents can be established, it must first be determined whether this system is to be found in other animals besides man and quadrupeds. Mr. Hewson claims the merit of having proved the affirmative of this question, by discovering the lymphatic system in birds, fish, and amphibious animals. See Phil. Trans. vol. lviii. and lxix. —And latterly, Mr. Cruikshank has traced the ramifications of that system in almost every part of the body; and from his dissections, figures have been made and lately published to the world. To Mr. Seldon also we are much in- debted for his illustration of this system, which promises to give great satisfaction, but of which only a part has yet been published. The gravid uterus is a subject likewise which has received considerable improvements, parti- cularly relating to one very important discovery; viz. that the internal membrane of the uterus, which Dr. Hunter has named decidua, consti- tutes the exterior part of the secundines or after- birth, and separates from the rest of the uterus every time that a woman either bears a child or suffers a miscarriage. This discovery includes another, to wit, that the placenta is partly made up of an excreccence or efflorescence from the uterus itself. These discoveries are of the utmost conse- 3 quence, 31 INTRODUCTION. quence, both in the physiological question about the connection between the mother and child, and likewise in expiring the phenomena of births and abortions, as well as in regulating ob- stetrieal practice. The anatomists of this century have improv- ed anatomy, and have made the study of it much more easy by giving us more correct as well as more numerous figures. It is amazing to think of what has been done in that time. We have had four large folio books of figures of the bones, viz. Cheselden's, Albinus's, Sue's and Trew's. Of the muscles, we have had two large folio's; one from Cowper, which is elegant; and one from Albinus, which from the accuracy and labour of the work, we may suppose will never be outdone. Of the blood vessels we have a large folio from Dr. Haller. We have had one upon the nerves from Dr. Meckel, and another by Dr. Monro, junior. We have had Albinus's, Roederer's, Jenty's, and Hunter's works upon the pregnant uterus; Weitbrecht and Leber on the joints and fresh bones; Soe- merring on the brain; Zin on the eye; Cotun- nius, Mekel, junior, &c. on the ear; Walterus on the nerves of the thorax and abdomen; Dr. Monro on the bursæ mucosæ, &c. It would be endless to mention the anatomical figures that have been published in this century, of particular and smaller parts of the body, by Morgagni, Ruysch, Valsalva, Sanctorini, Hei- ster, 32 INTRODUCTION. ster, Vater, Cant, Zimmerman, Walterus, and others. Those elegant plates of the brain, however, just published by M. Vicq. d'Azyr, must not pass without notice, especially as they form part of an universal system of anatomy and physi- ology, both human and comparative, proposed to be executed in the same splendid style. Up- on the brain alone 19 folio plates are employed; of which several are coloured. The figures are delineated with accuracy and clearness; but the colouring is rather beautiful than correct. Such parts of this work as may be published, cannot fail to be equally acceptable to the anatomist and the philosopher; but the entire design is appa- rently too extensive to be accomplished within the period of a single life. In Great Britain, also, a very great anatomical work is carrying on by Andrew Bell, F.S.A.S. engraver to his Royal Highness the Prince of Wales, with the approbation of Dr. Monro, and under the in- spection of his very ingenious assistant Mr. Fyfe. It is to compose a complete illustration, both ge- neral and particular, of the human body, by a selection from the best plates of all the greatest anatomists, as well foreign as British, exhibit- ing the latest discoveries in the science, and ac- companied with copious explanations. The whole number of plates mentioned in the Pro- spectus is 240, of which 152 are already done; all in royal folio. To 33 INTRODUCTION. To the foreign treatises already mentioned may be added those recently published by Sabbatier and Plenck on anatomy in general. In Great- Britain, the writings of Keil, Douglas, Chesel- den, the first Monro, Winslow, &c. are too well known to need description. The last of these used to be recommended as a standard for the Stu- dents of anatomy: but it has of late given place to a more accurate and comprehensive system, in three volumes, published by Mr Elliot of Edinburgh, upon a plan approved of by Dr Mon- ro, and executed by Mr Fyfe. Dr Simmons of London has also obliged the world with an excellent system of anatomy; and another work, under the title of "Elements of Anatomy and the Animal Œconomy: in which the subjects are treated with uncommon elegance and perspi- cuity. In the latter part of the last century, anatomy made two great steps, by the invention of injec- tions, and the method of making what we com- monly call preparations. These two modern arts have really been of infinite use to anatomy; and besides have introduced an elegance into our ad- ministrations, which in former times could not have been supposed to be possible. They arose in Holland under Swammerdam and Ruysch, and afterwards in England under Cowper, St. André, and others, where they have been greatly improved. C The 34 INTRODUCTION. The anatomists of former ages had no other knowledge of the blood-vessels, than what they were able to collect from laborious dissections, and from examining the smaller branches of them, upon some lucky occasion, when they were found more than commonly loaded with red blood. But filling the vascular system with a bright co- loured wax, enables us to trace the large vessels with great ease, renders the smaller much more conspicuous, and makes thousands of the very minute ones visible, which from their delicacy, and the transparency of their natural contents, are otherwise imperceptible. The modern art of corroding the fleshy parts with a menstruum, and of leaving the moulded wax entire, is so exceedingly useful, and at the fame time so ornamental, that it does great honour to the ingenious inventor Dr Nicholls. The wax-work art of the moderns might de- serve notice in any history of anatomy, if the masters in that way had not been so careless in their imitation. Many of the wax-figures are so tawdry with a show of unnatural colours, and so very incorrect in the circumstances of figure, Situation, and the like, that though they strike a vulgar eye with admiration, they must appear ridiculous to an anatomist. But those figures which are cast in wax, plaster, or lead, from the real subject, and which of late years have been frequently made, are, of course, very correct in all the principal parts, and may be considered as no 35 INTRODUCTION. no insignificant acquisition to modern anatomy. The proper, or principal use of this art is, to pre- serve a very perfect likeness of such subjects as we but seldom can meet with, or cannot well preserve in a natural state; a subject in pregnan- cy, for example. The modern improved methods of preserving animal bodies, or parts of them, has been of the greatest Service to anatomy; especially in saving the time and labour of the anatomist in the nicer dissections of the small parts of the bo- dy. For now, whatever he has prepared with care, he can preserve; and the object is ready to be seen at any time. And in the same manner he can preserve anatomical curiosities, or rarities of every kind; such as, parts that are uncom- monly formed; parts that are diseased; the parts of the pregnant uterus and its contents. Large collections of such curiosities which modern anatomists are striving almost every where to procure, are of infinite service to the art, espe- cially in the hands of teachers. They give stu- dents clear ideas about many things which it is very essential to know, and yet which it is im- possible that a teacher should be able to show otherwise, were he ever so well supplied with fresh subjects. C2 § 2. View 36 INTRODUCTION. § 2. View of the Subject in general, and Plan of the following Treatise. The etymology of the word anatomy, as above given, implies simply dissection; but by this term something more is usually understood. It is every day made use of to express a know- ledge of the human body; and a person who is said to understand anatomy, is supposed to be conversant with the structure and arrangement of the different solid parts of the body. It is commonly divided into Anatomy, pro- perly so called; and Comparative Anatomy: the first of these is confined solely to the human bo- dy; the latter includes all animals, so far as a knowledge of their structure may tend to perfect our ideas of the human body. See COMPARA- TIVE Anatomy. The term anatomy may also have another and more extensive signification: it may be employ- ed to express not only a knowledge of the struc- ture and disposition of the parts but likewise of their economy and use. Considered in this light, it will seldom fail to excite the curiosity of peo- ple of taste, as a branch of philosophy; since, if it is pleasing to be acquainted with the structure of the body, it is certainly more so to discover all the springs which give life and motion to the machine, and to observe the admirable mechan- ism by which so many different functions are ex- ecuted. Astronomy and anatomy, as Dr Hunter, after Fontenelle, 37 INTRODUCTION. Fontenelle, observes, are the studies which pre- sent us with the most striking view of the two greatest attributes of the Supreme Being. The first of these fills the mind with the idea of his immensity, in the largeness, distances, and num- ber of the heavenly bodies; the last, astonishes with his intelligence and art in the variety and delicacy of animal mechanism. The human body has been commonly enough known by the name of microcosmus, or the little world; as if it did not differ so much from the universal system of nature in the symmetry and number of its parts as in their size. Galen's excellent treatise De usu partium, was composed as a prose hymn to the Creator; and abounds with as irresistible proofs of a Supreme Cause and governing Providence, as we find in modern physicotheology. And Cicero dwells more on the structure and œconomy of animals than on all the productions of nature besides, when he wants to prove the existence of the gods from the order and beauty of the universe. He there takes a survey of the body of man in a most ele- gant synopsis of anatomy, and concludes thus: "Quibus rebus expositis, satis docuisse videor, hominis natura, quanto omnes anteiret animan- tes. Ex quo debet intelligi, nec figuram situmque membrorum, nec ingenii mentisque vim talem effici potuisse fortuna." The satisfaction of mind which arises from the study of anatomy, and the influence which it must 38 INTRODUCTION. must naturally have upon our minds as philoso- phers, cannot be better conveyed than by the following passage from the fame author: "Quæ contuens animus, accepit ab his cognitionem de- orem, ex qua oritur pietas: cui conjuncta justi- tia est, reliquæque virtutes: ex quibus vita beata exsistit, par et similes deorum, nulla alia re nisi immortalitate, quæ nihil ad bene vivendum per- tinet, cedens cœlestibus." It would be endless to quote the animated passages of this sort which are to be found in the physicians, philosophers, and theologists, who have considered the structure and functions of animals with a view towards the Creator. It is a view which must strike one with a most awful conviction. Who can know and consider the thousand evident proofs of the astonishing art of the Creator, in forming and sustaining an animal body such as ours, without feeling the most plea- sant enthusiasm? Can we seriously reflect upon this awful subject, without being almost lost in adoration? without longing for another life after this, in which we may be gratified with the highest enjoyment, which our faculties and na- ture seem capable of, the seeing and comprehend- ing the whole plan of the Creator, in forming the universe and in directing all its operations? But the more immediate purposes of anatomy concern those who are to be the guardians of health, as this study is necessary to lay a foun- dation for all the branches of medicine. The more 39 INTRODUCTION. more we know of our fabric, the more reason we have to believe, that if our senses were more acute, and our judgment more enlarged, we should be able to trace many springs of life which are now hidden from us: by the same sagacity we should discover the true causes and nature of diseases; and thereby be enabled to restore the health of many, who are now, from our more confined knowledge, said to labor under incura- ble disorders. By such an intimate acquaintance with the œconomy of our bodies, we should dis- cover even the seeds of diseases, and destroy them before they had taken root in the constitu- tion. That anatomy is the very basis of surgery eve- ry body allows. It is dissection alone that can teach us, where we may cut the living body with freedom and dispatch; and where we may venture with great circumspection and delicacy; and where we must not, upon any account, at- tempt it. This informs the head, gives dexterity to the hand, and familiarizes the heart with a sort of necessary inhumanity, the use of cutting- instruments upon our fellow-creatures. Besides the knowledge of our body, through all the variety of its structure and operations in a sound state, it is by anatomy only that we can ar- rive at the knowledge of the true nature of most of the diseases which afflict humanity. The symptoms of many disorders are often equivo- cal; and diseases themselves are thence frequent- ly 40 INTRODUCTION. ly mistaken, even by sensible, experienced, and attentive physicians. But by anatomical exa- mination after death, we can with certainty find out the mistake, and learn to avoid it in any simi- lar case. This use of anatomy has been so generally adopted by the moderns, that the cases already published are almost innumerable: Mangetus, Morgagni, indeed many of the best modern writings in physic, are full of them. And if we look among the physicians of the best charac- ter, and observe those who have the art itself, rather than the craft of the profession at heart; we Shall find them constantly taking pains to pro- cure leave to examine the bodies of their pati- ents after death. After having considered the rise and progress of anatomy; the various discoveries that have been made in it, from time to time; the great number of diligent observers who have applied themselves to this art; and the importance of the study, not only for the prevention and cure of diseases, but in furnishing the liveliest proofs of divine wisdom; the following questions seem naturally to arise: For what purpose is there such a variety of parts in the human body? Why such a complication of nice and tender machine- ry? Why was there not rather a more simple, less delicate, and less expensive frame (A)? In (A) The following beautiful representation is taken from the late Dr Hunter's Introductory Lecture in Anatomy. 41 INTRODUCTION. In order to acquire a satisfactory general idea of this subject, and find a solution of all such questions, let us, in our imaginations, make a man : in other words let us suppose that the mind, or immaterial part, is to be placed in a cor- poreal fabric, in order to hold a correspondence with other material beings by the intervention of the body; and then consider, a priori, what will be wanted for her accommodation. In this in- quiry, we shall plainly see the necessity or advan- tage, and therefore the final cause, of most of the parts which we actually find in the human body. And if we consider that, in order to an- swer some of the requisites, human wit and in- vention would be very insufficient; we need not be surprised if we meet with some parts of the body whose use we cannot yet perceive, and with some operations and functions which we cannot explain. We can see that the whole bears the most striking characters of excelling wisdom and ingenuity: but the imperfect senses and capacity of man cannot pretend to reach every part of a machine, which nothing less than the intelligence and power of the Supreme Being could contrive and execute. First, then, the mind, the thinking, immate- rial agent, must be provided with a place of im- mediate residence, which shall have all the re- quisites for the union of spirit and body; ac- accordingly she is provided with the brain, where she 42 INTRODUCTION. she dwells as governor and superintendant of the whole fabric. In the next place, as she is to hold a corres- pondence with all the material beings around her, she must be supplied with organs fitted to receive the different kinds of impressions which they will make. In fact, therefore, we see that she is pro- vided with the organs of sense, as we call them: the eye is adapted to light; the ear to sound; the nose to smell; the mouth to taste; and the skin to touch. Further: She must be furnished with organs of communication between herself in the brain and those organs of sense, to give her informa- tion of all the impressions that are made upon them: and she must have organs between herself in the brain and every other part of the body, fitted to convey her commands and influence over the whole. For these purposes the nerves are actually given. They are chords, which rise from the brain, the immediate residence of the mind, and disperse themselves in branches through all parts of the body. They convey all the different kinds of sensations to the mind, in the brain; and likewise carry out from thence all her commands or influence to the other parts of the body. They are intended to be occasion- al monitors against all such impressions as might endanger the well-being of the whole, or of any particular part; which vindicates the Creator of all things, in having actually subjected us to those 43 INTRODUCTION. those many disagreeable and painful sensations which we are exposed to from a thousand acci- dents in life. Moreover, the mind, in this corporeal system, must be endued with the power of moving from place to place, that she may have intercourse with a variety of objects; that she may fly from such as are disagreeable, dangerous or hurtful, and pursue such as are pleasant or useful to her. And accordingly she is furnished with limbs, and with muscles and tendons, the instruments of mo- tion, which are found in every part of the fabric where motion is necessary. But to support, to give firmness and shape to the fabric; to keep the softer parts in their pro- per places; to give fixed points for, and the pro- per direction to its motions, as well as to protect some of the more important and tender organs from external injuries; there must be some firm prop-work interwoven through the whole. And, in fact, for such purposes the bones are given. The prop-work must not be made into one rigid fabric, for that would prevent motion. Therefore there are a number of bones. These pieces must all be firmly bound toge- ther, to prevent their dislocation. And this end is perfectly well answered by the ligaments. The extremities of these bony pieces, where they move and rub upon one another, must have smooth and slippery surfaces for easy motion. This 44 INTRODUCTION. This is most happily provided for, by the carti- lages and mucus of the joints. The interstices of all those parts must be filled up with Some Soft and ductile matter, which shall keep them in their places, unite them, and at the same time allow them to move a little upon one another. And these purposes are answered by the cellular membrane or adipose substance. There must be an outward covering over the whole apparatus, both to give it compactness and to defend it from a thousand injuries: which, in fact, are the very purposes of the skin and other integuments. Lastly, the mind being formed for society and intercourse with beings of her own kind, she must be endued with powers of expressing and communicating her thoughts by some sensible marks or signs; which shall be both easy to her- self, and admit of great variety; and according- ly she is provided with the organs and faculty of speech, by which she can throw out signs with amazing facility, and vary them without end. Thus we have built up an animal body which would seem to be pretty complete: but as it is the nature of matter to be altered and worked upon by matter; so in a very little time such a living creature must be destroyed, if there is no provision for repairing the injuries which she must commit upon herself, and those which she must be exposed to from without. Therefore a treasure of blood is actually provided in the heart and 45 INTRODUCTION. and vascular system, full of nutritious and heal- ing particles, fluid enough to penetrate into the minutest parts of the animal; impelled by the heart, and conveyed by the arteries, it washes every part, builds up what was broken down, and sweeps away the old and useless materials. Hence we see the necessity or advantage of the heart and arterial system. What more there was of this blood than enough to repair the present damages of the ma- chine, must not be lost, but should be returned again to the heart; and for this purpose the ve- nous system is actually provided. These requi- sites in the animal explain, a priori, the circula- tion of the blood. The old materials which were become useless, and are swept off by the current of blood, must be separated and thrown out of the system. Therefore glands, the organs of secretion, are given for straining whatever is rudundant, vapid, or noxious, from the mass of blood; and when strained, they are thrown out by emunctories, called organs of Excretion. But now, as the machine must be constantly wearing the reparation must be carried on with- out intermission, and the strainers must always be employed. Therefore there is actually a per- petual circulation of the blood, and the secretions are always going on. Even all this provision, however, would not be sufficient; for that store of blood would soon be 46 INTRODUCTION. be consumed, and the fabric would break down, if there were not a provision made for fresh Sup- plies. These we observe, in fact, are profusely Scattered round her in the animal and vegetable kingdoms; and she is furnished with hands, the fittest instruments that could have been contrived, for gathering them, and for preparing them in a variety of ways for the mouth. But these supplies, which we call food, must be considerably changed; they must be convert- ed into blood. Therefore she is provided with teeth for cutting and bruising the food, and with a stomach for melting it down: In short, with all the organs subservient to digestion. The finer parts of the aliments only can be useful in the constitution: these must be taken up and con- veyed into the blood, and the dregs must be thrown off. With this view the intestinal canal is actually given. It separates the nutritious part, which we call chyle, to be conveyed into the blood by the system of absorbent vessels; and the feces pass downwards, to be conducted out of the body. Now we have got our animal not only furnish- ed with what is wanted for its immediate exist- ence, but also with the powers of protracting that existence to an indefinite length of time. But its duration, we may presume, must necessarily be limited: for as it is nourished, grows, and is raised up to its full Strength and utmost perfection; so it must, in time, in common with all material beings, 47 INTRODUCTION. beings, begin to decay, and then hurry on to fi- nal ruin. Hence we see the necessity of a scheme for renovation. Accordingly wise Providence, to perpetuate as well as preserve his work, be- sides giving a Strong appetite for life and self-pre- servation, has made animals male and female, and given them such organs and passions as will secure the propagation of the species to the end of time. Thus we see, that by the very imperfect sur- vey which human reason is able to take of this subject, the animal man must necessarily be complex in his corporeal system, and in its ope- rations. He must have one great and general system, the vascular, branching through the whole for cir- culation: Another, the nervous, with its append- ages the organs of sense, for every kind of feel- ing: And a third, for the union and connection of all those parts. Besides these primary and general systems, he requires others which may be more local or con- fined: One for Strength, Support, and protection; the bony compages: Another for the requisite motions of the parts among themselves, as well as for moving from place to place; the muscu- lar part of the body: Another to prepare nourish- ment for the daily recruit of the body; the di- gestive organs: And one for propagating the spe- cies; the organs of generation. And in taking this general Survey of what would 48 INTRODUCTION. would appear, a priori, to be necessary for adapt- ing an animal to the situations of life, we observe, with great satisfaction, that man is accordingly made of such systems, and for such purposes. He has them all; and he has nothing more, ex- cept the organs of respiration. Breathing it seemeth difficult to account for a priori: we only knew it to be in fact essentially necessary to life. Notwithstanding this, when we saw all the other parts of the body, and their functions, so well accounted for, and so wisely adapted to their se- veral purposes, there could be no doubt that res- piration was so likewise: And accordingly, the discoveries of Dr Priestley have lately thrown light upon this function also, as will be shown in its proper place. Of all the different systems in the human bo- dy, the use and necessity are not more apparent, than the wisdom and contrivance which has been exerted in putting them all into the most com pact and convenient form: in disposing them so, that they shall mutually receive, and give helps to one another; and that all, or many of the parts, shall not only answer their principal end or purpose, but operate successfully and usefully in a variety of secondary ways. If we consider the whole animal machine in this light, and compare it with any machine in which human art has exerted its utmost; sup- pose the best constructed ship that ever was built, we shall be convinced beyond the possibility of 3 doubt, 49 INTRODUCTION. doubt, that there are intelligence and power far surpassing what humanity can boast of. One superiority in the natural machine is pe- culiarly striking. In machines of human con- trivance or art, there is no internal power, no principle in the machine itself, by which it can alter and accommodate itself to any injury which it may suffer, or make up any injury which ad- mits of repair. But in the natural machine, the animal body, this is most wonderfully provided for, by internal powers in the machine itself; many of which are not more certain and obvious in their effects, than they are above all human comprehension as to the manner and means of their operation. Thus, a wound heals up of it- self; a broken bone is made firm again by a cal- lus; a dead part is separated and thrown off; noxious juices are driven out by some of the emunctories; a redundancy is removed by some spontaneous bleeding; a bleeding naturally stops of itself; and a great loss of blood, from any cause, is in some measure compensated, by a con- tracting power in the vascular system, which ac- commodates the capacity of the vessels to the quantity contained. The stomach gives informa- tion when the supplies have been expended; re- presents, with great exactness, the quantity and the quality of what is wanted in the present state of the machine; and in proportion as she meets with negect, rises in her demand, urges her pe- tition in a louder tone, and with more forcible D arguments. 50 INTRODUCTION. arguments. For its protection, an animal bo- dy resists heat and cold in, a very wonderful manner, and preserves an equal temperature in a burning and in a freezing atmosphere. A farther excellence or superiority in the na- tural machine, if possible, still more astonishing, more beyond all human comprehension, than what we have been, speaking of, is the follow- ing. Besides those internal powers of self-pre- servation in each individual, when two of them co-operate, or act in concert, they are endued with powers of making other animals or ma- chines like themselves, which again are possessed of the same powers of producing others, and so of multiplying the species without end. These are powers which mock all human in- vention or imitation. They are characteristics of the divine Architect. Having premised this general account of the subject, we shall next consider the method to be observed in treating it. The study of the human body, as already no- ticed, is commonly divided into two parts. The first, which is, called Anatomy, relates to the mat- ter and structure of its parts; the second, called Physiology and Animal œconomy, relates to the principles and laws of its internal operations and functions. As the body is a compound of solids and fluids, Anatomy, is divided into, 1. The Anatomy of the solids, and 51 INTRODUCTION. 2. The Anatomy of the fluids. I. The SOLIDS, by which we mean all parts of our body, which are not fluid, are generally divided into two classes, viz. 1. The hard solids or bones. This part of anatomy is called Osteology; which signifies the doctrine of the bones. 2. The softer solids; which part is called Sarcology, viz. the doctrine of flesh. This division of the solids, we may observe, has probably taken its origin from the vulgar ob- servation, that the body is made of bone and flesh. And as there are many different kinds of what are called soft or fleshy parts, Sarcology is subdivided into, (1.) Angeiology, or the doctrine of vessels; by which is commonly understood blood vessels: (2.) Adenology, of glands: (3.) Neurology, of nerves: (4.) Myology, of muscles: and, (5.) Splanchnology, of the viscera or bowels. There is, besides, that part which treats of the organs of sense and of the integuments. This division of the solids has been here men- tioned, rather for the sake of explaining so ma- ny words, which are constantly used by anato- mists, than for its importance or accuracy. For besides many other objections that might be urg- ed, there are in the body three species of solids, viz. gristle or cartilage, hair, and nails; which are of an intermediate nature between bone and D2 flesh; 52 INTRODUCTION. flesh; and therefore cannot so properly be brought into the osteology or the sarcology. The cartilages were classed with the bones; because the greatest number of them are appendages to bones: and for the like reason the hair and the nails were classed with the integuments. II. THE FLUIDS of the human body may be divided into three kinds, which Dr. Hunter calls the crude, the general or perfect, and the local or secreted fluid. 1. By the crude fluid is meant the chyle, and whatever is absorbed at the surface of the body, in other words, what is recently taken into the body, and is not yet mixed with or converted into blood. 2. The general or perfect fluid is the blood itself, to wit, what is contained in the heart, arteries, and veins, and is going on in the round of the circulation. 3. The local or secreted, are those fluids pe- culiar to particular parts of the body, which are strained off from the blood, and yet are very different in their properties from the blood. They are commonly called secretions; and some are useful, others excrementitious. In treating of the Physiology, it is very diffi- cult to say what plan should be followed; for every method which has been yet proposed, is attended with manifest inconvenience. The powers and operations of the machine have such a dependence upon one another, such connec- tions 53 INTRODUCTION. tions and reciprocal influence, that they cannot well be understood or explained separately. In this sense our body may be compared to a circular chain of powers, in which nothing is first or last, nothing solitary or indepen- dent; so that wherever we begin, we find that there is something preceding which we ought to have known. If we begin with the brain and the nerves, for example, we shall find that these cannot exist, even in idea, with- out the heart: if we set out with the heart and vascular-system, we shall presently be sensible, that the brain and nerves must be supposed: or, should we take up the mouth, and follow the course of the aliment, we should see that the very first organ which presents itself, supposes the existence of both the heart and the brain: where- fore we shall incorporate the Physiology with the Anatomy, by attempting to explain the functions after we have demonstrated the or- gans. PART I. 54 Part I. OSTEOLOGY. PART I. OSTEOLOGY. WE begin with the bones, which may be considered as the great support of the body, tend- ing to give it shape and firmness.—But before we enter into the detail of each particular bone, it will be necessary to describe their composition and connections, and to explain the nature of the different parts which have an immediate re- lation to them; as the cartilages, ligaments, periosteum, marrow, and synovial glands. Sect. I. Of the Bones in general, with their appendages, &c. THE BONES are of a firm and hard (B) sub- stance, of a white colour, and perfectly insen- sible. They are the most compact parts of the body, and serve for the attachment and support of all the other parts. Three different substances are usually distin- guished in them; their exterior or bony part, properly so called; their spongy cells; and their reticular (B) Mr. Scheele has lately discovered that bones contain the phosphoric acid united with calcareous earth; and that to this combination they owe their firmness. 55 Part I. OSTEOLOGY. reticular substance. The first of these is form- ed of many laminae or plates, composing a firm hard substance—The spongy or cellular part is so called on account of its resemblance to a sponge, from the little cells which compose it. This substance forms almost the whole of the extre- mities of cylindrical bones. The reticular part is composed of fibres, which cross each other in different directions. This net-work forms the internal surface of those bones which have cavities. The flat bones, as those of the head, are com- posed only of the laminæ and the cellular sub- stance. This last is usually found in the middle of the bone dividing it into two plates, and is there called diplöe. Gagliardi, who pretended to have discovered an infinite number of claviculi (c), or bony processes, which he describes as traversing the la- minæ to unite them together, has endeavoured to support this pretended discovery by the ana- logy of bones to the bark of trees, in which cer- tain woody nails have been remarked; but this opinion seems to be altogether fanciful. Some writers have supposed, that the bones are formed by layers of the periosteum, which gradually ossify, in the same manner as the tim- ber is formed in trees by the hardening of the white (c) In his Anat. ossium nov. invent. illustrat. he describes four kinds of these claviculi or nails, viz. the perpendicular, oblique, headed, and crooked. 56 Part I. OSTEOLOGY. white substance that is found between the inner bark and the wood. M. Duhamel, who has adopted this opinion, fed different animals with madder and their ordinary food alternately du- ring a certain time; and he asserts, that in dis- secting their bones, he constantly observed dis- tinct layers of red and white, which corresponded with the length of time they they had lived on madder or their usual aliment. But it has since been proved by Detleff, that M. Duhamel's ex- periments were inaccurate, and that neither the periosteum nor the cartilages are tinged by the use of madder, which is known to affect the bones only. We usually consider in a bone, its body and its extremities. The ancients gave the name of diaphysis to the body or middle part, and divi- ded the extremities into apophysis and epiphysis. An apophysis, or process, as it is more common- ly called, is an eminence continued from the bo- dy of the bone, whereas an epiphysis is at first a sort of appendage to the bone, by means of an intermediate cartilage. Many epiphyses, which appear as distinct bones in the fœtus, afterwards become apophyses; for they are at length so com- pletely united to the body of the bone as not to be distinguishable from it in the adult state. It is not unusual, however, at the age of 18 and even 20 years, to find the extremities of bones still in the state of epiphysis. The names given to the processes of bones are 57 Part I. OSTEOLOGY. are expressive of their shape, size, or use; thus if a process is large and of a spherical form, it is called caput, or head; if the head is flatted, it is termed condyle. Some processes, from their resemblance to a stiletto, a breast, or the beak of a crow, are called styloid, mastoid, or coracoid: others are styled ridges or spines. The two pro- cesses of the os femoris derive their name of trochanters from their use. A bone has its cavities as well as processes. These cavities either extend quite through its substance, or appear only as depressions. The former are called foramina or holes, and these foramina are sometimes termed canals or conduits, according to their form and extent. Of the de- pressions, some are useful in articulation. These are called cotyloid when they are deep, as is the case with the os innominatum, where it receives the head of the os femoris; or glenoid when they are superficial, as in the scapula, where it re- ceives os humeri. Of the depressions that are not designed for articulation, those which have small apertures are called sinuses; others that are large, and not equally surrounded by high brims, are styled fossæ; such as are long and narrow, furrows; or if broad and superficial without brims, sinuosities. Some are called digital im- pressions, from their resemblance to the traces of a finger on soft bodies. We shall abridge this article, which is exceed- ingly diffuse in the generality of anatomical books, 58 Part I. OSTEOLOGY. books, and will endeavour to describe it with all the clearness it will allow. The BONES composing the skeleton are so con- structed, that the end of every bone is perfect- ly adapted to the extremity of that with which it is connected, and this connection forms what is called their articulation. Articulation is divided into diarthrosis, synar- throsis, and amphiarthrosis, or moveable, im- moveable, and mixed articulation. Each of the two first has its subdivisions. Thus the Diar- throsis, or moveable articulation, includes, 1. The enarthrosis, as it is called, when a large head is admitted into a deep cavity, as in the articula- tion of the os femoris with the os innominatum. 2. Arthrodia, when a round head is articulated with a supeficial cavity, as is the case of the os humeri and scapula. 3. Ginglimus, or hinge- like articulation, as in the connection of the thigh-bone with the tibia. The enarthrosis and arthrodia allow of motion to all sides; the gin- glimus only of flexion and extension. The synarthrosis, or immoveable articulation, includes, 1. The future, when the two bones are indented into each other, as is the case with the parietal bones. 2. Gomphosis, when one bone is fixed into another, in the manner the teeth are placed in their sockets. The term amphiarthrosis is applied to those articulations which partake both of the synar- throsis and diarthrosis, as is the case with the bones 59 Part I. OSTEOLOGY. bones of the vertebræ, which are capable of mo- tion in a certain degree, although they are firm- ly connected together by intermediate carti- lages. What is called symphysis is the union of two bones into one; as in the lower jaw, for in- stance, which in the fœtus consists of two dis- tinct bones, but becomes one in a more advanc- ed age, by the ossification of the uniting carti- lage. When bones are thus joined by the means or cartilages, the union is styled synchondrosis; when by ligaments, syneurosis. CARTILAGES are white, solid, smooth, and elastic substances, between the hardness of bones and ligaments, and seemingly of a fibrous tex- ture. We are not able to trace any vessels into their Substance by injection, nor are they ever found tinged in animals that have been fed with madder. They may be distinguished into, 1st. Those which are connected with the bones; and 2dly. Those which belong to other parts of the body. The first serve either to cover the ends and ca- vities of bones intended for motion, as in the articulations, where by their smoothness they fa- cilitate motions, which the bones alone could not execute with so much freedom; or they serve to unite bones together, as in the symphysis pu- bis, or to lengthem them, as in the ribs. Many of them ossifying as we advance in life, these 60 Part I. OSTEOLOGY. their number is less in the adult than in the fœtus, and of course there are fewer bones in the old than in the young subject. Of the second class of cartilages or those be- longing to the soft parts, we have instances in the larynx, where we find them useful in the formation of the voice, and for the attachment of muscles. The PERIOSTEUM is a fine membrane of a compact cellular texture, reflected from one joint to another, and serving as a common covering to the bones. It has sanguiferous and lymphatic vessels, and is supplied with nerves from the neighbouring parts. It adheres very firmly to their surface, and by its smoothness facilitates the motion of muscles. It likewise supports the ves- sels that go to be distributed through the substance of the bones, and may serve to strengthen the articulations. At the extremities of bones, where it is found covering a cartilage, it has by some been improperly considered as a distinct mem- brane, and named perichondrium. This, in its use and structure, resembles the periosteum. Where it covers the bones of the skull, it has gotten the name of pericranium. The periosteum is not a production of the dura mater, as the ancients, and after them Havers, imagined; nor are the bones formed by the ossi- fication of this membrane, at least when it is in a sound state, as some late writers have supposed. The periosteum is deficient in the teeth above the 61 Part I. OSTEOLOGY. the sockets, and in those parts of bones to which ligaments or tendons are attached. The MARROW is a fat oily substance, filling the cavities of bones. In the great cavities of long bones it is of much firmer consistence than in the cells of their spongy part. In the former it inclines somewhat to a yellowish tinge, and is of the consistence of fat; in the latter it is more fluid, and of a red colour. This diffe- rence in colour and consistence is owing to ac- cidental causes; both kinds are of the same na- ture, and may both be described under the com- mon name of marrow, though some writers give the name only to the fat-like substance, and call the other the medullary juice. The marrow is contained in a very fine and transparent membrane, which is supplied with a great number of blood vessels, chiefly from the periosteum. This membrana medullaris adheres to the inner surface of the bones, and furnishes an infinite number of minute bags or vesicles for inclosing the marrow, which is likewise sup- ported in the cavities of the bones by the long filaments of their reticular substance. Besides the vessels from the periosteum, the membrana medullaris is furnished with others, which in the long bones may be seen passing in near the extremities of the bone, and sending off numerous branches that ramify through all the vesicles of this membrane. The bones, and the cells containing the mar- row 62 Part I. OSTEOLOGY. row, are likewise furnished with lymphatics. By their means, the marrow, like the fat, may be taken up in a greater quantity than it is secreted; and hence it is that so little are found in the bones of those who die of lingering diseases. It is still a matter of controversy, Whether the marrow is sensible or not? We are certain- ly not able to trace any nerves to it; and from this circumstance, and its analogy to fat, Haller has ventured to consider it as insensible. On the other hand, Duverney asserts, that an injury done to this substance in a living animal was at- tended with great pain. In this dispute physi- ologists do not seem to have sufficiently discri- minated between the marrow itself and the mem- branous cells in which it is contained. The former, like the fat, being nothing more than a secreted, and of course an inorganized, matter, may with propriety be ranked among the insensi- ble parts, as much as inspissated mucus or any other secreted matter in the body; whereas the membrana medullaris being vascular, though it possesses but an obscure degree of feeling in a sound state, is not perfectly insensible. The marrow was formerly supposed to be in- tended for the nourishment and renewal of the bones; but this doctrine is now pretty generally and deservedly exploded. It seems probable that the marrow is to the bones what fat is to the soft parts. They both serve for some important pur- poses in the animal œconomy; but their parti- cular 63 Part I. OSTEOLOGY. cular use has never yet been clearly ascertained. The marrow, from the transudation of the oil through the bones of a skeleton, is supposed to diminish their brittleness; and Havers, who has written professedly on the bones, describes the canals by which the marrow is conveyed through every part of their substance, and di- vides them into longitudinal and transverse ones. He speaks of the first as extending through the whole length of the bone; and of the latter, as the passages by which the longitudinal ones com- municate with each other. The similiarity of these to the large cancelli in burnt bones, and the transudation of the oil through the bones of the skeleton, seems to prove that some such pas- sages do actually exist. The SYNOVIAL GLANDS are small bodies (D), supposed to be of a glandular structure, and ex- ceedingly vascular, secreting a fluid of a clear mu- cilaginous nature, which serves to lubricate the joints. They are placed in small cavities in the articulations, so as to be capable of being gently compressed by the motion of the joint, which ex- presses their juice in proportion to the degree of friction. When the synovia is wanting, or is of too thick a consistence, the joint becomes stiff and (D) It is now much doubted, however, whether the ap- pearances in the joints, which are usually called glands, are any thing more than assemblages of fat. 64 Part I. OSTEOLOGY. and incapable of flexion or extension. This is what is termed anchylosis. LIGAMENTS are white, glistening, inelastic bands, of a compact substance, more or less broad or thick, and serving to connect the bones together. They are distinguished by different names adapted to their different forms and uses. Those of the joints are called either round or bursal. The round ligaments are white, tendi- nous, and inelastic. They are strong and flexi- ble, and are found only in the joint of the knee, and in the articulation of the os femoris with the os innominatum. The bursal, or capsular liga- ments, surround the whole joint like a purse, and are to be found in the articulations which allow motion every way, as in the articulation of the arm with the scapula. Of those sacs called BURSÆ MUCOSÆ, a few were known to former anatomists, but by much the greater number have been since discovered by Dr. Monro (E), who observes, that they are to be met with in the extremities of the body on- ly; that many of them are placed entirely on the inner sides of the tendons, between these and the bones. Many others cover not only the innner, but the outer sides of the tendons, or are interposed between the tendons and external parts, as well as between those and the bones. 3 Some (E) See Description of the Bursæ Mucosæ, &c. 65 Part I. OSTEOLOGY. Some are situated between the tendons and ex- ternal parts only or chiefly, some between con- tiguous tendons, or between the tendons or the ligaments and the joints. A few such sacs are observed where the processes of bones play upon the ligaments, or where one bone plays upon another. Where two or more tendons are con- tiguous, and afterwards separate from each other, we generally find a common bursa divided into branches, with which it communicates; and a few bursæ of contiguous tendons communicate with each other.—Some, in healthy children, communicate with the cavities of the joints; and in many old people be has seen such com- munications formed by use or worn by friction, independent of disease. Their proper membrane is thin and transpa- rent, but very dense and capable of confining air or any other fluid. It is joined to the neigh- bouring parts by the common cellular substance. Betweeen the bursa and the hard substance of bone, a thin layer of cartilage or of tough mem- brane is very generally interposed. To the cel- lular substance on the outside of the bursa, the adipose substance is connected; except where the bursa covers a tendon, cartilage, or bone, much exposed to pressure or friction. In several places a mass of fat, covered with the continuation of the membrane of the bursa, projects into its cavity. The edges of this are divided into fringes. E The 66 Part I. OSTEOLOGY. The inner side of the membrane is smooth, and is extremely slippery from the liquor secret- ed in it. The structure of the bursæ bears a strong re- semblance to the capsular ligaments of the joints. 1. The inner layer of the ligament, like that of the bursæ, is thin and dense. 2. It is connected to the external ligaments by the common cellu- lar substance. 3. Between it and the bones, lay- ers of cartilage, or the articular cartilages, are in- terposed. 4. At the sides of the joints, where it is not subject to violent pressure and friction, the adipose substance is connected with the cel- lular membrane. 5. Within the cavities of the joints we observe masses of fat projecting, cover- ed with similar blood-vessels, and with similar fimbriæ hanging from their edges. 6. In the knee the upper part of such a mass of fat forms what has been called the mucilaginous gland of the joint, and the under part projects into the bursæ behind the ligament which ties the patella to the tibia. 7. The liquor which lubricates the bursæ has the same colour, consistence, and pro- perties as that of the joints, and both are affected in the same manner by heat, mineral acids, and ardent spirits. 8. In some places the bursæ con- stantly communicate with the cavities of the joints, in others they generally do so; from which we may infer a sameness of structure. When we examine the fimbriæ common to the fatty bodies of the joints and bursæ, and which have been supposed to be the ducts of glands lodged 67 Part I. OSTEOLOGY. lodged within the masses of fat, we are not able to discover any glandular appearance within them. And although we observe many vessels dispersed upon the membranes of the fatty bo- dies and fimbriæ; and that we cannot doubt that these fimbriæ consist of ducts which contain a lubricating liquor, and can even press such a li- quor from them; yet their cavities and orifices are so minute, that they are not discoverable even by the assistance of magnifying-glasses. These fimbriæ appear, therefore, to be ducts like those of urethra, which prepare a mucila- inous liquor without the assistance of any knotty gr glandular organ. Upon the whole, the synovia seems to be fur- nished by invisible exhalent arteries by the ducts of the fimbriæ, and by oil exuding from the adi- pose follicles by passages not yet discovered. The word skeleton, which by its etymology simply implies a dry preparation, is usually ap- plied to an assemblage of all the bones of an animal united together in their natural order. It is said to be a natural skeleton, when the bones are connected together by their own proper li- gaments; and an artificial one, when they are joined by any other substance, as wire, &c. The skeleton is generally divided into the head, trunk, and extremities. The first division includes the bones of the cranium and face. The bones of the trunk are the spine, ribs, ster- num, and bones of the pelvis. E2 The 68 Part I. OSTEOLOGY. The upper extremity on each side consists of the two bones of the Shoulder, viz. the scapula and clavicle; the bone of the arm, or os hu- meri; the bones of the fore-arm, and those of the hand. The lower extremity on each side of the trunk consists of the thigh-bones and the bones of the leg and foot. SECT. II. Of the Bones of the Head. The head is of a roundish figure, and some- what oval (F). Its greatest diameter is from the forehead to the occiput; its upper part is called vertex, or crown of the head; its anterior or fore-part the face; and the upper part of this sinciput, or forehead; its sides the temples; its posterior, or hind-part, the occiput; and its in- ferior part the basis. The bones of the head may be divided into those of the cranium and face. § I Bones (F) The bones of the fœtus being perfectly distinct, and the muscles in young persons not acting much, the shape of the head has been supposed to depend much on the manage- ment of children when very young. Vesalius, who has remarked the difference in people of different nations, ob- serves, for instance, that the head of a Turk is conical, from the early use of the turban; whilst that of an English- man is flattened by the chin-stay. Some of the latest phy- siologists suppose, with good reason, that this difference is chiefly owing to certain natural causes with which we are as yet unacquainted. 69 Part I. OSTEOLOGY. § I. Bones of the Cranium and face. THERE are eight bones of the cranium, viz. the coronal bone, or os frontis; the two parietal bones; or ossa bregmatis; the os occipitis; the two temporal bones; the sphenoid bone; and the os ethmoides, or cribriforme. Of these, only the os occipitis and ossa breg- matis are considered as proper to the cranium; the rest being common both to the cranium and face. These bones are all harder at their surface than in their middle; and on this account they are divided into two tables, and a middle spongy substance called diplöe. Os Frontis. In this, as in all other bones, we shall consider its figure, structure, processes, de- pressions and cavities; and the manner in which it is articulated with the other bones. The os frontis has some resemblance in shape to the shell of the cockle. Externally it is con- vex, its concave side being turned towards the brain. This bone, in the places where it is uni- ted to the temporal bones, is very thin, and has there no diplöe. It is likewise exceedingly thin in that part of the orbit of the eye which is nearest to the nose. Hence it is, that a wound in the eye, by a sword or any other pointed in- strument, is sometimes productive of immedi- ate death. In these cases,the sword passing through the 70 Part I. OSTEOLOGY. the weak part of the bone, penetrates the brain, and divides the nerves at their origin; or per- haps opens some blood-vessel, the consequences of which are soon fatal. We observe on the exterior surface of this bone five apophyses or processes, which are ea- sily to be distinguished. One of these is placed at the bottom and narrowest part of the bone, and is called the nasal process, from its support- ing the upper end of the bones of the nose. The four others are called angular or orbitar pro- cesses. They assist to form the orbits, which are the cavities on which the eyes are placed. In each of these orbits there are two processes, one at the interior or great angle, and the other at the exterior or little angle of the orbit. They are called the angular processes. Between these a ridge is extended in form of an arch, and on this the eye-brows are placed. It is called the orbitar or superciliary ridge, and in some mea- sure covers and defends the globe of the eye. There is a hole in this for the passage of the frontal vessels and nerves. This arch is inter- rupted near the nose by a small pit, in which the tendon of the musculus obliquus major of the eye is fixed. From the under part of each su- perciliary ridge a thin plate runs a considerable way backwards, and has the name of orbitar; the external and fore-part of this plate forms a sinuosity for lodging the lacrymal gland. Between the orbitar plates there is a large discontinuation of 71 Part I. OSTEOLOGY. of the bone, which is filled up by the cribriform part of the os ethmoides. On examining the inner surface of this bone at its under and middle part, we observe an ele- vation in form of a ridge, which has been called the spinous process; it ascends for some way, di- viding the bone into two considerable fossæ, in which the anterior lobes of the brain are placed. To a narrow furrow in this ridge is attached the extremity of the falx, as the membrane is called, which divides the brain into two hemispheres. The furrow becoming gradually wider, is conti- nued to the upper and back part of the bone. It has the falx fixed to it, and part of the longitudi- nal sinus lodged in it. Besides the two fossæ, there are many depressions, which appear like digital impressions, and owe their formation to the prominent circumvolutions of the brain. In the fœtus, the forehead is composed of two distinct bones; so that in them the sagittal suture reaches from the os occipitis to the nose. This bone is almost every where composed of two ta- bles and a diplöe. These two tables separating from each other under the eyes, form two cavi- ties, one on each side of the face, called the frontal sinuses. These sinuses are lined with a soft membrane, called membrana pituitaria. In these sinuses a mucus is secreted, which is con- stantly passing through two small holes into the nostrils, which it serves to moisten. The os frontis is joined by future to many of the 72 OSTEOLOGY. Part I. the bones of the head, viz. to the parietal, max- illary, and temporal bones; to the os ethmoides; os sphenoides; os unguis; and ossa nasi. The suture which connects it with the parietal bones is called the coronal suture. The PARIETAL BONES are two in number; they are very thin, and even transparent in some places. The particular figure of each of these bones is that of an irregular square, bordered with indentations through its whole circumference, ex- cept at its lower part. It will be easily conceiv- ed, that these bones which compose the superior and lateral parts of the cranium, and cover the greatest part of the brain, form a kind of vault. On their inner surface we observe the marks of the vessels of the dura mater; and at their up- per edge the groove for the superior longitudinal sinus. The ossa parietalia are joined to each other by the sagittal future; to the os sphenoides and ossa temporum by the squamous suture; to the os occipitis by the lambdoidal suture (A), so called from its resemblance to the Greek letter lambda; and to the os frontis by the coronal sut- ture. In the fœtus, the parietal bones are separated from the middle of the divided os frontis by a portion of the cranium then unossified. The (A) The lambdoidal suture is sometimes very irregular, be- ing composed of many small sutures, which surround so many little bones called ossa triquetra, though perhaps improperly, as they are not always triangular. 73 Part I. OSTEOLOGY. The occipital bone forms the posterior and inferior parts of the skull; it approaches nearly to the shape of a lozenge, and is indented throughout three parts of its circumference. There is a considerable hole in the inferior portion of this bone, called the foramen magnum, through which the medulla oblongata passes into the spine. The nervi accessorii, and vertebral arteries, likewise pass through it. Behind the condyles are two holes for the passage of cervical veins into the lateral sinuses; and above them are two others for the passage of the eighth pair and accessory nerves out of the head. At the sides, and a little on the anterior part of the fora- men magnum, are two processes, called the con- dyles, one on each side; they are of an oval fi- gure, and are covered with cartilage. The external surface of this bone has a large transverse arched ridge, under which the bone is very irregular, where it affords attachment to se- veral muscles. On examining its inner surface, we may observe two ridges in form of a cross; one ascending from near the foramen magnum to the top of the bone; the upper end of this in which the falx is fixed, is hollow, for lodging the superior longitudinal sinus, and the under end has the third process of the dura mater fixed to it. The other ridge, which runs horizontally, is likewise hollow for containing the lateral sinuses. Four fossæ are formed by the cross, two above and two below. In the former are placed the pos- terior 74 Part I. OSTEOLOGY. terior lobes of the brain, and in the latter the lobes of the cerebellum. At the basis of the cranium, we observe the cuneiform process (which is the name given to the great apophysis at the fore part of this bone); it serves for the reception of the medulla oblon- gata. The os occipitis is of greater strength and thickness than either of the other bones of the head, though irregularly so: at its inferior part, where it is thinnest, it is covered by a great num- ber of muscles. This bone, from its situation, being more lia- ble to be injured by falls, than any other bone of the head, nature has wisely given it the greatest strength at its upper part, where it is most expos- ed to danger. It is joined to the parietal bones by the lamb- doidal suture, and to the ossa temporum, by the additamentum of the temporal suture. It is like- wise connected to the os sphenoides by the cunei- form process. It is by means of the os occipitis that the head is united to the trunk, the two con- dyles of this bone being connected to the supe- rior oblique processes of the first vertebra of the neck. There are two temporal bones, one on each side.—We may distinguish in them two parts; one of which is called the squamous or scaly part, and the other pars petrosa from its hardness. This last is shaped like a pyramid. Each 75 Part I. OSTEOLOGY. Each of these divisions affords processes and ca- vities: externally there are three processes; one anterior, called the zygomatic process, one pos- terior, called the mastoid or mamillary process, from its resemblance to a nipple; and one infe- rior, called the syloid process, because it is shaped like a stiletto, or dagger. The cavities are, 1. The meatus auditorius ex- ternus. 2. A large fossa which serves for the ar- ticulation of the lower jaw; it is before the mea- tus auditorius, and immediately under the zygo- matic process. 3. The stylo-mastoid hole, so called from its situation between the styloid and mastoid processes; it is likewise styled the aquæ- duct of Fallopius, and affords a passage to the portio dura of the auditory, or seventh pair of nerves. 4. Below, and on the fore-part of the last foramen, we observe part of the jugular fossa, in which the beginning of the internal jugular vein is lodged. Anterior and superior to this fossa is the orifice of a foramen, through which passes the carotid artery. This foramen runs first upwards and then forwards, forming a kind of elbow, and terminates at the end of the os pe- trosum.—At this part of each temporal bone, we may observe the opening of the Eustachian tube, a canal which passes from the ear to the back part of the nose. In examining the internal surface of these bones, we may remark the triangular figure of their petrous part which separates two fossæ; one superior 76 Part I. OSTEOLOGY. superior and anterior; the other inferior and posterior: the latter of these composes part of the fossa, in which the cerebellum is placed; and the former, a portion of the least fossa for the basis of the brain. On the posterior side of the pars petrosa, we observe the meatus auditorus internus, into which enters the double nerve of the seventh pair. On the under side of this pro- cess, part of a hole appears, which is common to the temporal and occipital bones; through it the lateral sinus, the eighth pair, and accessory nerves, pass out of the head. The pars petrosa contains several little bones called the bones of the ear; which, as they do not enter into the formation of the cranium, shall be described when we are treating of the organs of hearing. The ossa temporum are joined to the ossa ma- larum, by the zygomatic sutures; to the pa- rietal bones, by the squamous sutures; to the os occipitis, by the lambdoidal suture; and to the sphenoid bone, by the suture of that name. Os sphenoides. This bone, from its situation amidst the other bones of the head, has sometimes been called cuneiforme. It is of a very irregular figure, and has been compared to a bat with its wings extended. It is commonly divided into its middle part or body, and its sides or wings. The fore part of the body has a spine or ridge, which makes part of the septum narium. The upper 77 Part I. OSTEOLOGY. upper part of each wing forms a share of the temple. The fore part of this belongs to the or- bit; while the under and back part, termed spinous process, is lodged in the base of the skull at the point of the pars petrosa. But two of the most remarkable processes are the ptergoid or ali- form, one on each side of the body of the bone, and at no great distance from it. Each of these processes is divided into two wings, and of these the exterior one is the widest. The other ter- minates in a hook-like process. The internal surface of this bone affords three fossæ. Two of these are formed by the wings of the bone, and make a part of the lesser fossæ of the basis of the cranium. The third, which is smaller, is on the top of the body of the bone; and is called sella turcica, from its resemblance to a Turkish saddle. This fossa, in which the pituitary gland is placed, has posteriorly and an- teriorly processes called the clinoid processes. There are twelve holes in this bone, viz. six on each side. The first is the passage of the op- tic nerve and ocular artery; the second, or large slit, transmits the third, fourth, sixth, and first part of the fifth pair of nerves with the ocular vein; the third hole gives passage to the second branch of the fifth pair; and the fourth hole to the third branch of the fifth pair of nerves. The fifth hole is the passage of the artery of the dura mater. The sixth hole is situated above the ptergoid process of the sphenoid bone; through it 78 Part I. OSTEOLOGY. it a reflected branch of the second part of the fifth pair passes. Within the substance of the os sphenoides there are two sinuses separated by a bony plate. They are lined with the pituitary membrane; and, like the frontal sinuses, separate a mucus which passes into the nostrils. The os sphenoides is joined to all the bones of the cranium; and likewise to the ossa maxillaria, ossa malarum, ossa palati, and vomer. This bone makes part of the basis of the skull, assists in forming the orbits, and affords attach- ment to several muscles. The os ethmoides is situated at the fore part of the basis of the cranium, and is of a very irre- gular figure. From the great number of holes with which it is pierced, it is sometimes called os cribriforme or sieve-like bone The os ethmoides or cribriforme consists of a middle part and two sides. The middle part is formed of a thin bony plate, in which are an infinite number of holes that afford a passage to filaments of the olfactory nerve. From the mid- dle of this plate, both on the outside and from within, there rises up a process, which may be easily distinguished. The inner one is called crista galli, from its supposed resemblance to a cock's comb. To this process the falx of the dura mater is attached. The exterior process, which has the same common basis as the crista galli, 79 Part I. OSTEOLOGY. galli, is a fine lamella which is united to the vomer; and divides the cavity of the nostrils, though unequally, it being generally a little in- clined to one side. The lateral parts of this bone are composed of a cellular substance; and these cells are so ve- ry intricate, that their figure or number cannot be described. Many writers have on this account called this part of the bone the labyrinth. These cells are externally covered with a very thin bony lamella. This part of the bone is called the os planum, and forms part of the orbit. The different cells of this bone, which are numerous, and which are every where lined with the pituitary membrane, evidently serve to en- large the cavity of the nose, in which the organ of smelling resides. This bone is joined to the os sphenoides, os frontis, ossa maxillaria, ossa palati, ossa nasi, ossa unguis, and vomer. The ancients, who considered the brain as the seat of all the humours, imagined that this viscus discharged its redundant moisture through the holes of the ethmoid bone. And the vulgar still think, that abscesses of the brain discharge them- selves through the mouth and ears, and that snuff is liable to get into the head; but neither snuff nor the matter of an abscess are more capable of passing through the cribriform bone, than the serosity which they supposed was discharged through it in a common cold.—All the holes of the 80 Part I. OSTEOLOGY. the ethmoid bone are filled up with the branches of the olfactory nerve. Its inner part is likewise covered with the dura mater, and its cells are every where lined with the pituitary membrane; so that neither matter nor any other fluid can possibly pass through this bone either externally or internally. Matter is indeed sometimes dis- charged through the nostrils; but the seat of the disease is in the sinuses of the nose, and not in the brain; and imposthumations are observed to take place in the ear, which suppurate and dis- charge themselves externally. Before we leave the bones of the head, we wish to make some general observations on its struc- ture and figure. As the cranium might have been composed of a single bone, the articulation of its several bones being absolutely without mo- tion, it may be asked perhaps, Why such a mul- tiplicity of bones, and so great number of su- tures? Many advantages may possibly arise from this plurality of bones and sutures, which may not yet have been observed. We are able, how- ever, to point out many useful ends, which could only be accomplished by this peculiarity of struc- ture.—In this, as in all the other works of na- ture, the great wisdom of the Creator is evinced, and cannot fail to excite our admiration and gra- titude. The cranium, by being divided into several bones, grows much faster and with greater faci- lity, than if it was composed of one piece only. 3 In 81 Part I. OSTEOLOGY. In the fœtus, the bones, as we have before ob- served, are perfectly distinct from each other. The ossification begins in the middle of each bone, and proceeds gradually to the circumfer- ence. Hence the ossification, and of course the increase of the head, is carried on from an infi- nite number of points at the same time, and the bones consequently approach each other in the same proportion. To illustrate this doctrine more clearly, if it can want further illustration, suppose it necessary for the parietal bones which compose the upper part of the head, to extend their ossification, and form the fore part of the head likewise.—Is it not evident, that this pro- cess would be much more tedious than it is now, when the os frontis and the parietal bones are both growing at the same time? Hence it hap- pens, that the heads of young people, in which the bones begin to touch each other, increase slowly; and that the proportionate increase of the volume of the head is greater in three months in the fœtus, than it is perhaps in twenty-four months at the age of fourteen or fifteen years. The sutures, exclusive of their advantage in suspending the processes of the dura mater, are evidently of great utility in preventing the too great extent of fractures of the skull.—Suppose, for instance, that by a fall or blow, one of the bones of the cranium becomes fractured. The fissure, which in a head composed of only one bone, would be liable to extend itself through F the 82 Part I. OSTEOLOGY. the whole of it, is checked, and sometimes per- haps stopped by the first suture it meets, and the effects of the injury are confined to the bone on which the blow was received. Ruysch indeed, and some others, will not allow the sutures to be of any such use; but cases have been met with where they seemed to have had this effect, and in young subjects their utility in this respect must be still more obvious. The spherical shape of the head seems likewise to render it more capable of resisting external vi- olence than any other shape would do. In a vault, the parts mutually support and strengthen each other, and this happens in the cranium. § 2. Proper Bones of the Face. THE FACE, which consists of a great num- ber of bones, is commonly divided into the up- per and lower jaws. The upper jaw consists of thirteen bones, exclusive of the teeth. Of these, six are placed on each side of the maxilla superior, and one in the middle. The bones, which are in pairs, are the ossa malarum, ossa maxillaria, ossa nasi, ossa unguis, ossa palati, and ossa spongiosa inferiora. The single bone is the vomer. The ossa malarum are the prominent square bones which are placed under the eyes, forming part of the orbits and the upper part of the cheeks. Each of them affords three surfaces; one exte- rior and a little convex; a second superior and concave, 83 Part I. OSTEOLOGY. concave, forming the inferior part and sides of the orbit; and a third posterior, irregular, and hollowed for the lodgement of the lower part of the temporal muscle. The angles of each bone form four processes, two of which may be called orbitar processes; of these the upper one is joined by suture to the os frontis, and that below to the maxillary bone. The third is connected with the os sphenoides by means of the transverse suture; and the fourth is joined to the zygomatic process of the tempo- ral bone, with which it forms the zygoma. The ossa maxillaria superiora. These bones, which are of a very irregular figure, are so called because they form the most considerable por- tion of the upper jaw. They are two in number, and generally remain distinct through life. Of the many processes which are to be seen on these bones, and which are connected with the bones of the face and skull, we shall describe only the most remarkable. One of these processes is at the upper and fore part of the bone, making part of the side of the nose, and called the nasal process. Another forms a kind of circular sweep at the inferior part of the bone, in which are the alveoli or sockets for the teeth: this is called the alveolar process. A third process is united to the os malæ on each side. Be- tween this and the nasal process there is a thin plate, which forms a Share of the orbit, and lies F2 over 84 Part I. OSTEOLOGY. over a passage for the superior maxillary vessels and nerves.—The alveolar process has posteriorly a considerable tuberosity on its internal surface, called the maxillary tuberosity. Behind the alveolar process we observe two horizontal lamellæ, which uniting together, form a part of the roof of the mouth, and divide it from the nose. The hollowness of the roof of the mouth is owing to this partition's being seat- ed somewhat higher than the alveolar process.— At the fore part of the horizontal lamellæ there is a hole called foramen incisivum, through which small blood-vessels and nerves go between the mouth and nose. In viewing these bones internally, we ob- serve a fossa in the inferior portion of the nasal process, which, with the os unguis and os spon- giosum inferius, forms a passage for the lachrymal duct. Where these two bones are united to each other, they project somewhat upwards and for- wards, leaving between them a furrow, into which the lower portion of the septum nasi is admitted. Each of these bones being hollow, a consider- able sinus is formed under its orbitar part. This cavity, which is usually named after Highmore, though it was described by Fallopius and others before his time, is lined with the pituitary mem- brane. It is intended for the same purposes as the 85 Part I. OSTEOLOGY. the other sinuses of the nose, and opens into the nostrils. The ossa maxillaria are connected with the greater part of the bones of the face and crani- um, and assist in forming not only the cheeks, but likewise the palate, nose, and orbits. The ossa nasi form two irregular squares. They are thicker and narrower above than below. Ex- ternally they are somewhat convex, and inter- nally slightly concave. These bones constitute the upper part of the nose. At their fore part they are united to each other, above to the os frontis, by their sides to the ossa maxillaria supe- riora, posteriorly and interiorly to the septum na- rium, and below to the cartilages that compose the rest of the nostrils. The ossa unguis. These little transparent bones owe their name to their supposed resem- blance to a finger-nail. Sometimes they are called ossa lachrymalia, from their concurring with the nasal process of each maxillary bone in forming a lodgement for the lachrymal sac and duct. The ossa unguis are of an irregular figure. Their external surface consists of two smooth parts, divided by a middle ridge. One of these parts, which is concave and nearest to the nose, serves to support the lachrymal sac and part of the lachrymal duct. The other, which is flat, forms a small part of the orbit. Each 86 Part I. OSTEOLOGY. Each of these bones is connected with the os frontis, os ethmoides, and os maxillare su- perius. The ossa palati. These bones, which are si- tuated at the back part of the roof of the mouth, between the os sphenoides and the ossa maxillaria superiora, are of a very irregular shape, and serve to form the nasal and maxillary fossa, and a small portion of the orbit. Where they are united to each other, they rise up into a spine on their in- ternal surface. This spine appears to be a con- tinuation of that of the superior maxillary bones, and helps to form the septum narium. These bones are joined to the ossa maxillaria superiora, os ethmoides, os sphenoides, and vo- mer. The vomer. This bone derives its name from its resemblance to a ploughshare. It is a long and flat bone, somewhat thicker at its back than at its fore part. At its upper part we observe a furrow extending through its whole length. The posterior and largest part of this furrow receives a process of the sphenoid bone. From this the furrow advances forwards, and becoming nar- rower and shallower, receives some part of the nasal lamella ethmoidea; the rest serves to sup- port the middle cartilage of the nose. The inferior portion of this bone is placed on the nasal spine of the maxillary and palate bones, which we mentioned in our description of the ossa palati. The 87 Part I. OSTEOLOGY. The vomer is united to the os sphenoides, os ethmoides, ossa maxillaria superiora, and ossa palati. It forms part of the septum narium, by dividing the back part of the nose into two nostrils. The ossa spongiosa inferiora. The parts which are usually described by this name, do not seem to deserve to be distinguished as distinct bones, except in young subjects. They consist of a spongy lamella in each nostril, which is united to the Spongy lamina of the ethmoid bone, of which they are by some considered as a part. Each of these lamellæ is longest from be- hind forwards; with its convex surface turn- ed towards the septum narium, and its con- cave part towards the maxillary bone, co- vering the opening of the lachrymal duct into the nose. These bones are covered with the pituita- ry membrane; and, besides their connection with the ethmoid bone, are joined to the ossa maxillaria superiora, ossa palati, and ossa un- guis. The maxilla inferior, or lower jaw, which in its shape resembles a horse-shoe, consists of two distinct bones in the fœtus; but these unite to- gether soon after birth, so as to form only one bone. The upper edge of this bone, like the os maxillare superius, has an alveolar process, fur- nished with sockets for the teeth. On 88 Part I. OSTEOLOGY. On each side, the posterior part of the bone rises almost perpendicularly into two processes, The highest of these called the coronoid process, is pointed and thin, and serves for the insertion of the temporal muscle. The other, or condy- loid process, as it is called, is shorter and thicker, and ends in an oblong rounded head, which is received into a fossa os the temporal bone, and is formed for a moveable articulation with the cranium. This joint is furnished with a move- able cartilage. At the bottom of each coronoid process, on its inner part, we observe a foramen extending under the roots of all the teeth, and terminating at the outer surface of the bone near the chin. Each of these canals transmits an ar- tery, vein, and nerve, from which branches are sent off to the teeth. The lower jaw is capable of a great variety of motion. By sliding the condyles from the cavi- ty towards the eminences on each side, we bring it horizontally forwards, as in biting; or we may bring the condyles only forward, and tilt the rest of the jaw backward, as in opening the mouth. We are likewise able to slide the con- dyles alternately backwards and forwards, from the cavity to the eminence, and vice versa, as in grinding the teeth. The cartilages, by adapt- ing themselves to the different inequalities in these several motions of the jaw, serve to secure the articulation, and to prevent any injuries from friction. The 89 Part I. OSTEOLOGY. The alveolar processes are composed of an outer and inner bony plate, united together by thin partitions, which at the fore part of the jaw divide the processes into as many sockets as there are teeth. But at the back part of the jaw, where the teeth have more than one root, we find a distinct cell for each root. In both jaws these processes begin to be formed with the teeth; they likewise accompany them in their growth, and gradually disappear when the teeth are removed. § 3. Of the Teeth. THE TEETH are bones of a particular structure, formed for the purposes of mastication and the articulation of the voice. It will be necessary to consider their composition and figure, their num- ber and arrangement, and the time and order in which they appear. In each tooth we may distinguish a body, a neck, and a root or fangs. The body of the tooth is that part which ap- pears above the gums. The root is fixed into the socket, and the neck is the middle part be- tween the two. The teeth are composed of two substances, viz. enamel and bone. The enamel, or the vi- terous or cortical part of the tooth, is a white and very hard and compact substance peculiar to the teeth, and appears fibrous or striated when broken. This substance is thickest on the grinding sur- face, 90 OSTEOLOGY. Part I. face, and becoming gradually thinner, terminates insensibly at the neck of the tooth. Ruysch * affirmed, that he could trace the arteries into the hardest part of the teeth; Liewenhoeck † sus- pected the fibres of the enamel to be so many vessels : and Monro ‡ says, he has frequently in- jected the vessels of the teeth in children, so as to make the inside of the cortex appear perfectly red. But it is certain, that it is not tinged by a madder diet; and that no injection will ever reach it, so that it has no appearance of being vascular §. The bony part, which composes the inner substance of the body, neck, and root of the tooth, resembles other bones in its structure, but it is much harder than the most compact part of bones in general. As a tooth when once formed receives no tinge from a madder diet, and as the minutest injections do not penetrate into its substance, this part of the tooth has, like the enamel, been supposed not to be vascular. But when we consider that the fangs of a tooth are inverted by a periosteum, and that the swell- ings of these fangs are analogous to the swellings of other bones, we may reasonably conclude, that there is a similarity of structure ; and that this bony part has a circulation through its sub- stance, * The saur 10. no. 27. † Arcan. Natur. continuat. Epistol. ‡ Anat. of the Human Bones. § Hunter on the Teeth. 91 Part I. OSTEOLOGY. stance, although from its hardness we are unable to demonstrate its vessels. In each tooth we find an inner cavity, into which enter an artery, vein, and nerve. This cavity begins by a small opening, and becoming larger, terminates in the body of the tooth. In advanced life this hole sometimes closes, and the tooth is of course rendered insensible. The periosteum surrounds the teeth from their fangs to a little beyond their bony sockets, where we find it adhering to the gums. This mem- brane, while it incloses the teeth, serves at the same time to line the sockets, So that it may be considered as common to both. The teeth are likewise secured in their sock- ets by means of the gums; a red, vascular, firm, and elastic substance, that possesses but little sen- sibility. In the gums of infants we find a hard ridge extending through their whole length, but no such ridge is to be seen in old people who have lost their teeth. The number of the teeth in both jaws at full maturity, usually varies from twenty-eight to thirty-two. They are commonly divided into three classes, viz. incisores, canini, and grind- ers or molares (H). The incisores are the four teeth in the fore part of each jaw. They have each (H) Mr. Hunter has thought proper to vary this division. He retains the old name incisores to the four fore teeth, but he distinguishes the canine teeth by the name of the cuspidati. The 92 Part I. OSTEOLOGY. each of them two surfaces; one anterior and convex, the other posterior and slightly concave, both of which terminate in a sharp edge. They are called incisores from their use in dividing the food. They are usually broader and thicker in the upper than in the under jaw; and, by be- ing placed somewhat obliquely, generally fall over the latter. The canini derive their name from their re- semblance to a dog's tusks, being the longest of all the teeth. We find one on each side of the incisores, so that there are two canini in each jaw. Their fang resembles that of the incisores, but is much larger; and in their shape they appear like an incisor with its edge worn off, so as to terminate in a narrow point. These teeth not being calculated for cutting and dividing the food like the incisores, or for grinding it like the molares, seem to be intend- ed for laying hold of substances (I). The molares or grinders, of which there are ten in each jaw, are so called, because from their shape and size they are fitted for grinding the food. Each of the incisores and canini is fur- nished The two teeth which are next to these, and which have been usually ranked with the molares, he calls the bicuspides; and he gives the name of grinders only to the three last teeth on each side. (I) Mr. Hunter remarks of these teeth, that we may trace in them a similarity in shape, situation, and use, from the most imperfectly carnivorous animal, which we believe to be the human species, to the lion, which is the most perfectly carnivorous. 93 Part I. OSTEOLOGY. nished only with one fang; but in the molares of the under jaw we constantly find two fangs, and in those of the upper jaw three fangs. These fangs are sometimes separated into two points, and each of these points has sometimes been de- scribed as a distinct fang. The two first of the molares, or those near- est to the canine teeth on each side, differ from the other three, and are with great propriety named bicuspides by Mr. Hunter. They have sometimes only one root, and seem to be of a middle nature between the incisores and the larger molares. The two next are much larger. The fifth or last grinder on each side is smaller and shorter than the rest; and from its not cut- ting the gum till after the age of twenty, and sometimes not till much later in life, is called dens sapientiæ. There is in the structure and arrangement of all these teeth an art which cannot be sufficiently admired. To understand it properly, it will be necessary to consider the under jaw as a kind of lever, with its fixed points at its articulations with the temporal bones:—it will be right to observe, too, that its powers arise from its different mus- cles, but in elevation chiefly from the tempora- lis and masseter; and that the aliment constitutes the object of resistance. It will appear, then, that the molares, by being placed nearest the cen- tre of motion, are calculated to press with a much greater force than the other teeth, independent of their grinding powers which they possess by means 94 Part I. OSTEOLOGY. means of the pterygoid muscles; and that it is for this reason we put between them any hard body we wish to break. The canini and incisores are placed farther from this point, and of course cannot exert so much force; but they are made for cutting and tearing the food, and this form seems to make amends for their deficiency in Strength. There are examples of children who have come into the world with two, three, and even four teeth; but these examples are very rare; and it is seldom before the seventh, eighth, or ninth month after birth, that the incisores, which are the first formed, begin to pass through the gum. The Symptoms of dentition, however, in consequence of irritation from the teeth, fre- quently take place in the fourth or fifth month. About the twentieth or twenty-fourth month, the canini and two molares make their appear- ance. The dangerous symptoms that sometimes ac- company dentition, are owing to the pressure of the teeth on the gum, which they irritate so as to excite pain and inflammation. This irri- tation seems to occasion a gradual wasting of the gum at the part, till at length the tooth makes its appearance. The symptoms are more or less alarming, in proportion to the resistance which the gum affords to the teeth, and according to the number of teeth which may chance to seek a passage at the same 95 Part I. OSTEOLOGY. same time. Were they all to appear at once, children would fall victims to the pain and ex- cessive irritation; but nature has so very wisely disposed them, that they usually appear one af- ter the other, with some distance of time between each. The first incisor that appears is generally in the lower jaw, and is followed by one in the upper jaw. Sometimes the canini, but more commonly one of the molares, begins to pass through the gum first. These 20 teeth, viz. eight incisores, four ca- nini, and eight molares, are called temporary or milk teeth, because they are all shed between the age of seven and fourteen, and are succeeded by what are called the permanent or adult teeth. The latter are of a firmer texture, and have larger fangs. These adult teeth being placed in a distinct set of alveoli, the upper sockets gradually disappear, as the under ones increase in size, till at length. the temporary, or upper teeth, having no longer any Support, consequently fall out. To these 20 teeth, which succeed the tempo- rary ones, 12 others are afterwards added, viz. three molares on each side in both jaws : and in order to make room for this addition, we find that the jaws gradually lengthen in proportion to the growth of the teeth; so that with 20 teeth, they seem to be as completely filled as they are afterwards with 32. This is the reason why 3 the 96 Part I. OSTEOLOGY. the face is rounder and and flatter in children than in adults. With regard to the formation of the teeth, we may observe, that in a fœtus of four months, the alveolar process appears only as a shallow longi- tudinal groove, divided by minute ridges into a number of intermediate depressions; in each of which we find a small pulpy substance, surround- ed by a vascular membrane. This gradually ossifies, and its lower part is lengthened out to form the fang. When the bony part of the tooth is formed, its surface begins to be incrust- ed with the enamel. How the latter is formed and deposited, we are not yet able to determine. The rudiments of some of the adult teeth be- gin to be formed at a very early period, for the pulp of one of the incisores may generally be perceived in a fœtus of eight months, and the ossification begins in it soon after birth. The first bicuspis begins to ossify about the fifth or sixth, and the second about the seventh year. The first adult grinder cuts the gum about the 12th, the second about the 18th, and the third, or dens sapientiæ, usually between the 20th and 30th year. The teeth, like other bones, are liable to be affected by disease. Their removal is likewise the natural consequence of old age ; for as we advance in life, the alveoli fill up, and the teeth, especially the incisores, fall out. When this hap- pens. 97 Part I. OSTEOLOGY. pens, the chin projects forward, and the face is much shortened. § 4. Of the Os Hyoides. (K) The os hyoides, which is placed at the root of the tongue, was so called by the the ancients on account of its supposed resemblance to the Greek letter υ. It will be necessary to distinguish in it, its bo- dy, horns, and appendices. The body, which is the middle and broadest part of the bone, is so placed that it may be ea- sily felt at the fore part of the throat. Anteri- orly it is irregularly convex, and its inner sur- face is unequally concave. Its cornua, or horns, which are flat and a little bent, being much longer than the body part, may be described as forming the sides of the υ. The appendices, or little horns, as they are called by M. Winslow, and some other writers, are two processes which rise up from the articulations of the cornua with the body, and are usually connected with the sty- loid process on each side by means of a liga- ment. G The (K) This bone is very seldom preserved with the skeleton, and cannot be included among the bones of the head, or inany other division of the skeleton. Thomas Bartholin has per- haps very properly described it among parts contained in the mouth; but the generality of anatomical writers have plac- ed it, as it is here, after the bones of the face. 98 Part I. OSTEOLOGY. The uses of this bone are to support the tongue, and afford attachment to a great number of mus- cles; some of which perform the motions of the tongue, while others act on the larynx and fauces. Sect. III. Of the Bones of the Trunk. The trunk of the skeleton consists of the spine, the thorax, and the pelvis. § 1. Of the Spine. The spine is composed of a great number of bones called vertrebæ, forming a long bony co- lumn, in figure not much unlike the letter s. This column, which extends from the head to the lower part of the body, may be said to con- sist of two irregular and unequal pyramids, uni- ted to each other in that part of the loins where the last lumbar vertebra joins the os sacrum. The vertebrae of the upper and longest pyra- mid are called true vertebræ, in contradistinction to those of the lowermost pyramid, which, from their being immoveable in the adult, are styled false vertebræ. It is upon the bones of the spine that the body turns; and it is to this cir- cumstance they owe their name, which is deriv- ed from the Latin verb vertere, to turn. The true vertebræ are divided into three classes of cervical, dorsal, and lumbar vertebræ.—The false 99 Part I. OSTEOLOGY. false vertebræ consist of the os sacrum and os oc- cygis. In each vertebra, as in other bones, it will be necessary to remark the body of the bone, its pro- cesses, and cavities. The body, which is convex before, and con- cave behind, where it assists in forming the ca- vity of the spine, may be compared to part of a cylinder cut off transversely. Each vertebra affords seven processes. The first is at the back part of the vertebra, and from its shape and direction is named the spinous pro- cess. On each side of this are two others, which, from their situation with respect to the spine, are called transverse processes. The four others are styled oblique or articular processes. They are much smaller than the spinous or transverse ones. Two of them are placed on the upper, and two on the lower part of each vertebra, ri- sing from near the basis of each transverse pro- cess. They have gotten the name of oblique pro- cesses, from their situation with respect to the processes with which they are articulated; and they are sometimes styled articular processes, from the manner in which they are articulated with each other; the two superior processes of one vertebra being articulated with the two other in- ferior processes of the vertebra above it. Each of these processes is covered with cartilage at its articulation, and their articulations with each other are by a species of ginglimus. G2 In 100 Part I. OSTEOLOGY. In each vertebra, between its body and its processes, we find a hole large enough to admit a linger. These holes or foramina, correspond with each other through all the vertebrae, and form the long bony channel in which the spinal marrow is placed. We may likewise observe four notches in each vertebra. Two of these notches are at the upper, and two at the lower part of the bone, between the oblique processes and the body of the vertebra. Each of those notches meeting with a similar opening in the vertebra above or below it, forms a foramen for the passage of blood-vessels, and of the nerves out of the spine. The bones of the spine are united together by means of a substance, which in young subjects appears to be of a ligamentous, but in adults more of a cartilaginous nature. This interver- tebral substance, which forms a kind of partition between the several vertebræ, is thicker and more flexible between the lumbar vertebræ than in the other parts of the spine, the most considerable motions of the trunk being performed on those vertebræ. This substance being very elastic, the extension and flexion of the body, and its motion backwards and forwards, or to either side, are performed with great facility. This elasti- city seems to be the reason why people who have been long standing, or have carried a considera- ble weight, are found to be shorter than when they have been long in bed. In the two first instances 101 Part I. OSTEOLOGY. instances the intervertebral cartilages (as they are usually called) are evidently more exposed to compression than when we are in bed in an ho- rizontal posture. In advanced life these cartilages become shri- velled, and of course lose much of their elasti- city. This may serve to account for the decrease in stature and the stooping forward which are usually to be observed in old people. Besides the connection of the several vertebræ by means of this intervertebral substance, there are likewise many strong ligaments, both exter- nal and internal, which unite the bones of the spine to each other. Their union is also strength- ed by a variety of strong muscles that cover and surround the spine. The bones of the spine are found to diminish in density, and to be less firm in their texture in proportion as they increase in bulk; so that the lowermost vertebræ, though the largest, are not so heavy in proportion as the upper ones. By this means the size of these bones is increased without adding to their weight: a circumstance of no little importance in a part like the spine, which, besides flexibility and suppleness, seems to require lightness as one of its essential proper- ties. In very young children, each veterbra con- sists of three bony pieces united by cartilages which afterwards ossify. There are seven vertebra of the neck—they are 102 Part I. OSTEOLOGY. are of a firmer texture than the other bones of the spine. Their transverse processes are forked for the lodgement of muscles, and at the bottom of each we observe a foramen, through which pass the cervical artery and vein. The first and se- cond of these vertebrae must be described more particularly. The first approaches almost to an oval shape—On its superior surface it has two cavities which admit the condyles of the occipi- tal bone with which it is articulated. This ver- tebra, which is called atlas from its supporting the head, cannot well be described as having either body or spinous process, being a kind of bony ring. Anteriorly, where it is articulated to the odontoid process of the second vertebra, it is very thin. On its upper surface it has two cavities which admit the condyles of the occipi- tal bone. By this connection the head is allow- ed to move forwards and backwards, but has very little motion in any other direction. The second vertebra has gotten the name of dentata, from its having, at its upper and anterior part, a process called the odontoid or tooth-like process, which is articulated with the atlas, to which this second vertebra may be said to serve as an axis. This odontoid process is of a cylindri- cal shape, somewhat flattened, however, ante- riorly and posteriorly. At its fore-part where it is received by the atlas, we may observe a smooth, convex, articulating surface. It is by means of this articulation that the head performs its 103 Part I. OSTEOLOGY. its rotatory motion, the atlas in that case moving upon this odontoid process as upon a pivot. But when this motion is in any considerable degree, or, in other words, when the head moves much either to the right or left, all the cervical verte- bræ seem to assist, otherwise the spinal marrow would be in danger of being divided transverse- ly by the first vertebra. Vertebræ of the back. The spinous process of each of the cervical vertebræ is shorter, and their articular processes more oblique, than in the other bones of the spine. These 12 vertebræ are of a middle size be- tween those of the neck and loins. At their sides we may observe two depressions, one at the upper and the other at the lower part of the bo- dy of each vertebra; which uniting with simi- lar depressions in the vertebræ above and below, form articulating surfaces, covered with cartilages for receiving the heads of the ribs; and at the fore-part of their transverse process (excepting the two last) we find an articulating surface for receiving the tuberosity of the ribs. Lumbar vertebræ. These five vertebræ differ only from those of the back in their being larger, and in having their spinous processes at a greater distance from each other. The most consider- able motions of the trunk are made on these vertebræ; and these motions could not be per- formed with so much ease, were the processes placed nearer to each other. The 104 Part I. OSTEOLOGY. The OS SACRUM which is composed of five or six pieces in young subjects, becomes one bone in more advanced age. It is nearly of a triangular figure, its inferior portion being bent a little forwards. Its superior part has two oblique processes which are articu- lated with the last of the lumbar vertebræ; and it has likewise commonly three small spinous processes, which gradually become shorter, so that the lowermost is not so long as the second, nor the second as the uppermost. Its transverse processes are formed into one oblong process, which becomes gradually smaller as it descends. Its concave or anterior side is usually smooth, but its posterior convex side has many prominences (the most remarkable of which are the spinous processes just now mentioned), which are filled up and covered with the muscular and tendinous parts behind. This bone has five pair of holes, which af- ford a passage to blood-vessels, and likewise to the nerves that are derived from the spinal marrow, which is continued even here, being lodged in a triangular cavity, that becomes smaller as it descends, and at length terminates oblique- ly at the lower part of this bone. Below the third division of the os sacrum, this canal is not completely bony as in the rest of the spine, be- ing secured as its back part only by a very strong membrane, so that a wound at this part must be extremely dangerous. The 105 Part I. OSTEOLOGY. The os sacrum is united laterally to the ossa innominata or hip-bones, and below to the coccyx. Os Coccyx. The coccyx, which, like the os sacrum, is in young people made up of three or four distinct parts, usually becomes one bone in the adult state. It serves to support the intestinum rectum; and, by its being capable of some degree of motion at its articulation with the sacrum, and being like that bone bent forwards, we are enabled to fit with ease. This bone is nearly of a triangular shape, be- ing broadest at its upper part, and from thence growing narower to its apex, where it is not bigger than the little finger. It has got its name from its supposed resem- blance a cuckow's beak. It differs greatly from the vertebræ, being commonly without any pro- cesses, and having no cavity for the spinal mar- row, or foramina for the transmission of nerves. The spine, of which we have now finished the anatomical description, is destined for many great and important uses. The medulla spinalis is lodged in its bony canal secure from external injury. It serves as a defence to the abdominal and thoracic viscera, and at the same time sup- ports the head, and gives a general firmness to the whole trunk. We have before compared it to the letter s, and its different turns will be found to render it not very 106 Part I. OSTEOLOGY. very unlike the figure of that letter.—In the neck we see it projecting somewhat forward to Support the head, which without this assistance would require a great number of muscles.—Lower down, in the thorax, we find it taking a curved direction backwards, and of course increasing the cavity of the chest. After this, in the loins, it again projects forwards in a direction with the centre of gravity, by which means we are easi- ly enabled to keep the body in an erect posture, for otherwise we should be liable to fall forward. Towards its inferior extremity, however, it again recedes backward, and thus assists in form- ing the pelvis, the name given to the cavity in which the urinary bladder, intestinum rectum, and other viscera are placed. If this bony column had been formed only of one piece, it would have been much more easi- ly fractured than it is now: and by confining the trunk to a stiff situation, a variety of motions would have been altogether prevented, which are now performed with ease by the great num- ber of bones of which it is composed. It is firm, and yet to this firmness there is added a perfect flexibility. If it be required to carry a load upon the head, the neck becomes stiff with the assistance of its muscles, and ac- commodates itself to the load, as if it was com- posed only of one bone— n stooping likewise, or turning to either side, the spine turns in itself every 107 Part I. OSTEOLOGY. every direction, as if all its bones were separated from each other. In a part of the body like the spine, that is made up of so great number of bones, and in- tended for such a variety of motion, there must be a greater danger of dislocation than fracture; but we shall find, that is very wisely guarded against in every direction by the processes belong- ing to each vertebra, and by the ligaments, car- tilages, &c. by which these bones are connect- ed with each other. § 2. Of the Bones of the Thorax. The thorax, or chest, is composed of many bones, viz. the sternum which is placed at its an- terior part, twelve ribs on each side which make up its lateral parts, and the dorsal vertebræ which constitute its posterior part. These last have been already described. The sternum is the long bone which extends itself from the upper to the lower part of the breast anteriorly, and to which the ribs and the clavicles are articulated. In children it is composed of several bones united by cartilages; but as we advance in life, most of these cartilages ossify, and the sternum in the adult state is found to consist only of three pieces, and sometimes become one bone. It is however generally described as being composed of three parts—one superior, which is broad, thick, 108 Part I. OSTEOLOGY. thick, and short; and one in the middle, which is thinner, narrower, and longer than the other. It terminates at its lower part by a third piece, which is called the xyphoid, or sword-like carti- lage, from its supposed resemblance to the blade of a sword, and because in young subjects it is commonly in a cartilaginous state. We have already observed, that this bone is articulated with the clavicle on each side. It is likewise joined to the fourteen true ribs, viz. seven on its right and seven on its left side. The ribs are bones shaped like a bow, form- ing the sides of the chest. There are twelve on each side. They are distinguished into true and false ribs: the seven upper ribs which are arti- culated to the sternum are called true ribs, and the five lower ones that are not immediately at- tached to that bone are called false ribs. On the inferior and interior surface of each rib, we observe a sinuosity for the lodgement of an artery, vein, and nerve. The ribs are not bony through their whole length, their anterior part being cartilaginous, They are articulated with the vertebræ and ster- num. Every rib (or at least the greater num- ber of them) has at its posterior part two pro- cesses; one at its extremity called the head of the rib, by meanss of which it is articulated with the body of two vertebræ; and another, call- ed its tuberosity, by which it is articulated with the transverse process of the lowest of these two vertebræ. 109 Part I. OSTEOLOGY. vertebræ. The first rib is not articulated by its extremity to two vertebræ, being simply attach- ed to the upper part of the first vertebra of the back. The seven superior or true ribs are arti- culated anteriorly with the sternum by their car- tilages; but the false ribs are supported in a dif- ferent manner—the eighth, which is the first of these ribs, being attached by its cartilages to the seventh; the ninth to the eighth, &c. The two lowermost ribs differ likewise from all the rest in the following particulars: They are articulated only with the body of the vertebra, and not with a transverse process; and anterior- ly, their cartilage is loose, not being attached to the cartilages of the other ribs; and this seems to be, because the most considerable motions of the trunk are not performed on the lumbar verte- bræ alone, but likewise on the two last vertebræ of the back; so that if these two ribs had been confined at the fore part like the other ribs, and had been likewise articulated with the bodies of two vertebræ, and with the transverse processes, the motion of the two last vertebræ, and con- sequently of the whole trunk, would have been impeded. The ribs help to form the cavity of the thorax; they afford attachment to different muscles; they are useful in respiration; and they serve as a security to the heart and lungs. § 3. Of 110 Part I. OSTEOLOGY. § 3. Of the Bones of the Pelvis. The pelvis is composed of the os sacrum, os coccygis, and two ossa innominata. The two first of these bones were included in the account of the spine, to which they more properly be- long. In children, each os innominatum is compos- ed of three distinct bones; but as we advance in life the intermediate cartilages gradually ossify, and the marks of the original separation disap- pear, so that they become one irregular bone; still however continuing to retain the names of ilium, ischium, and pubis, by which their divi- sions Were originally distinguished, and to be de- scribed as three different bones by the generality of anatomists. The os ilium forms the upper and most considerable part of the bone, the os ischium its lower and posterior portion, and the os pubis its fore part. The os ilium or haunch bone, is articulated posteriorly to the os sacrum by a firm cartilagin- ous substance, and is united to the os pubis be- fore and to the os ischium below. Its superior portion is thin, and terminates in a ridge called the crista or spine of the ilium, and more com- monly known by the name of the haunch. This crista rises up like an arch; being turned some- what outwards, so as to resemble the wings of a phaeton. Externally this bone is unequally prominent and 111 Part I. OSTEOLOGY. and hollowed for the lodgement of muscles; in- ternally we find it smooth and concave. At its lower part there is a considerable ridge en its in- ner surface. This ridge extends from the os sa- crum, and corresponds with a similar prominence both on that bone and the ischium; forms with the inner part of the ossa pubis what in midwisery is termed the brim of the pelvis. The crista, or spine, which at first is an epi- physis, has two considerable tuberosities; one anteriorly, and the other posteriorly, which is the largest of the two: These, from their pro- jecting more than the parts of the bone below them, have gotten the name of spinal processes. From the anterior spinous process, the sartorius and tensor vagina femoris muscles have their ori- gin; and below the posterior process we observe a considerable niche in the bone, which, in the recent subject, is formed into a large foramen, by means of a strong ligament that is stretched over its lower part from the os sacrum to the sharp-pointed process of the ischium. This hole affords a passage to the great sciatic nerve, and to the posterior crural vessels under the py- riform muscle, part of which likewise passes out here. The os ischium, or hip-bone, which is of a very irregular figure, constitutes the lower lateral parts of the pelvis, and is commonly divided into its body, tuberosity, and ramus. The body forms the lower and most considerable portion of the acetabulum, 112 Part I. OSTEOLOGY. acetabulum, and sends a Sharp-pointed process backwards, called the spine of the ischium. To this process the ligament adheres, which was just now spoken of, as forming a foramen for the passage of the sciatic nerve.—The tuberosity, which is the lowest part of the trunk, and sup- ports us when we sit, is large and irregular, af- fording origin to several muscles. From this tu- berosity we find the bone becoming thinner and narrower. This part, which has the name of ramus or branch, passes forwards and upwards, and concurs with the ramus of the os pubis, to form a large hole called the foramen magnum ischii, or thyroideum, as it is sometimes named, from its resemblance to a door or shield. This hole, which in the recent subject is closed by a strong membrane called the obturator ligament, affords through its whole circumference attach- ment to muscles. At its upper part where we observe a niche in the bone, it gives passage to the obturator vessels and nerves, which go to the inner part of the thigh. Nature seems every where to avoid an unnecessary weight of bone, and this foramen, no doubt, serves to lighten the bones of the pelvis. The os pubis, or share-bone, which with its fellow forms the fore-part of the pelvis, is the smallest division of the os innominatum. It is united to its fellow by means of a strong car- tilage, which forms what is called the symphysis pubis. 3 In 113 Part I. OSTEOLOGY. In each os pubis we may distinguish the body of the bone, its angle, and ramus. The body or outer part is united to the os ilium. The angle comes forward to form the symphysis, and the ramus is a thin process which unites with the ramus of the ischium, to form the foramen thy- roideum. The three bones we have described as compos- ing each os innominatum, all assist in forming the acetabulum, in which the head of the os femoris is received. This cavity is every where lined with smooth cartilage, excepting at its inner part, where we may observe a little fossa, in which are lodged the mucilaginous glands of the joint. We may likewise notice the pit or depression made by the round ligament, as it is improperly called, which, by adhering to this cavity and to the head of the thigh-bone, helps to secure the latter in the socket. These bones, which are united to each other and to the spine by many very strong ligaments, serve to support the trunk, and to connect it with the lower extremities; and at the same time to form the pelvis or bason, in which are lodg- ed the intestines and urinary bladder, and in women the uterus; so that the study of this part of osteology is of the utmost importance in mid- wifery. It is worthy of observation, that in women the os sacrum is usually shorter, broader, and H more 114 Part I. OSTEOLOGY. more hollowed, the ossa ilia more expanded, and the inferior opening of the pelvis larger than in men. Sect. IV. Of the Extremities. These parts of the Skeleton consist of the up- per extremity and the lower. § 1. Of the UPPER EXTREMITY. This consists of the Shoulder, the arm, and the hand. 1. Of the Shoulder. The shoulder consists of two bones, the cla- vicula and the scapula. The former, which is so named from its re- semblance to the key in use amongst the ancients, is a little curved at both its extremities like an italic s. It is likewise called jugulum, or collar- bone, from its situation. It is about the size of the little finger, but longer, and being of a very spongy substance is very liable to be fractured. In this, as in other long bones, we may distinguish a body and two extremities. The body is rather flattened than rounded. The anterior extremity is formed into a slightly convex head, which is nearly of a triangular shape. The inferior sur- face of the head is articulated with the sternum. The posterior extremity, which is flatter and broader than the other, is connected to a pro- cess of the scapula, called acromion. Both these 115 Part I. OSTEOLOGY. these articulations are secured by ligaments, and in that with the sternum we meet with a move- able cartilage, to prevent any injury from fric- tion. The clavicle serves to regulate the motions of the scapula, by preventing it from being brought: too much forwards, or carried too far back- wards. It affords origin to several muscles, and helps to cover and protect the subclavian vessels, which derive their name from their situation un- der this bone. The scapula, or shoulder-blade, which is near- ly of a triangular shape, is fixed to the posterior part of the true ribs, somewhat in the manner of a buckler. It is of a very unequal thickness, and, like all other broad, flat bones, is somewhat cellular. Exteriorly it is convex, and interiorly concave, to accommodate itself to the convexity of: the ribs. We observe in this bone three un- equal sides, which are thicker and stronger than the body of the bone, and are therefore termed its costæ. The largest of the three, called also the basis, is turned towards the vertebræ. An- other, which is less than the former, is below this; and the third, which is the least of the three, is at the upper part of the bone. Exter- nally the bone is elevated into a considerable spine, which rising small at the basis of the scapula, be- comes gradually higher and broader, and divides the outer surface of the bone into two fossæ. H2 The 116 Part I. OSTEOLOGY. The superior of these, which is the smallest, serves to lodge the supra spinatus muscle; and the in- ferior fossa, which is much larger than the other, gives origin to the infra spinatus. This spine terminates in a broad and flat process at the top of the shoulder, called the processus acromion, to which the clavicle is articulated. This process is hollowed at its lower part to allow a passage to the supra and infra spinati muscles. The scapula has likewise another considerable process at its upper part, which, from its resemblance to the beak of a bird, is called the coracoid process. From the outer side of this coracoid process, a strong ligament passes to the processus acromion, which prevents a luxation of the os humeri upwards. A third process begins by a narrow neck, and ends in a cavity called glenoid, for the connection of the os humeri. The scapula is articulated with the clavicle and os humeri, to which last it serves as a fulcrum; and by varying its position it affords a greater scope to the bones of the arm in their different motions. It likewise gives origin to several muscles, and posteriorly serves as a defence to the trunk. 2. Bones of the Arm. The arm is commonly divided into two parts, which are articulated to each other at the elbow. The upper part retains the name of arm, proper- ly 117 Part I. OSTEOLOGY. ly so called, and the lower part is usually called the fore arm. The arm is composed of a single bone called os humeri. This bone, which is almost of a cy- lindrical shape, may be divided into its body and its extremities. The upper extremity begins by a large, round smooth head, which is admitted into the glenoid cavity of the scapula. On the upper and fore part of the bone there is a groove for lodging the long head of the biceps muscle of the arm; and on each side of the groove, at the upper end of the bone, there is a tubercle to which the spinata muscles are fixed. The lower extremity has several processes and cavities. The principal processes are its two con- dyles, one exterior and the other interior, and of these the last is the largest. Between these two we observe two lateral protuberances, which, together with a middle cavity, form as it were a kind of pully upon which the motions of the fore-arm are chiefly performed. At each side of the condyles, as well exteriorly as interiorly, there is another eminence which gives origin to several muscles of the hand and fingers. Poste- riorly and superiorly, speaking with respect to the condyles, we observe a deep fossa which re- ceives a considerable process of the ulna: and anteriorly and opposite to this fossa, we observe another, which is much less and receives another process of the same bone. The 118 Part I. OSTEOLOGY. The body of the bone has at its upper and and anterior part a furrow which begins from be- hind the head of the bone, and serves to lodge the tendon of a muscle. The body of the os humeri is hollow through its whole length, and, like all other long bones, has its marrow. This bone is articulated at its upper part to the scapula. This articulation, which allows motion every way, is surrounded by a capsular ligament; that is sometimes torn in luxation, and becomes an obstacle to the easy reduction of the bone Its lower extremity is articulated with the bones of the fore-arm. The fore-arm is composed of two bones, the ulna and radius. The ulna or elbow-bone is much less than the os humeri, and becomes gradually smaller as it descends to the wrist. At is upper part it has two processes and two cavities. Of the two pro- cesses, the largest, which is situated posteriorly and called the olecranon, is admitted into the pos- terior fossa of the os humeri. The other process is placed anteriorly, and is called the coronoid pro- cess. In bending the arm it enters into the ante- rior fossa of the os humeri. This process being much smaller than the other, permits the fore- arm to bend inwards; whereas the olecranon, which is shaped like a hook, reaches the bottom of its fossa in the os humeri as soon as the arm becomes straight, and will not permit the fore-arm to 119 Part I. OSTEOLOGY. to be bent backwards. The ligaments likewise oppose this motion. Between the two processes we have described, there is a considerable cavity called the sygmoid cavity, divided into two fossæ by a small eminence, which passes from one process to the other; it is by means of this cavity and the two processes, that the ulna is articulated with the os humeri by ginglimus. At the bottom of the coronoid process inte- riorly, there is a small sygmoid cavity, which serves for the articulation of the ulna with the radius. The body of the ulna is of a triangular shape: Its lower extremity terminates by a small head and a little styloid process. The ulna is articu- lated above to the os humeri—both above and below to the radius, and to the wrist at its lower extremity. All these articulations are secured by means of ligaments. The chief use of this bone seems to be to support and regulate the motions of the radius. The radius, which is so named from its sup- posed resemblance to the spoke of a wheel, is placed at the inside of the fore-arm. It is some- what larger than the ulna, but not quite so long as that bone. Its upper part is cylindrical, hol- lowed superiorly to receive the outer condyle of the os humeri. Laterally it is admitted into the little sygmoid cavity of the ulna, and the cylin- drical part of the bone turns in this cavity in the motions 120 Part I. OSTEOLOGY. motions of pronation and supination (I). This bone follows the ulna in flexion and extension, and may likewise be moved round its axis in any direction. The lower extremity of the radius is much larger and stronger than its upper part; the ulna, on the contrary, is smaller and weak- er below than above; so that they serve to sup- ply each others deficiencies in both those parts. On the external side of this bone, we observe a small cavity which is destined to receive the lower end of the ulna; and its lower extremity is formed into a large cavity, by means of which it is articulated with the bones of the wrist, and on this account it is sometimes called manubrium manus. It supports the two first bones of the wrist on the side of the thumb, whereas the ulna is articulated with that bone of the wrist which corresponds with the little finger. Through the whole length both of this bone and the ulna, a ridge is observed, which af- fords attachment to an interosseous ligament. This ligament fills up the Space between the two bones. 3. Bones of the Hand. The carpus or wrist consists of eight small bones (I) The motions of pronation and supination may be easi- ly described. If the palm of the hand, for instance, is placed on the surface of a table, the hand may be said to be in a state of pronation ; but if the back part of the hand is turned to- wards the table, the hand will be then in a state of supination. 121 Part I. OSTEOLOGY. bones of an irregular shape, and disposed in two unequal rows. Those of the upper row are articulated with the bones of the fore-arm, and those of the lower one with the metacarpus. The ancient anatomists described these bones numerically; Lyserus seems to have been the first who gave to each of them a particular name. The names he adopted are founded on the fi- gure of the bones, and are now pretty general- ly received, except the first, which instead of ΧΟΤνλΟξΙδξς (the name given to it by Lyserus, on account of its sinus that admits a part of the os magnum), has by later writers been named Scap- boides or Naviculare. This, which is the outer- most of the upper row (considering the thumb as the outer side of the hand) is articulated with the radius; on its inner side it is connected with the os lunare, and below to the trapezium and trapezoides. Next to this is a smaller bone, called the os lunare; because its outer side, which is connected with the scaphoides, is shaped like a crescent. This is likewise articulated with the radius. On its inner side it joins the os cunei- forme, and anteriorly, the os magnum and os unciforme. The os cuneiforme, which is the third bone in the upper row, is compared to a wedge, from its being broader above, at the back of the hand, than it is below. Posteriorly it is articu- lated with the ulna, and anteriorly with the os unciforme. These 122 Part I. OSTEOLOGY. These three bones form an oblong articulating surface, covered by cartilage, by which the hand is connected with the fore-arm. The os pisiforme, or pea-like bone, which is smaller than the three just now described, though generally classed with the bones of the upper row, does not properly belong to either series, being placed on the under surface of the os cuneiforme, so as to project into the palm of the hand. The four bones of the second row correspond with the bones of the thumb and fingers; the first, second, and fourth, are from their shapes named trapezium, trapezoides, and unciforme; the third, from its being the largest bone of the carpus, is styled os magnum. All these bones are convex towards the back, and slightly concave towards the palm of the hand; their articulating surfaces are covered with cartilages, and secured by many strong ligaments, particularly by two ligamentous expansions, call- ed the external and internal annular ligaments of the wrist. The former extends in an oblique direction from the os pisiforme to the styloid pro- cess of the radius, and is an inch and an half in breadth; the latter or internal annular ligament is stretched from the os pisiforme and os unci- forme, to the os scaphoides and trapezium. These annular ligaments likewise serve to bind down the tendons of the wrist and fingers. The metacarpus consists of four bones, which support the fingers; externally they are a little convex, 123 Part I. OSTEOLOGY. convex, and internally somewhat concave, where they form the palm of the hand. They are hol- low, and of a cylindrical shape. At each extremity they are a little hollowed for their articulation; superiorly with the bones of the carpus, and inferiorly with the first phalanx of the fingers, in the same manner as the several phalanges of the fingers are articulated with each other. The five fingers of each hand are composed of fifteen bones, disposed in three ranks called phalanges: The bones of the first phalanx, which are articulated with the metacarpus, are the largest, and those of the last phalanx the smallest. All these bones are larger at their extremities than in their middle part. We observe at the extremities of the bones of the carpus, metacarpus, and fingers, seve- ral inequalities that serve for their articulation with each other; and these articulations are strengthened by means of the ligaments which surround them. It will be easily understood that this multipli- city of bones in the hand (for there are 27 in each hand) is essential to the different motions we wish to perform. If each finger was composed only of one bone instead of three, it would be impossi- ble for us to grasp any thing. § 2. Of 124 Part I. OSTEOLOGY. § 2. Of the LOWER EXTREMITIES. EACH lower extremity is divided into four parts, viz. the os femoris, or thigh bone: the rotula, or knee-pan; the leg and the foot. 1. Of the Thigh. The os femoris. The thigh is composed only of this bone, which is the largest and strongest we have. It will be necessary to distinguish its body and extremities: Its body, which is of a cylindrical shape, is convex before and con- cave behind, where it serves to lodge several muscles. Throughout two-thirds of its length we observe, a ridge called linea aspera, which originates from the trochanters, and after running for some way downwards, divides into two branches, that ter- minate in the tuberosities at the lower extremity of the bone. At its upper extremity we must describe the neck and smooth head of the bone, and likewise two considerable processes: The head, which forms the greater portion of a sphere unequally divided, is turned inwards, and received into the great cotyloid cavity of the os innominatum. At this part of the bone there is a little fossa to be observed, to which the round ligament is attach- ed, and which we have already described as tend- ing to secure the head of this bone in the great acetabulum. 125 Part I. OSTEOLOGY. acetabulum. The neck is almost horizontal, con- sidered with respect to its situation with the body of the bone. Of the two processes, the external one, which is the largest, is called trochanter major ; and the other, which is placed on the inside of the bone, trochanter minor. They both afford attachment to muscles. The articulation of the os femoris with the trunk is strengthened by means of a capsular ligament, which adheres every where round the edge of the great cotyloid cavity of the os innominatum, and surrounds the head of the bone. The os femoris moves upon the trunk in every direction. At the lower extremity of the bone are two processes called the condyles, and an intermedi- ate smooth cavity, by means of which it is arti- culated with the leg by ginglimus. All round the under end of the bone there is an irregular surface where the capsular ligament of the joint has its origin, and where blood-vessels go into the substance of the bone. Between the condyles there is a cavity poste- riorly, in which the blood-vessels and nerves are placed, secure from the compression to which they would otherwise be exposed in the action of bending the leg, and which would not fail to be hurtful. At the side of each condyle externally, there is a tuberosity, from whence the lateral ligaments originate, which are extended down to the tibia. A 126 Part I. OSTEOLOGY. A ligament likewise arises from each condyle posteriorly. One of these ligaments passes from the right to the left, and the other from the left to the right, so that they intersect each other, and for that reason are called the cross li- gaments. The lateral ligaments prevent the motion of the leg upon the thigh to the right or left; and the cross ligaments, which are also attached to the tibia, prevent the latter from being brought forwards. In new-born children all the processes of this bone are cartilaginous. 2. The Rotula, or Knee-pan. The rotula, patella, or knee-pan, as it is dif- ferently called, is a flat bone about four or five inches in circumference, and is placed at the fore- part of the joint of the knee. In its shape it is somewhat like the common figure of the heart, with its point downward. It is thinner at its edge than in its middle part; at its fore-part it is smooth and somewhat con- vex ; its posterior surface, which is more un- equal, affords an elevation in the middle which is admitted between the two condyles of the os femoris. This bone is retained in its proper situation by a strong ligament which every where surrounds it, and adheres both to the tibia and os femoris; it 127 Part I. OSTEOLOGY. it is likewise firmly connected with the tibia by means of a strong tendinous ligament of an inch in breadth, and upwards of two inches in length, which adheres to the lower part of the patella, and to the tuberosity at the upper end of the tibia. On account of this connection, it is very pro- perly considered as an appendage to the tibia, which it follows in all its motions, so as to be to it what the olecranon is to the ulna. There is this difference, however, that the olecranon is a fixed process; whereas the patella is moveable, being capable of sliding from above downwards and from below upwards. This mobility is es- sential to the rotatory motion of the leg. In very young children this bone is entirely cartilaginous. The principal use of the patella seems to be to defend the articulation of the the knee from external injury; it likewise tends to increase the power of the extensor muscles to the leg, by re- moving their direction farther from the centre of motion in the manner of a pulley. 3. Of the leg. The leg is composed of two bones: Of these the inner one, which is the largest, is called ti- bia; the other is much smaller, and named fi- bula. The TIBIA, which is so called from its resem- blance to the musical pipe of the ancients, has three 128 Part I. OSTEOLOGY. three surfaces, and is not very unlike a triangu- lar prism. Its posterior surface is the broadest; anteriorly it has a considerable ridge called the shin, between which and the skin there are no muscles. At the upper extremity of this bone are two surfaces, a little concave, and separated from each other by an intermediate elevation. The two little cavities receive the condyles of the os femoris, and the eminence between them is admitted into the cavity which we spoke of as being between the two condyles; so that this articulation affords a specimen of the com- plete ginglimus. Under the external edge of the upper end of this bone is a circular flat sur- face, which receives the head of the fibula. At the lower and inner portion of the tibia, we observe a considerable process called mal- leolus internus. The basis of the bone termi- nates in a large tranverse cavity, by which it is articulated with the uppermost bone of the foot. It has likewise another cavity at its lower end and outer side, which is somewhat oblong, and receives the lower end of the fibula. The tibia is hollow through its whole length. The FIBULA is a small long bone situated on the outside of the tibia. Its superior extremity does not reach quite so high as the upper part of the tibia, but its lower end descends somewhat lower. Both above and below, it is articulated with the tibia by means of the lateral cavities we noticed in our description of that bone. 3 Its 129 Part I. OSTEOLOGY. Its lower extremity is stretched out into a co- ronoid process, which is flattened at its inside, and is convex externally, forming what is called the malleolus externus, or outer ancle. This is rather lower than the malleolus internus of the tibia. The body of this bone, which is irregularly triangular, is a little hollow at its internal sur- face, which is turned towards the tibia; and it affords like that bone, through its whole length, attachment to a ligament, which from its situa- tion is called the interosseous ligament. 4. Of the foot. The foot consists of the tarsus, metatarsus, and toes. The TARSUS is composed of seven bones, viz. the astragalus, or os calcis, os naviculare, os cuboides, and three others called cuneiform bones. The ASTRAGALUS is a large bone with which both the tibia and fibula are articulated. It is the uppermost bone of the foot; it has several surfaces to be considered; its upper, and some- what posterior part, which is smooth and con- vex, is admitted into the cavity of the tibia. Its lateral parts are connected with the malleoli of the two bones of the leg; below, it is articula- ted with the os calcis, and its anterior surface is I received 130 Part I. OSTEOLOGY. received by the os naviculare. All these arti- culations are secured by means of ligaments. The os calcis, or calcaneum, which is of a very irregular figure, it is the largest bone of the foot. Behind, it is formed into a consider- able tuberosity called the heel; without this tu- berosity, which supports us in an erect posture, and when we walk, we should be liable to fall backwards. On the internal surface of this bone, we ob- serve a considerable sinuosity, which affords a passage to the tendon of a muscle: and to the pos- terior part of the os calcis, a strong tendinous cord called tendo achillis (M) is attached, which is formed by the tendons of several muscles uni- ted together. The articulation of this with the other bones is secured by means of ligaments. The OS NAVICULARE, or scaphoides, (for these two terms have the same signification), is so called on account of its resemblance to a little bark. At its posterior part, which is concave, it receives the astragalus; anteriorly it is articu- lated with the cuneiform bones, and laterally it is connected with the os cuboides. The OS CUBOIDES forms an irregular cube. Posteriorly it is articulated with the os calcis; anteriorly it supports the two last bones of the metatarsus, and laterally it joins the third cunei- form bone and the os naviculare. Each (M) This tendon is sometimes ruptured by jumping, danc- ing, or other violent efforts. 131 Part I. OSTEOLOGY. Each of the OSSA CUNEIFORMIA, which are three in number, resembles a wedge, and from this similitude their name is derived. They are placed next to the metatarsus by the sides of each other, and are usually distinguished into os cu- neiforme externum, medium or minimum, and in- ternum or maximum. The superior surface of these bones, from their wedge-like shape, is broader than that which is below, where they help to form the sole of the foot; posteriorly they are united to the os naviculare, and ante- riorly they support the three first metatarsal bones. When these seven bones composing the tarsus are viewed together in the skeleton, they appear convex above, where they help to form the up- per part of the foot; and concave underneath, where they form the hollow of the foot, in which the vessels, tendons and nerves of the foot are placed secure from pressure. They are united to each other by very strong ligaments, and their articulation with the foot is secured by a caspular and two lateral ligaments; each of the latter is covered by an annular liga- ment of considerable breadth and thickness, which serves to bind down the tendons of the foot, and at the same time to strengthen the articula- tion. The os cuneiforme externum is joined laterally to the os cuboides. These bones complete our account of the tar- I2 sus. 132 Part I. OSTEOLOGY. sus. Though what we have said of this part of the osteology has been very simple and concise, yet many readers may not clearly understand it: but if they will be pleased to view these bones in their proper situation in the skeleton, all that we have said of them will be easily understood. The METATARUS is made up of five bones, whereas the metacarpus consists only of four. The cause of this difference is, that in the hand the last bone of the thumb is not included among the metacarpal bones; whereas in the foot the great toe has only two bones. The first of these bones supports the great toe and is much larger than the rest, which nearly resemble each other in size. These bones are articulated by one extremity with the cuneiform bones and the os cuboides, and by their other end with the toes. Each of the TOES, like the fingers, consists of three bones, except the great toe, which is formed of two bones. Those of the other four are distinguished into three phalanges. Although the toes are more confined in their motion than the fin- gers, yet they appear to be perfectly fitted for the purposes they are designed for. In walking, the toes bring the centre of gravity perpendicular to the advanced foot; and as the soles of the foot are naturally concave, we can at pleasure increase this cavity, and form a kind of vault, which ad- justs itself to the different inequalities that occur to us in walking; and which, without this mode of 133 Part I. OSTEOLOGY. of arrangement, would incommode us exceed- ingly, especially when bare-footed. § 4. Of the Ossa SESAMOIDEA. BESIDES the bones we have already describ- ed, there are several small ones that are met with only in the adult skeleton, and in persons who are advanced in life; which, from their suppos- ed general resemblance to the seeds of the sesa- mum, are called ossa sesamoidea. They are com- monly to be seen at the first joint of the great toe, and sometimes at the joints of the thumb; they are likewise now and then to be found at the lower extremity of the fibula, upon the con- dyles of the thigh-bone, under the os cuboides of the tarsus, and in other parts of the body. Their size and number seem constantly to be in- creased by age and hard labour; and as they are generally found in situations where tendons and ligaments are most exposed to the action of muscles, they are now generally considered as ossified portions of ligaments or tendons. The upper surface of these bones is usually convex, and adherent to the tendon that covers it; the side which is next to the joint is smooth and flat. Though their formation is accidental, yet they seem to be of some use, by raising the tendons farther from the centre of motion, and consequently increasing the power of the mus- cles. In the great toe and thumb they are like- wise 134 Part I. OSTEOLOGY. wise useful, by forming a groove for the flexor tendons. EXPLANATION of the PLATES of OSTEOLOGY. PLATE XIX. FIG. 1. A front view of the MALE SKELETON. A, The os frontis. B, The os parietale. C, The coronal suture. D, The squamous part of the temporal bones. E, The squamous suture. F, The zygoma. G, The mastoid process. H, The temporal process of the spenoid bone. I, The orbit. K, The os malæ. L, The os max- illare superius. M. Its nasal process. N, The ossa nasi. O, The os unguis, P, The maxilla in- ferior. Q, The teeth, which are sixteen in number in each jaw. R, The seven cervical ver- tebrae, with their intermediate cartilages. S, Their transverse processes. T, The twelve dor- sal vertebrae, with their intermediate cartilages, U, The five lumbar vertebræ. V, Their trans- verse processes. W, The upper part of the os sacrum. X, Its lateral parts. The holes seen on its fore part are the passages of the under- most spinal nerves and small vessels. Opposite to the holes, the marks of the original divisions of the bone are seen. Y, The os ilium. Z, Its crest or spine. a, The anterior spinous pro- cesses. 135 Part I. OSTEOLOGY. esses. b, The brim of the pelvis, c, The is- hiatic niche. d, The os ischium. e, Its tube- osity. f, Its spinous process. g, Its crus. h, The foramen thyroideum. i, The os pubis. k, The symphysis pubis. l, The crus pubis, m, The acetabulum, n, The seventh or last true rib. o, The twelfth or last false rib. p, The up- per end of the sternum, q, The middle piece. , The under end, or cartilage ensiformis. s, The clavicle. t, The internal surface of the scapula. u, Its acromion, v, Its coracoid process. w, Its cervix. x, The glenoid cavity, y, The os hu- meri. z, Its head, which is connected to the glenoid cavity. 1, Its external tubercle. 2, Its internal tubercle. 3, The groove for lodging the long head of the biceps muscle of the arm, 4, The internal condyle. 5, The external con- dyle. Between 4 and 5, the trochlea. 6. The radius. 7, Its head. 8, Its tubercle. 9, The ulna. 10, Its coronoid process. 11, 12, 13, 14, 15, 16, 17, 18, The carpus ; composed of os na- viculare, os lunare, os cuneiforme, os pisiforme, os trapezium, os trapezoides, os magnum, os unciforme. 19, The five bones of the metacar- pus, 20, The two bones of the thumb. 21, The three bones of each of the fingers. 22, The os femoris. 23, Its head. 24, Its cervix. 25, The trochanter major. 26, The trochanter, mi- nor. 27, The internal condyle. 28, The ex- ernal condyle. 29, The rotula. 30, The tibia. 31, Its head. 32, Its tubercle. 33, Its spine. 34, The 136 Part I. OSTEOLOGY. 34, The malleolus internus. 35, The fibula. 36, Its head. 37. The malleolus externus. The tarsus is composed of, 38, The astragalus; 39, The os calcis; 40, The os naviculare; 41, Three ossa cuneiformia, and the os cuboides, which is not seen in this figure. 42, The five bones of the metatarsus. 43, The two bones of the great toe. 44, The three bones of each of the small toes. FIG. 2. A front view of the SKULL. A, The os frontis. B, The lateral part of the os frontis, which gives origin to part of the tem- poral muscle. C, The superciliary ridge. D, The superciliary hole through which the frontal vessels and nerves pass. EE, The orbitar pro- cesses. F, The middle of the transverse suture. G, The upper part of the orbit. H, The fora- men opticum. I, The foramen lacerum. K, The inferior orbitar fissure. L, The os unguis M, The ossa nasi. N, The os maxillare superius. O, Its nasal process. P, The external orbitar hole through which the superior maxillary ves- sels and nerves pass. Q, The os malæ. R, A passage for small vessels into, or out of the orbit. S, The under part of the left nostril. T, The septum narium. U. The os spongiosum supe- rius. V, The os spongiosum infernus. W, The edge of the alveoli, or spongy sockets, for the teeth. 137 Part I. OSTEOLOGY. teeth. X, The maxilla inferior. Y, The pas- sage for the inferior maxillary vessels and nerves. FIG. 3. A side-view of the SKULL. A, The os frontis. B, The coronal suture. C, The os parietale. D, An arched ridge which gives origin to the temporal muscle. E, The squamous suture. F, The squamous part of the temporal bone; and, farther forwards, the tem- poral process of the sphenoid bone, G, The zy- gomatic process of the temporal bone. H, The zygomatic suture. I, The mastoid process of the temporal bone. K, The meatus auditorius ex- ternum L, The orbitar plate of the frontal bone, under which is seen the transverse suture. M, The pars plana of the ethmoid bone. N, The os unguis. O, The right os nasi. P, The su- perior maxillary bone. Q, Its nasal process. R, The two dentes incisores. S, The dens ca- ninus. T, The two molares. U, The three large molares. V, The os malæ. W, The lower jaw. X, Its angle. Y, The coronoid process. Z, The condyloid process, by which the jaw is articulated with the temporal bone. FIG. 4. The posterior and right side of the SKULL. A, The os frontis. BB, The ossa parietalia. C, The sagittal suture. D, The parietal hole, through which a small vein runs to the superior longitu- 138 Part I. OSTEOLOGY. longitudinal sinus. E. The lambdoid suture. FF, Ossa triquetra. G, The os occipitis. H, The squamous part of the temporal bone. I, The mastoid process. K, The zygoma. L, The os malæ. M, The temporal part of the sphenoid bone. N, The superior maxillary bone and teeth. FIG. 5. The external surface of the OS FRONTIS, A, The convex part. B, Part of the tempo- ral fossa. C, The external angular process. D, The internal angular process. E, The nasal process. F, The superciliary hole. H, The orbitar plate. FIG. 6. The interior surface of the OS FRONTIS. AA, The serrated edge which assists to form the coronal suture. B, The external angular process. C, The internal angular process. D, The nasal process. E, The orbitar plate. F, The cells which correspond with those of the eth- moid bone. G, The passage from the frontal sinus. H, The opening which receives the cri- briform plate of the ethmoid bone. I, The ca- vity which lodges the fore part of the brain. K, The spine to which the falx is fixed. L, The groove which lodges the Superior longitudinal sinus. PLATE   139 Part I. OSTEOLOGY. PLATE XX. FIG. 1. A back view of the SKELETON. AA, The ossa parietalia. B, The sagittal su- ture. C, The lambdoid suture. D, The occi- pital bone. E, The squamous suture. F, The mas- toid process of the temporal bone. G, The os malæ. H, The palate plate of the superior max- illary bones. I, The maxilla inferior. K, The teeth of both jaws. L, The seven cervical ver- tebræ. M, Their spinous processes. N, Their transverse and oblique processes. O, The last of the twelve dorsal vertebræ. P, The fifth or last lumbar vertebra. Q, The transverse pro- cesses. R, The oblique processes. S, The spi- nous processes. T, The upper part of the os sacrum. U, The posterior holes which trans- mit small blood-vessels and nerves. V, The un- der part of the os sacrum which is covered by a membrane. W, The os coccygis. X, The os ilium, Y, Its spine or crest. Z, The ischi- atic niche. a, The os ischium. b, Its tubero- sity. c, Its spine. d, The os pubis, e, The fora- men hydroideum. f, The seventh or last true rib. g, The twelfth or last false rib. h, The clavicle. i, The scapula. k, its spine. l, Its acromion. m, Its cervix, n, Its superior costa. o, Its pofterior costa. p, Its inferior costa. q, The os humeri. r, The radius, s, The ulna, t, Its olecranon. u, All the bones of the carpus, exceting the os pisiforme, 140 Part I. OSTEOLOGY. pisiforme, which is seen in Plate XIX. fig. 1. v, The five bones of the metacarpus, w, The two bones of the thumb, x, The three bones of each of the fingers, y, The two sesamoid bones at the root of the left thumb, z, The os femoris. 1, The trochanter major. 2, The tro- chanter minor. 3, The linea aspera. 4, The internal condyle. 5, The external condyle. 6 6, The semilunar cartilages. 7, The tibia. 8, The malleolus internus. 9, The fibula. 10, The malleolus externus. 11, The tarsus. 12, The metatarsus. 13, The toes. FIG . 2. The external Surface of the left OS PA- RIETALE. A, The convex smooth surface. B, The pa- rietal hole. C, An arch made by the beginning of the temporal muscle. FIG. 3. The internal Surface of the same bone. A, Its superior edge, which, joined with the other, forms the sagittal suture. B, The anterior edge, which assists in the formation of the co- ronal suture. C. The inferior edge for the squa- mous suture. D, The posterior edge for the lambdoid suture. E, A depression made by the lateral sinus. FF, The prints of the arteries of the dura mater. FIG. 4. 141 Part I. OSTEOLOGY. FIG. 4. The external Surface of the Left OS TEM- PORUM. A, The squamous part. B, The mastoid pro- cess. C, The zygomatic process. D, The sty- loid process. E, The petrosal process. F, The meatus auditorius externus. G, The glenoid cavity for the articulation of the lower jaw. H, The foramen stylo-mastoideum for the portio dura of the seventh pair of nerves. I, Passages for blood-vessels into the bone. K, The fora- men mastoideum through which a vein goes to the lateral sinus. FIG. 5. The Internal surface of the Left OS TEM- PORUM. A, The squamous part; the upper edge of which assists in forming the squamous suture. B, The mastoid process. C, The styloid process. D, The pars petrosa. E, The entry of the seventh pair, or auditory nerve. F, The fossa, which lodges a part of the lateral sinus. G, The for- amen mastoideum. FIG. 6. The External Surface of the OSSEOUS CIRCLE, which terminates the meatus audi- trius externus. A, The anterior part. B, A small part of the groove in which the membrana tympani is fixed. N. B. This, with the subsequent bones of the ear, are here delineated as large as the life. FIG. 7. 142 Part I. OSTEOLOGY. FIG. 7. The Internal Surface of the OSSEOUS CIRCLE. A, The anterior part. B, The groove in which the membrana tympani is fixed. FIG. 8. The situation and connection of the small Bones of the EAR. A, The malleus. B, The incus. C, The os orbiculare. D, The stapes. FIG. 9. The MALLEUS, with its Head, Han- dle, and Small Processes. FIG. 10. The INCUS, with its Body, Superior and Inferior branches. FIG. 11. The OS ORBICULARE. FIG. 12. The STAPES, with its Head, Base, and two cura. FIG. 13. An Internal View of the LBYARINTH of the EAR. A, The hollow part of the cochlea, which forms a share of the meatus auditorius internus. B, The vestibulum. CCC, The semicircular canals. FIG. 14. An External View of the LABYRINTH. A, The semicircular canals. B, The senestra ovalis 143 Part I. OSTEOLOGY. ovalis which leads into the vestibulum. C, The senestra rotunda which opens into the cochlea. D, The different turns of the cochlea. FIG 15. The Internal Surface of the OS SPHE- NOIDES. AA, The temporal processes. BB, The pte- rygoid processes. CC, The spinous processes. DD, The anterior clinoid processes. E. The posterior clinoid process. F, The anterior pro- cess which joins the ethmoid bone. G, The sella turcica for lodging the glandula pituitaria. H, The foramen opticum. K, The foramen lacerum, L, The foramen rotundum. M, The foramen ovale. N, The foramen spinale. FIG. 16. The External Surface of the OS SPHE- NOIDES. AA, The temporal processes. BB, The pterygoid processes. CC, The spinous pro- cesses. D, The processus azygos. E, The small triangular processes which grow from the body of the bone. FF, The orifices of the sphenoid sinuses. G, The foramen lacerum. H, The foramen rotundum. I, The foramen ovale. K, The foramen pterygoideum. FIG. 17. The External Tiew of the OS ETH- MOIDES. A, The nasal lamella. BB, The grooves be- tween 144 Part I. OSTEOLOGY. tween the nasal lamella and ossa spongiosa supe- riora. CC, The ossa spongiosa superiora. DD, The sphenoidal cornua. See Fig. 16. E. FIG. 18. The Internal View of the OS ETH- MOIDES. A, The crista galli. B, The cribriform plate, with the different passages of the olfactory nerves. CC, Some of the ethmoidal cells. D, The right os planum. EE, The sphenoidal cornua. FIG. 19. The right SPHENOIDAL CORNU. FIG. 20. The left SPHENOIDAL CORNU. FIG. 21.The external Surface of the OS OCCI- PITIS. A, The upper part of the bone. B, The su- perior arched ridge. C, The inferior arched ridge. Under the arches are prints made by the muscles of the neck. DD, The two condy- loid processes which articulate the head with the spine. E, The cuneiform process. F, The foramen magnum through which the spinal mar- row passes. GG, The posterior condyloid for- amina which transmit veins into the lateral si- nuses. HH, The foramina lingualia for the passage of the nine pair of nerves. 3 FIG. 22.   145 Part I. OSTEOLOGY. FIG. 22. The Internal surface of the OS OCCI- PITIS. AA, The two sides which assist to form the lambdoid suture. B, The point of the cunei- form process, where it joins the sphenoid bone. CC, The prints made by the posterior lobes of the brain. DD, Prints made by the lobes of the cerebellum. E, The cruciform ridge for the attachment of the processes of the dura mater. F, The course of the superior longitudinal si- nuses. GG, The course of the two lateral si- nuses. H, The foramen magnum. II, The posterior condyloid foramina. PLATE XXI. FIG. 1. A Side-view of the SKELETON. AA, The ossa parietalia. B, The sagittal su- ture. C, The os occipitis, DD, The lambdoid suture. E, The squamous part of the temporal bone. F, The mastoid process. G, The mea- tus auditorius externus. H, The os frontis. I, The os malæ. K, The os maxillare supe- rius. L, The maxilla inferior. M, The teeth of both jaws. N, The seventh, or last cervical vertebra. O, The spinous processes. P, Their Transverse and oblique processes. Q, The twelfth or last dorsal vertebra. R, The fifth or last lumbar vertebra. S, The spinous processor K T, Open- 146 Part I. OSTEOLOGY. T, Openings between the vertebræ for the pas- sage of the spinal nerves. U, The under end of the os sacrum. V, The os coccygis. W, The os ilium. X, The anterior spinous processes. Y, The posterior spinous processes. Z, The ischiatic niche, a, The right os ilium. b, The ossa pubis. c, The tuberosity of the left os is- chium. d, The scapula. e, Its spine. f, The os humeri. g, The radius. h, The ulna. i, The carpus. k, The metacarpal bone of the thumb. 1, The metacarpal boneof the fingers. m, The two bones of the thumb. n, The three bones of each of the fingers. o, The os femoris. p, Its head. q, The trochanter major. r, The external condyle. s, The rotula. t, The tibia. u, The fibula. v, The malleolus externus. w, The astragalus. x, The os calcis. y, The os naviculare, z, The three ossa cuneiformia. 1, The os cuboides. 2, The five metatarsal bones. 3, The two bones of the great toe. 4, The three bones of each of the small toes. FIG. 2. A View of the Internal Surface of the Base of the SKULL. AAA, The two tables of the skull with the diplöe. BB, The orbitar plates of the frontal bone. C, The crista galli, with cribriform plate of the ethmoidal bone on each side of it, through which the first pair of nerves pass. D, The cuneiform process of the occipital bone. E. The cruciform ridge. F, The foramen mag- num 147 Part I. OSTEOLOGY. num for the passage of the spinal marrow. G, The zygoma, made by the joining of the zygo- matic processes of the os temporum. I, The pars mamillaris. K, The pars petrosa. L, The temporal process of the sphenoid bone. MM, The anterior clinoid processes. N, The poste- rior clinoid process. O, The sella turcica. P, The foramen opticum, for the passage of the op- tic nerve and occular artery of the left side. Q, The foramen lacerum, for the third, fourth, sixth, and first of the fifth pair of the nerves and ocular vein. R, The foramen rotundum, for the second of the fifth pair. T, The fora- men spinale, for the principal artery of the du- ra mater. U, The entry of the auditory nerve. V, The passage for the lateral sinus. W, The passage of the eighth pair of nerves. X, The passage of the ninth pair. FIG. 3. A View of the External surface of the Base of the SKULL. A, The two dentes incisores of the right side. B, The dens caninus. C, The two small mo- lares. D, The three large molares. E, The foramen incisivum, which gives passage to small blood-vessels and nerves. F, The palate-plates oS the ossa maxillaria and palati, joined by the longitudinal and transverse palate sutures. G, The foramen palatinum posterius, for the pala- tine vessels and nerves. H, The os maxillare K2 superius 148 Part I. OSTEOLOGY. superius of the right side. I, The os malæ. K, The zygomatic process of the temporal bone. L, The posterior extremity of the ossa spongiosa. M, The posterior extremity of the vomer, which forms the back-part of the septum nasi. N, The pterygoid process of the right side of the sphenoid bone. OO, The foramina ovalia. P P, The foramina spinalia. QQ, The passages of the internal carotid arteries. R, A hole between the point of each pars petrosa and cuneiform pro- cess of the occipital bone, which is filled up with a ligamentous substance in the recent subject. S, The passage of the left lateral sinus. T, The posterior condyloid foramen of the left side. U, The foramen mastoideum. V, The foramen magnum. W, The inferior orbitar fissure. X, The glenoid cavity, for the articulation of the lower jaw. Y, The squamous part of the tem- poral bone. Z, The mastoid process, at the inner side of which is a fossa for the posterior bel- ly of the digastric muscle. a, The styloid pro- cess. b, The meatus auditorius externus. c, The left condyle of the occipital bone. d, The perpendicular occipital spine. ee, The inferior horizontal ridge of the occipital bone. ff, The superior horizontal ridge, which is opposite to the crucial ridge where the longitudinal sinus di- vides to form the lateral sinuses. ggg, The lambdoid suture, h, The left squamous suture. i, The parietal bone. FIG. 4. 149 Part I. OSTEOLOGY. FIG. 4. The anterior surface of the OS NASI. A, The upper part, which joins the os frontis. B, The under end, which joins the cartilage of the nose. C, The inner edge, where they join each other. FIG. 5. The posterior surface of the OSSA NASI. AA, Their cavity, which forms part of the arch of the nose. BB, Their ridge or spine, which projects a little to be fixed to the fore-part of the septum narium. FIG. 6. The external surface of the OS MAXIL- LARE SUPERIUS of the left side. A, The nasal process. B, The orbitar plate. C, The unequal surface which joins the os malæ. D, The external orbitar hole. E, The opening into the nostril. F, The palate-plate. G, The maxillary tuberosity. H, part of the os palati. I, The two dentes incisores. K, The dens ca- ninus. L, The two small dentes molares. M, The three large dentes molares. FIG. 7. The internal surface of the OS MAXIL- LARE SUPERIUS and OS PALATI. A, The nasal process. BB, Eminences for the connection of the os spongiosum inferius. D, The under end of the lachrymal groove. E, The 150 Part I. OSTEOLOGY. The antrum maxillare. F, The nasal spine, be- tween which and B is the cavity of the nostril. G, The palate-plate. H, The orbitar part of the os palati. I, The nasal plate. K, The su- ture which unites the maxillary and palate bones. L, The pterygoid process of the palate bones. FIG. 8. The external surface of the right OS UNGUIS. A, The orbitar part. B, The lachrymal part. C, The ridge between them. FIG. 9. The internal surface of the right OS UNGUIS. This side of the bone has a furrow opposite to the external ridge ; all behind that is irregular, where it covers part of the ethmoidal cells. FIG. 10. The external surface of the left OS MALÆ. A, The superior orbitar process. B, The inferior orbitar process. C, The malar process. D, The zygomatic process. E, The orbitar plate. F, A passage for small vessels into or out of the orbit. FIG. 11. The internal surface of the left OS MALÆ. A, The superior orbitar process. B, The in- ferior 151 Part I. OSTEOLOGY. ferior orbitar process. C, The malar process. D, The zygomatic process. E, The internal or- bitar plate or process. FIG. 12. The external surface of the right OS SPONGIOSUM INFERIUS. A, The anterior part. B, The hook-like pro- cess for covering part of the antrum maxillare. C, A small process which covers part of the un- der end of the lachrymal groove. D, The in- ferior edge turned a little outwards. FIG. 13. The internal surface of the OS SPON- GIOSUM INFERIUS. A, The anterior extremity. B, The upper edge which joins the superior maxillary and pa- late bones. FIG. 14. The posterior and external surface of the right OS PALATI. A, The orbitar process. B, The nasal lam- ella. C, The pterygoid process. D, The pa- late process. FIG. 15. The anterior and external surface of the right OS PALATI. A, The orbitar process. B, An opening through which the lateral nasal vessels and nerves pass. 152 Part I. OSTEOLOGY. pass. C, The nasal lamella. D, The pterygoid process. E, The posterior edge of the palate process for the connection of the velum palati. F, The inner edge by which the two ossa palati are connected. FIG. 16. The right side of the VOMER. A, The upper edge which joins the nasal lam- ella of the ethmoid bone and the middle cartilage of the nose. B, The inferior edge, which is connected to the superior maxillary and palate bones. C, The superior and posterior part which receives the processus azygos of the sphe- noid bone. FIG. 17. The MAXILLA INFERIOR. A, The chin. B, The base and left side. C, The angle. D, The coronoid process. E, The condyloid process. F, The beginning of the inferior maxillary canal of the right side, for the entry of the nerve and blood-vessels. G, The termination of the left canal. H, The two dentes incisores. I, The dens caninus. K, The two small molares. L, The three large mo- lares. FIG. 18. The different classes of the TEETH. 1, 2, A sore and back view of the two ante- rior dentes incisores of the lower jaw. 3, 4, Similar 153 Part I. OSTEOLOGY. Similar teeth of the upper jaw. 5, 6, A fore and back view of the dentes canini. 7, 8, The anterior dentes molares. 9, 10, 11, The poste- rior dentes molares. 12, 13, 14, 15, 16, Un- usual appearances in the shape and size of the teeth. FIG. 19. The external surface of the OS HY- OIDES. A, The body. BB, The cornua. CC, The appendices. PLATE XXII. FIG. 1. A Posterior View of the STERNUM and CLAVICLES, with the ligament connecting the clavicles to each other. a, The posterior surface of the Sternum, bb, The broken ends of the clavicles. cccc, The tubercles near the extremity of each clavicle. d, The ligament connecting the clavicles. FIG. 2. A Fore-view of the LEFT SCAPULA, and of a half of the CLAVICLE, with their Ligaments. a, The spine of the scapula. b, The acro- mion, c, The inferior angle, d, Inferior costa. e, Cervix. f, Glenoid cavity, covered with cartilage for the arm-bone, gg, The capsular ligament of the joint. h, Coracoid process. i, The 154 Part I. OSTEOLOGY. The broken end of the clavicle. k, Its extremity joined to the acromion. 1, A ligament coming out single from the acromion to the coracoid pro- cess. m, A ligament coming out single from the acromion, and dividing into two, which are fixed to the coracoid process. FIG. 3. The Joint of the Elbow of the LEFT ARM, with the Ligaments. a, The os humeri. b, Its internal condyle. cc, The two prominent parts of its trochlea ap- pearing through the capsular ligament. d, The ulna. e, he radius, f, The part of the liga- ment including the head of the radius. FIG. 4. The Bones of the RIGHT-HAND, with the PALM in view. a, The radius. b, The ulna. c, The scap- hoid bone of the carpus. d, The os lunare. e, The os cuneiforme. f, The os pisiforme. g, Trapezium. h, Trapezoides. i, Capitatum. k, Unciforme. 1, The four metacarpal bones of the fingers. m, The first phalanx. n, The second phalanx. o, The third phalanx. p, The metacarpal bone of the thumb. q, The first joint, r, The second joint. FIG. 5. The Posterior View of the Bones of the LEFT-HAND. The explication of Fig. 4. serves for this fi- gure; 155 Part I. OSTEOLOGY. gure; the same letters pointing out the same bones, though in a different view. FIG. 6. The Upper Extremity of the TIBIA, with the Semilunar Cartilages of the Joint of the Knee, and some Ligaments. a, The strong ligament which connects the rotula to the tubercle of the tibia, bb, The parts of the extremity of the tibia, covered with cartilage, which appear within the semilunar cartilages, cc, The semilunar cartilages, d, The two parts of what is called the cross liga- ment. FIG. 7. The Posterior View of the Joint of the RIGHT-KNEE. a, The os femoris cut. b, Its internal con- dyle, c, Its external condyle, d, The back- part of the tibia. e, The superior extremity of the fibula. f, The edge of the internal semilu- nar cartilage. g, An oblique ligament. h, A larger perpendicular ligament. i, A ligament connecting the femur and fibula. FIG. 8. The Anterior View of the Joint of the RIGHT-KNEE. b, The internal condyle. c, Its external con- dyle. d, The part of the os femoris, on which the patella moves. e, A perpendicular ligament. ff, The 156 OSTEOLOGY. Part I. ff, The two parts of the crucial ligaments, gg, The edges of the two moveable semilunar carti- lages, h, The tibia, i, The strong ligament of the patella, k, The back part of it where the fat has been diffected away. 1, The exter- nal depression. m, The internal one. n, The cut tibia. FIG. 9. A View of the inferior part of the Bones of the RIGHT FOOT. a, The great knob of the os calcis. b, A pro- minence on its outside. c, The hollow for the tendons, nerves, and blood-vessels. d, The an- terior extremity of the os calcis. e, Part of the astragalus. f, Its head covered with cartilage. g, The internal prominence of the os naviculare. h, The os cuboides. i, The os cuneiforme inter- num ; k,—Medium ;—l,— Externum. m, The metatarsal bones of the four lesser toes, n, The first—o, The second—p, The third phalanx of the four lesser toes. q, The metatarsal bones of the great toe. r, Its first—s, Its second joint. FIG. 10. The Inferior Surface of the two large SESAMOID BONES, at the first Joint of the Great Toe. FIG. 11, The Superior View of the Bones of the RIGHT FOOT. a, b, as in Fig. 9. c, The superior head of the astragalus.—d, & c. as in Fig. 9. FIG. 12. 157 Part I. OSTEOLOGY. FIG. 12. The View of the SOLE of the FOOT, with its Ligaments. a, The great knob of the os calcis. b, The hollow for the tendons, nerves, and blood-vessels. c, The sheaths of the flexores pollicis and digi- torum longi opened. d, The strong cartilagin- ous ligament supporting the head of the astraga- lus. e, h, Two ligaments which unite into one, and are fixed to the metatarsal bone of the great toe. f, A ligament from the knob of the os cal- cis to the metatarsal bone of the little toe. g, A strong triangular ligament, which supports the bones of the tarsus. i, The ligaments of the joints of the five metatarsal bones. FIG. 13. a, The head of the thigh bone of a child. b, The ligamentum rotundum connect- ing it to the acetabulum. c, The capsular liga- ment of the joint with its arteries injected. d, The numerous vessels of the mucilaginous gland injected. FIG. 14. The Back-view of the Cartilages of the LARYNX, with the OS HYOIDES. a, The posterior part of the base of the os hyoides. bb, Its cornua. c, The appendix of the right side, d, A ligament sent out from the appendix of the left side, to the styloid process of the temporal bone. e, The union of the base with the left cornu. ff, The posterior sides of 158 Part II. OF THE INTEGUMENTS. of (g) the thyroid cartilage, hh, Its superior cornua. ii, Its inferior cornua. k, The cricoid cartilage. ll, The arytenoid cartilages. m, The entry into the epiglottis. o trachea. p, It FIG. 15. The Superior Concave surface of the SESAMOID BONES at the first joint of the Great Toe, with their Ligaments. a, Three sesamoid bones, b, The ligament- ous substance in which they are formed. PART II. OF the SOFT PARTS IN GENERAL; Of the COMMON INTEGUMENTS, with their Appendages; And of the MUSCLES. ANATOMICAL writers usually proceed to a description of the muscles after having finished the osteology ; but we shall deviate a little from the common method, with a view to describe every thing clearly and distinctly, and to avoid a tautology which would otherwise be unavoidable. All the parts of the body are so intimately connected with each other, that it seems     159 Part II. AND MUSCLES. seems impossible to convey a just idea of any one of them, without being in some measure oblig- ed to say something of others; and on this ac- count we wish to mention in this place the names and situation of the principal viscera of the body, that when mention is hereafter made of any one of them in the course of the work, the reader may at least know where they are placed. After this little digression, the common inte- guments, and after them the muscles will be de- scribed; we then propose to enter into an exa- mination of the several viscera and their differ- ent functions. In describing the brain, occasion will be taken to speak of the nerves and animal spirits. The circulation of the blood will follow the anatomy of the heart, and the secretions and other matters will be introduced in their proper places. The body is divided into three great cavi- ties. Of these the uppermost is formed by the bones of the cranium, and incloses the brain and cerebellum. The second is composed of the vertebræ of the back, the sternum, and true ribs, with the ad- ditional assistance of muscles, membranes, and common integuments, and is called the thorax— It contains the heart and lungs. The third, and inferior cavity, is the abdo- men. It is separated from the thorax by means of the diaphragm, and is formed by the lumbar vertebræ, the os sacrum, the ossa innominata, and 160 Part II. OF THE INTEGUMENTS. and the false ribs, to which we may add the pe- ritonæum, and a variety of muscles. This ca- vity incloses the stomach, intestines, omentum or cawl, liver, pancreas, spleen, kidneys, urinary bladder, and parts of generation. Under the division of common integuments are usually included the epidermis, or scarf-skin, the reticulum mucosum of Malpighi, the cutis or true skin, and the membrana adiposa.—The hair and nails, as well as the sebaceous glands may be considered as appendages to the skin. SECT. I. Of the SKIN. § 1. Of the SCARF-skin. The epidermis, cuticula, or scarf-skin, is a fine transparent, and insensible pellicle, destitute of nerves and blood-vessels, which invests the body, and everywhere covers the true skin. This scarf-skin, which seems to be very simple, ap- pears, when examined with a microscope, to be composed of several laminæ or scales which are increased by pressure, as we may observe in the hands and feet, where it is frequently much thickened, and becomes perfectly callous. It seems to adhere to the cutis by a number of very minute filaments, but may easily be separated from it by heat, or by maceration in water. Some anatomical writers have supposed that it is formed by a moisture exhaled from the whole surface of I the 161 Part II. AND MUSCLES. the body, which gradually hardens when it comes into contact with the air. They were per- haps induced to adopt this opinion, by observing the speedy regeneration of this part of the body when it has been by any means destroyed, it ap- pearing to be renewed on all parts of the surface at the same time; whereas other parts which have been injured, are found to direct their growth from their circumference only towards their cen- tre. But a demonstrative proof that the epider- mis is not a fluid hardened by means of the ex- ternal air, is that the fœtus in utero is found to have this covering. Lieuwenhoeck supposed its formation to be owing to the expansion of the extremities of the excretory vessels which are found every where upon the surface of the true skin. Ruysch attributed its origin to the nervous papillæ of the skin; and Heister thinks it pro- bable, that it may be owing both to the papillæ and the excretory vessels. The celebrated Mor- gagni, on the other hand, contends*, that it is nothing more than the surface of the cutis, har- dened and rendered insensible by the liquor am- nii in utero, and by the pressure of the air. This is a subject, however, on which we can advance nothing with certainty. The cuticle is pierced with an infinite number of pores or little holes, which afford a passage to the hairs, sweat, and insensible perspiration, and L likewise * Adversar. Anat. II. Animadver. 2. 162 PART II. OF THE INTEGUMENTS. likewise to warm water, mercury, and whatever else is capable of being taken in by the absorbents of the skin. The lines which we observe on the epidermis belong to the true skin. The cu- ticle adjusts itself to them, but does not form them. § 2. Of the Rete Mucosum. BETWEEN the epidermis and cutis we meet with an appearance to which Malpighi, who first described it, gave the name of rete mucosum, supposing it to be of a membranous structure, and pierced with an infinite number of pores; but the fact is, that it seems to be nothing more than a mucous substance which may be dissolved by macerating it in water, while the cuticle and cutis preserve their texture. The colour of the body is found to depend on the colour of this rete mucosum; for in negroes it is observed to be perfectly black, whilst the true skin is of the ordinary colour. The blisters which raise the skin when burnt or scalded, have been supposed by some to be owing to a rarefaction of this mucus; but they are more probably occasioned by an increased action of the vessels of the part, together with an afflux and effusion of the thinner parts of the blood. § 3. Of 163 Part II. AND MUSCLES. § 3. Of the CUTIS, or True Skin. The cutis is composed of fibres closely com- pacted together, as we may observe in leather, which is the prepared Skin of animals. These fibres form a thick net-work, which every where admits the filaments of nerves, and an infinite number of blood-vessels and lymphatics. The cutis, when the epidermis is taken off, is found to have, throughout its whole surface, in- numerable papillæ, which appear like very mi- nute granulations, and seem to be calculated to receive the impressions of the touch, being the most easily observed where the sense of feeling is the most delicate, as in the palms of the hands and on the fingers. These papillæ are supposed by many anatomi- cal writers to be continuations of the pulpy sub- stance of nerves, whose coats have terminated in the cellular texture of the skin. The great sensibility of these papillæ evidently proves them to be exceedingly nervous; but surely the nerv- ous fibrillæ of the skin are of themselves scarcely equal to the formation of these papillæ, and it seems to be more probable that they are formed like the rest of the cutis. These papillæ being described, the uses of the epidermis and the reticulum mucosum will be more easily understood; the latter serving to keep them constantly moist, while the former L2 protects 164 Part II. OF THE INTEGUMENTS. protects them from the external air, and modifies their too great sensibility. § 4. Of the GLANDS of the Skin. IN different parts of the body we meet, with- in the substance of the skin, with certain glands or follicles, which discharge a fat and oily humor that serves to lubricate and soften the skin. When the fluid they secrete has acquired a certain de- gree of thickness, it approaches to the colour and consistence of suet; and from this appearance they have derived their name sebaceous glands. They are found in the greatest number in the nose, ear, nipple, axilla, groin, scrotum, vagina, and prepuce. Besides these sebaceous glands, we read, in anatomical books, of others that are described as small spherical bodies placed in all parts of the skin, in much greater abundance than those just now mentioned, and named miliary, from their supposed resemblance to millet-seed. Steno, who first described these glands, and Malpighi, Ruysch, Verheyen, Winslow, and others, who have adopt- ed his opinions on this subject, speak of them as having excretory ducts, that open on the surface of the cuticle, and distil the sweat and matter of insensible perspiration; and yet, notwithstand- ing the positive manner in which these pretend- ed glands have been spoken of, we are now suf- ficiently 165 Part II. AND MUSCLES. ficiently convinced that their existence is altoge- ther imaginary. § 5. Of the INSENSIBLE Perspiration and SWEAT. THE matter of insensible perspiration, or in other words, the subtile vapour that is continu- ally exhaling from the surface of the body, is not secreted by any particular glands, but seems to be derived wholly from the extremities of the minute arteries that are every where dispersed through the skin. These exhaling vessels are easily demonstrated in the dead subject, by throw- ing water into the arteries; for then small drops exude from all parts of the skin, and raise up the cuticle, the pores of which are closed by death ; and in the living subject, a looking-glass placed against the skin, is soon obscured by the vapour. Bidloo fancied he had discovered ducts leading from the cutis to the cuticle, and trans- mitting this fluid; but in this he was mistaken. When the perspiration is by any means in- creased, and several drops that were insensible when separate, are united together and condens- ed by the external air, they form upon the skin small, but visible, drops called sweat (N). This particularly happens after much exercise, or whatever occasions an increased determination of fluids (N) Lieuwenhoeck asserts that one drop of sweat is formed by the conflux of fifteen drops of perspirable vapour. 166 Part II. OF THE INTEGUMENTS. fluids to the surface of the body; a greater quan- tity of perspirable matter being in such cases carried through the passages that are destined to convey it off. It has been disputed, indeed, whether the in- sensible perspiration and sweat are to be consider- ed as one and the same excretion, differing only in degree ; or whether they are two distinct ex- cretions derived from different sources. In sup- port of the latter opinion, it has been alledged, that the insensible perspiration is agreeable to na- ture, and essential to health, whereas sweat may be considered as a species of disease. But this argument proves nothing; and it seems probable, that both the insensible vapour and the sweat are exhaled in a similar manner, though they differ in quantity, and probably in their qualities; the former being more limpid, and seemingly less impregnated with salts than the latter: at any rate we may consider the skin as an emunctory through which the redundant water, and some- times the other more saline parts of the blood, are carried off. But the insensible perspiration is not confined to the skin only—a great part of what we are constantly throwing off in this way is from the lungs. The quantity of fluid exhal- ed from the human body by this insensible per spiration is very considerable. Sanctorius (O) an (O) The insensible perspiration is sometimes distinguished by the name of this physician, who was born in the territo- ries 167 Part II. AND MUSCLES. an Italian physician, who indefatigably passed a great many years in a series of statical experiments, demonstrated long ago what has been confirmed by later observations, that the quantity of vapour exhaled from the skin and from the surface of the lungs, amounts nearly to 5-8ths of the ali- ment we take in. So that if in the warm climate of Italy a person eats and drinks the quantity of eight pounds in the course of a day, five pounds of it will pass off by insensible perspiration, while three pounds only will be evacuated by stool, u- rine, saliva, &c. But in countries where the degree of cold is greater than in Italy, the quan- tity of perspired matter is less; in some of the more northern climates, it being found not to equal the discharge by urine. It is likewise ob- served to vary according to the season of the year, and according to the constitution, age, sex, dis- eases, diet, exercise, passions, &c. of different people. From ries of Venice, and was afterwards a professor in the uni- versity of Padua. After estimating the aliment he took in, and the sensible secretions and discharges, he was enabled to ascertain with great accuracy the weight or quantity of in- sensible perspiration by means of a statical chair which he contrived for this purpose; and from his experiments, which were conducted with great industry and patience, he was led to determine what kinds of solid or liquid aliment in- creased or diminished it. From these experiments he form- ed a system, which he published at Venice in 1614, in the form of aphorisms, under the title of "Ars de Medicina Statica." 168 Part II. OF THE INTEGUMLNTS. From what has been said on this subject, it will be easily conceived, that this evacuation can- not be either much increased or diminished in quantity without affecting the health. The perspirable matter and the sweat are in some measure analogous to the urine, as appears from their taste and saline nature (P). And it is worthy of observation, that when either of these secretions is increased in quantity, the other is di- minished ; so that they who perspire the least, usually pass the greatest quantity of urine, and vice versa. § 6. Of the NAILS. The nails are of a compact texture, hard and transparent like horn. Their origin is still a subject of dispute. Malpighi supposed them to be formed by a continuation of the papillæ of the skin : Ludwig, on the other hand, main- tained, that they were composed of the extre- mities of blood-vessels and nerves; both these opinions are now deservedly rejected. They seem to possess many properties in com- mon with the cuticle; like it they are neither vascular nor sensible, and when the cuticle is se- parated from the true skin by maceration or other means, the nails come away with it. They (P) Minute chrystals have been observed to shoot upon the cloaths of men who work in glass-houses. Haller Elem. Phys. 169 Part II. AND MUSCLES. They appear to be composed of different lay- ers, of unequal size, applied one over the other. Each layer seems to be formed of longitudinal fibres. In each nail we may distinguish three parts, viz. the root, the body or middle, and the extre- mity. The root is a soft, thin, and white sub- stance terminating in the form of a crescent; the epidermis adheres very strongly to this part; the body of the nail is broader, redder and thicker, and the extremity is of still greater firm- ness. The nails increase from their roots, and not from their upper extremity. Their principal use is to cover and defend the ends of the fingers and toes from external in- jury. § 7. Of the HAIR. The hairs, which from their being general- ly known do not seem to require any definition, arise from distinct capsules or bulbs seated in the cellular membrane under the skin (Q). Some of (Q) Malpighi, and after him the celebrated Ruysch, sup- posed the hairs to be continuations of nerves, being of opi- nion that they originated from the papillæ of the skin, which they considered as nervous; and as a corroborating proof of what they advanced, they argued the pain we feel in plucking them out; but later anatomists seem to have re- jected this doctrine, and consider the hairs as particular bo- dies, not arising from the papillæ (for in the parts where the papillæ; abound most there are no hairs), but from bulbs or capsule, which are peculiar to them. 170 Part II. OF THE INTEGUMENTS. of these bulbs inclose several hairs. They may be observed at the roots of the hairs which form the beard or whiskers of a cat. The hairs, like the nails, grow only from be- low by a regular propulsion from the root, where they receive nourishment. Their bulbs, when viewed with a microscope, are found to be of various shapes. In the head and scrotum they are roundish; in the eye-brows they are oval; in the other parts of the body they are nearly of a cylindrical shape. Each bulb seems to con- sist of two membranes, between which there is a certain quantity of moisture. Within the bulb the hair separates into three or four fibrillæ; the bodies of the hairs, which are the parts without the skin, vary in softness and colour according to the difference of climate, age, or tempera- ment of the body (R). Their general use in the body does not seem to be absolutely determined ; but hairs in parti- cular parts, as on the eye-brows and eye-lids, are destined for particular uses, which will be mentioned when those parts are described. § 8. Of (R) The hairs differ likewise from each other, and may not be improperly divided into two classes; one of which may in- clude the hair of the head, chin, pubes, and axillæ; and the other, the softer hairs, which are to be observed almost every where on the surface of the body. 171 PART II. AND MUSCLES. § 8. Of the CELLULAR MEMBRANE and FAT. THE CELLULAR MEMBRANE is found to in- vest the most minute fibres we are able to trace; so that by modern physiologists, it is very pro- perly considered as the universal connecting me- dium of every part of the body. It is composed of an infinite number of mi- nute cells united together, and communicating with each other. The two diseases peculiar to this membrane are proofs of such a communica- tion; for in the emphysema all its cells are filled with air, and in the anasarca they are univer- sally distended with water. Besides these proofs of communication from disease, a familiar in- stance of it may be observed amongst butchers, who usually puncture this membrane, and by inflating it with air add to the good appearance of their meat. The cells of this membrane serve as refer- voirs to the oily part of the blood or Fat, which seems to be deposited in them, either by transu- dation through the coats of the arteries, that ra- mify through these cells, or by particular ves- sels, continued from the end of arteries. These cells are not of a glandular structure, as Malpi- ghi and others after him have supposed. The fat is absorbed and carried back into the system by the lymphatics. The great waste of it in many diseases, particularly in the consumption, is a sufficient proof that such an absorption takes place. The 172 PART II. OF THE INTEGUMENTS. The fullness and size of the body are in a great measure proportioned to the quantity of fat contained in the cells of this membrane. In the living body it seems to be a fluid oil, which concretes after death. In graminivorous animals, it is found to be of a firmer consistence than in man. The fat is confined to the skin alone, being met with every where in the interstices of mus- cles, in the omentum, about the kidneys, at the basis of the heart, in the orbits, &c. The chief uses of the fat seem to be to afford moisture to all the parts with which it is con- nected; to facilitate the action of the muscles; and to add to the beauty of the body, by making it every where smooth and equal. SECT. II. Of the MUSCLES. THE MUSCLES are the organs of motion. The parts that are usually included under this name consist of distinct portions of flesh, susceptible of contraction and relaxation; the motions of which, in a natural and healthy state, are sub- ject to the will, and for this reason they are call- ed voluntary muscles. But besides these, there are other parts of the body that owe their power of contraction to their muscular fibres; thus the heart is of a muscular texture, forming what is called a hollow muscle; and the urinary blad- der, stomach, intestines, &c. are enabled to act upon their contents, merely because they are provided 173 PART II. AND MUSCLES. provided with muscular fibres. These are called involuntary muscles, because their motions are not dependent on the will. The muscles of res- piration, being in some measure influenced by the will, are said to have a mixed motion. The names by which the voluntary muscles are distinguished, are founded on their size, fi- gure, situation, use, or the arrangement of their fibres, or their origin and insertion. But be- sides these particular distinctions, there are cer- tain general ones that require to be noticed. Thus, if the fibres of a muscle are placed pa- rallel to each other in a straight direction, they form what is styled a rectilinear muscle; if the fibres cross and intersect each other, they consti- tute a compound muscle; a radiated one, if the fibres are disposed in the manner of rays; or a penniform muscle, if, like the plume of a pen, they are placed obliquely with respect to the tendon. Muscles that act in opposition to each other, are called antagonistæ; thus every extensor or muscle has a flexor for its antagonist, and vice versa. Muscles that concur in the same action are styled congeneres. The muscles being attached to the bones, the latter may be considered as levers that are mov- ed in different directions by the contraction of those organs. The end of a muscle which adheres to the most fixed part is usually called the origin, and that 174 PART II. OF THE INTEGUMENTS. that which adheres to the more moveable part the insertion, of the muscle. In every muscle we may distinguish two kinds of fibres; the one soft, of a red colour, sensi- ble, and irritable, called fleshy fibres; the other of a firmer texture, of a white glistening colour, insensible without irritability or the power of contracting, and named tendinous fibres. They are occasionally intermixed; but the fleshy fi- bres generally prevail in the belly or middle part of the muscle, and the tendinous ones in the ex- tremities. If these tendinous fibres are formed into a round slender chord, they form what is called the tendon of the muscle; on the other hand, if they are spread into a broad flat sur- face, the extremity of the muscle is styled apo- neurosis. The tendons of many muscles, especially when they are long and exposed to pressure or friction in the grooves formed for them in the bones, are surrounded by a tendinous sheath or fascia, in which we sometimes find a small mucous sac or bursa mucosa, which obviates any inconvenience from friction. Sometimes we find whole mus- cles, and even several muscles, covered by a fas- cia of the same kind, that affords origin to ma- ny of their fibres, dipping down between them, adhering to the ridges of the bones, and thus preventing them from swelling too much when in action. The most remarkable instance of such a covering is the fascia lata of the the thigh. Each 175 PART II. AND MUSCLES. Each muscle is inclosed by a thin covering of cellular membrane, which has been sometimes improperly considered as peculiar to the muscles, and described under the name of propria mem- brana musculofsa. This cellular covering dips down into the substance of the muscle, connect- ing and surrounding the most minute fibres we are able to demonstrate, and affording a support to their vessels and nerves. Lieuwhenhock fancied he had discovered, by means of his microscope, the ultimate division of a muscle, and that he could point out the sim- ple fibre, which appeared to him to be an hun- dred times less than a hair; but he was after- wards convinced how much he was mistaken on this subject, and candidly acknowledged, that what he had taken for a simple fibre was in fact a bundle of fibres. It is easy to observe several of these fasculi or bundles in a piece of beef, in which, from the coarseness of its texture, they are very evident. The red colour which so particularly distin- guishes the muscular or fleshy parts of animals, is owing to an infinite number of blood-vessels that are dispersed through their substance. When we macerate the fibres of a muscle in wa- ter, it becomes of a white colour like all other parts of the body divested their blood. The blood-vessels are accompanied by nerves, and they are both distributed in such abundance to these parts, that in endeavouring to trace the course 176 PART II. OF THE INTEGUMENTS. course of the blood-vessels in a muscle, it would appear to be formed altogether by their ramifica- tions ; and in an attempt to follow the branches of its nerves, they would be found to be equal in proportion. If a muscle is pricked or irritated, it imme- diately contracts. This is called its irritable principle; and this irritability is to be consider- ed as the characteristic of muscular fibres, and may serve to prove their existence in parts that are too minute to be examined by the eye. This power, which disposes the muscles to contract when stimulated, independent of the will, is supposed to be inherent in them; and is there- fore named vis insita. This property is not to be confounded with elasticity, which the mem- branes and other parts of the body possess in a greater or less degree in common with the mus- cles; nor with sensibility, for the heart though the most irritable, seems to be the least sensible of any of the muscular parts of the body. After a muscular fibre has contracted, it soon returns to a state of relaxation, till it is excited afresh, and then it contracts and relaxes again. We may likewise produce such a contraction, by irritating the nerve leading to a muscle, al- though the nerve itself is not affected. This principle is found to be greater in small than in large, and in young than in old, ani- mals. In the voluntary muscles these effects of con- I traction 177 PART II. AND MUSCLES. traction and relaxation of the fleshy fibres are produced in obedience to the will, by what may be called the vis nervosa, a property that is not to be confounded with the vis insita. As the existence of a vis insita different from a vis ner- vea, was the doctrine taught by Doctor Haller in his Elem. Phys. but is at present called in ques- tion by several, particularly Doctor Monro, we think it necessary to give a few objections, as sta- ted in his Observations on the Nervous System: Vis Nervea. "The chief experiment (says the Doctor) which seems to have led Dr. Hal- ler to this opinion, is the well known one, that the heart and other muscles, after being detach- ed from the brain, continue to act spontaneously, or by stimuli may be roused into action for a considerable length of time; and when it cannot be alledged, says Dr. Haller, that the nervous fluid is by the mind, or otherwise, impelled into the muscle. "That in this instance we cannot compre- hend by what power the nervous fluid or energy can be put in motion, must perhaps be granted: But has Dr. Haller given a better explanation of the manner in which his supposed vis insita be- comes active? "If it be as difficult to point out the cause of the action of the vis insita as that of the action of the vis nervea, the admission of that new power, instead of relieving, would add to our perplexity. M "We 178 PART II. OF THE INTEGUMENTS. "We Should then have admitted, that two causes of a different nature were capable of pro- ducing exactly the same effect; which is not in general agreeable to the laws of nature. "We should find other consequences arise from such an hypothesis, which tend to weaken the credibility of it. For instance, if in a sound animal the vis nervea alone produces the contrac- tion of the muscles, we will ask what purpose the vis insita serves? If both operate, are we to suppose that the vis nervea, impelled by the mind or living principle, gives the order, which the vis insita executes, and that the nerves are the internuntii; and so admit two wise agents employed in every the most simple action? But instead of speculating farther, let us learn the ef- fect of experiments, and endeavour from these to draw plain conclusions. "1. When I poured a solution of opium in water under the skin of the leg of a frog, the mus- cles, to the surface of which it was applied, were very soon deprived of the power of contraction. In like manner, when I poured this solution in- to the cavity of the heart, by opening the vena cava, the heart was almost instantly de- prived of its power of motion, whether the ex- periment was performed on it fixed in its place, or cut out of the body. "2. I opened the thorax of a living frog; and then tied or cut its aorta, so as to put a stop to the circulation of its blood. "I then 179 Part II. AND MUSCLES. "I then opened the vena cava, and poured the solution of opium into the heart; and found not only that this organ was instantly deprived of its powers of action, but that in a few mi- nutes the most distant muscles of the limbs were extremely weakened. Yet this weakness was not owing to want of circulation, for the frog could jump about for more than an hour after the heart was cut out. "In the first of these two experiments, we observed the supposed vis insita destroyed by the opium; in the latter, the vis nervea; for it is evident that the limbs were affected by the sym- pathy of the brain, and of the nervous system in general, with the nerves of the heart. "3. When the nerve of any muscle is first divided by a transverse section, and then burnt with a hot iron, or punctured with a needle, the muscle in which it terminates contracts vio- lently, exactly in the same manner as when the irritation is applied to the fibres of the muscle. But when the hot iron, or needle, is confined to the nerve, Dr. Haller himself must have ad- mitted, that this vis nervea, and not the vis in- sita, was excited. But here I would ask two questions. "First, Whether we do not as well under- stand how the vis nervea is excited when irrita- tion is applied to the muscle as when it is appli- ed to the trunk of the nerve, the impelling M2 power 180 Part II. OF THE INTEGUMENTS. power of the mind seeming to be equally want- ing in both cases? "Secondly, If it appears that irritation ap- plied to the trunk of a nerve excites the vis ner- vea, why should we doubt that it can equally well excite it when applied to the small and ve- ry sensible branches and terminations of the nerve in the muscle? "As, therefore, it appears that the supposed vis insita is destroyed or excited by the same means as the vis nervea; nay, that when, by the application of the opium to the heart of a frog, after the aorta is cut and the circulation in- terrupted, we have destroyed the vis insita, the vis nervea is so much extinguished, that the animal cannot act with the distant muscles of the limb; and that these afterward grow very tor- pid, or lose much of their supposed vis insita; it seems clearly to follow, that there is no just ground for supposing that any other principle produces the contraction of a muscle." The vis nervosa, or operation of the mind, if we may so call it, by which a muscle is brought into contraction, is not inherent in the muscle like the vis insita; neither is it perpetual, like this latter property. After long continued or violent exercise, for example, the voluntary muscles become painful, and at length incapa- ble of further action; whereas the heart and other involuntary muscles, the motions of which depend solely on the vis insita, continue through life 181 Part II. AND MUSCLES. life in a constant state of action, without any inconvenience or waste of this inherent principle; The action of the vis nervosa on the volun- tary muscles, constitutes what is called muscular motion; a subject that has given rise to a variety of hypotheses, many of them ingenious, but none of them satisfactory. Borelli and some others have undertaken to explain the cause of contraction, by supposing that every muscular fibre forms as it were a chain of very minute bladders, while the nerves which are distributed through the muscle, bring with them a supply to animal spirits, which at our will fill these bladders, and by increasing their diameter in width, shorten them, and of course the whole fibre. Borelli supposes these bladders to be of a rhomboidal shape; Bernouilli on the other hand contends that they are oval. Our countryman, Cowper, fancied he had filled them with mer- cury; the cause of this mistake was probably owing to the mercury's insinuating itself into some of the lymphatic vessels. The late inge- nius Mr. Elliot undertook to account for the phe- nomena of muscular motion on principles very different from those just now mentioned. He supposed that a deplogisticated state of the blood is requisite for muscular action, and that a com- munication of phlogiston to the blood is a neces- sary effect of such action. We know that the muscular fibre is shorten- ed, 183 Part II. OF THE INTEGUMENTS, &c. ed, and that the muscle itself swells when in action: but how these phenomena are produced, we are unable to determine. We likewise know that the nerves are essential to muscular motion; for upon dividing or making a ligature round the nerve leading to a muscle, the latter becomes incapable of motion. A ligature made on the artery of a muscle produces a similar effect; a proof this, that a regular supply of blood is also equally necessary to muscular motion. The cause of palsy is usually not to be fought for in the muscle affected, but in the nerve leading to that muscle, or in that part of the brain or spi- nal marrow from which the nerve derives its origin. Of the particular Muscles. As the enumeration and description of the the particular muscles must be dry and uninter- taining to the generality of readers, yet cannot be altogether omitted in a work of this nature, it appeared eligible to throw this part of the sub- ject into the form of a table; in which the name, origin, insertion, and principal use of each muscle, will be found described in few words, and occasionally its etymology when it is of Greek derivation or difficult to be understood. A TABLE 183 PART II. A TABLE of the MUSCLES. A TABLE of the MUSCLES arranged according to their SITUATION. [N. B. This table does not include all the muscles of the body; those belonging to the eyes, internal ear, intestinom rectum, and the male and female organs of generation, being described in other parts of the work. The reader will be pleased to observe likewise, that although all the muscles (a few only excepted) we in pairs, mention is here made of the muscles of one side.] Muscles situated Name. Origin. Insertion Use. under the integu- ments of the cra- nium- 1. Occipito frontalis. From the transvere ridge of the os oc- cipitis. Into the skin of the eye-brows. To pull the skin of the head back- wards, and to raise the eye-brows and skin of the fore- head. 2. Corrugator super- cilii. From above the join- ing of the os fron- tis, os nasi, and os maxillare. Into the inner part of the occipito- frontalis. To draw the eye- brows towards each other, and to wrinkle the forehead. -of the eye-lids- 1. Orbicularis palpe- brarum. From around the edge of the orbit. Into the nasal pro- cess of the os maxillare. To shut the eye. 2. Levator palpebræ superioris. From the bottom of the orbit, near the optic foramen. Into the cartilage of the upper eye-lid. To open the eye. 184 Part II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. —of external ear 1. Attolens auri- culam. From the tendon of the occipito fron- talis near the os temporis. Into the upper part of the ear. To raise the ear. 2. Anterior auriculæ. From near the back part of the zygoma. Into an eminance be- hind the helix. To raise this emi- nence, and to pull it forwards. 3. Retrahentes (s) au- riculæ. From the outer and back part of the root of the mastoid process. Into the convex part of the concha. To stretch the con- cha, and pull the ear backwards. MUSCLES of the car- tilages of the ear 1. Tragicus. From the outer and middle part of the concha, near the tragus. Into the upper part of the tragus. To depress the con- cha, and pull the point of the tragus a little outwards. 2. Anti-tragicus. From the root of the inner part of the helix. Into the upper part of the anti-tragus. To dilate the mouth of the concha. 3. Transversus-auri- culæ From the upper part of the concha. Into the inner part of the helix. To stretch the conc and scapha, and likewise to pull the (s) These are three small slender muscles. The inferior one is sometimes wanting. 185 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. parts it is connect- ed with towards each other. 4. Helicis major. From the upper, ante- rior, and acute part of the helix. Into the cartilage of the helix, a little above the tragus. To depress the upper part of the helix. 5. Helicis minor. From the lower and fore part of the he- lix. Into the helix, near the fissure in its cartilage. To contract the fis- sure. —of the nose, 1. Compressor (T) naris. From the outer part of the root of the ala nasi. Into the nasal pro- cess of the os max- illare, and anterior extremity of the os nasi. To straighten the nos- trils, and likewise to corrugate the skin of the nose. —of the mouth and lips, 1. Levator labii supe- rioris, alæque nasi. From the outer part of the orbitar pro- cess of the os max- illare, and from the nasal process of that bone, where it joins the os fron- tis. Into the upper lip and ala of the nose. To draw the upper lip and skin of the nose upwards and out- wards. (T) The nose is affected by fibres of the occipito frontalis, and by several muscles of the face; but this pair, the com- pressores, is the only one that is proper to it. 186 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 2. Levator anguli oris. From the os maxil- lare superlus, be- tween the orbitar foramen and the first dens molares. Into the orbicularis oris at the angle of the mouth. To draw the corner of the mouth. 3. Zygomaticus ma- jor. From the os malæ near the zygoma- tic suture. Into the angle of the mouth. To raise the angle of the mouth, and make the cheek prominent, as in laughing. 4. Zygomaticus mi- nor. Immediately above the origin of the zyg. major. Into the angle of the mouth. To raise the angle of the month oblique- ly outwards. 5. Buccinator. From the alveoli of the dentes molares in the upper and lower jaws. Into the angle of the mouth. To contract the mouth and draw the angle of it outwards and backwards. 6. Depressor labii su- perioris, alæque nasi. From the os maxill. super. immediately above the gums of the dentesincisores. Into the root of the ala nasi and upper lip. To draw the ala nasi and upper lip down- wards. 7. Depressor auguli oris. At the side of the chin from the low- er edge of the max- illa inferior. Into the angle of the mouth. To draw the corner of the mouth down- wards. 187 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 8. Depressor labii in- ferioris. From the lower and anterior part of the maxilla infe- rior. Into the under lip. To draw the under lip downwards and somewhat out- wards. 9. Levator labii infe- noris. From near the gums of the incisores and caninnus of the max- illa inferior. Into the under lip and skin of the chin. To raise the under lip and skin of the chin. 10. Orbicularis O- ris (U). To shut the mouth by constringing the lips. Muscles of the low- er jaw, 1. Temporalis. From part of the os bregmatis and os frontis; squamous part of the os tem- poris; back part of the os malæ, and the temporal pro- cess of the os sphe- noides (V) Into the coronoid pro- cess of the lower jaw. To move the lower jaw upwards. (U) This muscle is, in a great measure, if not wholly, formed by the buccinator, zygomatici, depressores, and other muscles that move the lips. Its fibres surround the mouth like a ring. (V) Some of its fibres likewise have their origin from a strong fascia that covers the muscle and adheres to the bone round the whole circumference of its origin. When we remove this covering, we find the muscle of a semicircular shape with its fibres, converging and forming a strong middle tendon. 188 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 2. Masseter (W). From the malar pro- cess of the os max- illare, and the low- er edges of the os malæ, and of the zygomatic process of the os temporis. Into the basis of the coronoid process, and that part of the jaw which supports that and the con- dyloid process. To raise and likewise to move the jaw a little forwards and backwards. 3. Pterygoideus in- ternus. From the inner surface of the outerwing of the pterygoid pro- cess of the os sphe- noides, and from the process of the os palati that helps to form the ptery- goid fossa Into the lower jaw on its inner side and near its angle. To raise the lower jaw and draw it a little to one side. 4. Pterygoideus ex- ternus. From the external ala of the pterygoid process, a small part of the adjacent os maxillare, and a ridge in the tem- Into the fore part of the condyloid pro- cess of the lower jaw, and likewise of the capsular li- gament. To move the jaw for- wards and to the opposite side (X); and at the same time to prevent the liga- ment of the joint (W) So called from its use in chewing, its derivation being from μαδιδαομαι, manduco, "to eat." (X) This happens when the muscle acts singly. When both act, the jaw is brought horizontally forwards. 189 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. poral of the os sphenoides. from being pinch- ed. MUSCLES situated at the fore part of the neck. 1. Latissimus colli (Y). From the cellular membrane covering the pectoral, del- toid, and trapezius muscles. Into the side of the chin and integu- ments of the cheek. To draw the cheeks and skin of the face downwards; and when the mouth is shut, to draw all that part of the skin to which it is connect- ed below the lower jaw upwards. 2. Mastoideus (Z). From the upper part of the sternum,and from the upper and fore part of the clavicle. Into the mastoid pro- cess, and as far back as the lamb- doidal suture. To move the head to one side, or when both muscles act, to bend it forwards. (Y) This broad and thin muscular expansion, which is situated immediately under the common integuments, is by Wins- low named muscules cutaneus. Galen gave it the name of ωλατνσμα μνωδες (Platysma-myoides); the etymology of which is from ωλατνσμος, dilatatio, and , μνς musculus, and ειδος, forma. (Z) This, on account of two origins, is by Albinus described as two distinct muscles, which he names sterno-mastoideus and cleido-mastoideus. 190 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion Use. MUSCLES situated between the trunk and the os hyoides. 1. Omo-hyoides (A) From the upper costa of the scapula near its niche; from part of a ligament that extends across this niche, and some- times by a few fi- bres, from the co- racoid process. Into the basis of the os hyoides. To draw the os hyoi- des in an oblique di- rection downwards. 2. Sterno-hyoideus. From the cartilage of the first rib, the in- ner and upper part of the sternum ,and a small part of the clavicle. Into the basis of the os hyoides. To draw the os hyoi- des downwards. 3. Hyo-thyroideus. From part of the ba- sis and horn of the os hyoides. Into a rough oblique line at the side of the thyroid carti- lage. To raise the thyroid cartilage, or depress the os hyoides. (A) This muscle does not always arise from the coracoid process, it seems to have been improperly named coraco-hyoides by Douglas and Albinus. Winslow calls it omo-hyoideus, on account of its general origin from the scapula. 191 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 4. Sterno-thyroideus From between the cartilages of the 1st and 2d ribs at the upper and inner part of the sternum. Immediately under the hyo-thyroideus. To pull the tyroid cartilage down- wards. 5. Crico-thyroideus. From the anterior part and side of the cricoid cartilage. Into the lower part and inferior horn of the thyroid car- tilage. To pull the cricoid cartilage upwards and backwards, or the thyroid for- wards and down- wards —situated be- tween the os hyoi- des and lower jaw, 1. Diagrasticus (B) From a fossa at the root of the mastoid process, and like- wise from the os hyoides. Into the lower and anterior part of the chin. To draw the lower jaw downwards. 2. Stylo-hyoideus (C) From the basis of the styloid process. Into the side and fore part of the os hy- oides near its base. To draw the os hy- oides obliquely up- wards. (B) From δις and ναςμρ (biventer), because it has two fleshy bellies with a middle tendon. This tendon passes through the stylo-hyoideus. (C) In some subjects we meet with another muscle, which from its having nearly the same origin, insertion and use as this, has been named stylo-hyoideus alter. 192 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 3. Mylo-hyoideus (D) From the inside of the lower jaw, be- tween the last dens molaris and the chin. Into the basis of the os hyoides. To move the os hyoi- des to either side, forwards or up- wards. 4. (E) Genio-hyoide- us. From the inside of the chin. Into the base of the os hyoides. To move the os hy- oides forwards or upwards. 5. Genio-glossus. From the inside of the chin. Into the tongue and basis of the os hy- oides. To move the tongue in various direc- tions. 6. Hyo-glossus (F) From the horn, basis, and appendix of the os hyoides. Into the tongue late- rally. To draw the tongue downwards and in- wards. 7. Lingualis. Laterally from the root of the tongue. Into the extremity of the tongue. To shorten the ton- gue and draw it backwards. 8. Stylo-glossus. From the styloid pro- cess,and sometimes also from a liga- ment that extends Into the side of the tongue from the root to near its tip. To move the tongue backwards and to one side. (D) So named from its arising near the dens molares (μνλοι) and its being inserted into the os hyoides. (E) From γενειον, mentum, the "chin." (F) From χερας, cornu, and γλωσσα, lingua, "the tongue." 3 193 PART II. A TABLE OF THE MUSCLES. from thence to the angle of the lower jaw. 9. Stylo-pharyngæ- us. From the basis of the styloid process. Into the side of the pharynx and poste- rior part of the thy- roid cartilage. To raise the thyroid cartilage and pha- rynx, and likewise to dilate the latter. To dilate and draw the velum obliquely downwards. 10. Circumflexus- palati. From near the bony part of the Eusta- chian tube, and from the spinous process of the os sphenoides. Into the semilunar edge of the os palati and the velum pen- dulum palati (G). 11. Levator palati. From the membra- nous part of the Eustachian tube, and the extremity of the os petrosum. Into the velum pen- dulum palati. To pull the velum backwards. MUSCLES situated a- bout the fauces, 1 Palato-pharyn- gæus. From the lower and anterior part of the cartilaginous extre- Into the upper and posterior part of the thyroid cartilage. To raise the pharynx and thyroid carti- lage, or to pull the (G) This muscle in its course forms a round tendon, which, after crossing over a kind of hook formed by the inner plate of the pterygoid process of the spenoid bone, expands into a tendinous membrane. N 194 PART II. A TABLE OF THE MUSCLES. Names. Origin. Insertion. Use. mity of the Eusti- chian tube (H); the tendinous expan- sion of the circum- flexus palati; and the velum pendu- lum palati near the basis and black part of the uvula. velum and uvula backwards and downwards. 2. Constrictor isthmi faucium. From near the basis of the tongue late- rally. Into the velum pen- dulum palati, near the basis and fore part of the uvula. To raise the tongue and draw the ve- lum towards it (I). 3. Azygos uvulæ. From the end of the suture that unites the ossa palati. Into the extremity of the uvula. To shorten the uvula, and bring it for- wards and upwards. MUSCLES at the back part of the pharynx 1. Constrictor pharyn- gis superior. From the cuneiform process of the oc- Into the middle of the pharynx. To move the pharynx upwards and for- (H) The few fibres that arise from the Eustachian tube are described as a distinct muscle by Albinus, under the name of salpingo pharyngeus. They serve to dilate the mouth of the tube. (I) This muscle, and the palato-pharyngæus, likewise serve to close the passage into the fauces, and to carry the food into the pharynx. 195 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. cipital bone; the pterygoid process of the os sphenei- des, and from each jaw near the last dens molares (K). wards, and to com- pressits upper part. 2. Constrictor pharyn- gis medius (L). From the horn and appendix of the os hyoides, and from the ligament that unites it with the thyroid cartilage. Into the middle of the processus cunei- formis of the occi- pital bone, about its middle and be- fore the great fo- ramen. To draw theos hyoi- des and pharynx upwards, and to compress the latter. 3. Constrictor pharyn- gis inferior (M). From the cricoid and thyroid cartilages. Into the middle of the pharynx. To compress part of the pharynx. —about the glottis 1. Crico-arytænoide- us lateralis. From the side of the cricoid cartilage. Into the basis of the arytænoid carti- lage laterally. To open the glottis. (K) The three orders of fibres here mentioned, with a few others |derived from the tongue, have given occasion to Douglas to describe them as four distinct muscles, under the names of cephalo-pharyngæus, mylo-pharyngæus, ptery-pharyn- gæus and glosso-pharyngæus. (L) Douglas makes two muscles of this, the hyo-pharyngæus and syndesmo-pharyngæus. (M) The crico-pharyngæus, and thyro-pharyngus of Douglas. N2 196 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 2. Crico-arytænoide- us posticus. From the cricoid car- tilage posteriorly. Into the basis of the arytænoid carti- lage posteriorly. To open the glottis. 3. Arytænoideus ob- liquis. From the basis of one of the arytænoid cartilages. Near the extremity of the other arytæ- noid cartilage. To draw the parts it is connected with towards each other. 4. Arytænoideus transversus. From one of the ary- tænoid cartilages laterally. Into the other arytæ- noid cartilage late- rally. To shut the glottis. 5. Thyreo-arytænoi- deus. From the posterior and under part of the thyroid carti- lage. Into the arytænoid cartilage. To draw the arytæ- noid cartilage for- wards. 6. Arytæno-epiglot- tideus. From the upper part of the arytænoid cartilage laterally. Into the side of the epiglottis. To move the epiglot- tis outwards. 7. Thyreo-epiglotti- deus. From the thyroid car- tilage. Into the side of the epiglottis. To pull the epiglot- tis oblique down- wards (N). MUSCLES at the fore part of the neck close to the vertebræ 1. Rectus capitis in- ternus major. From the anterior ex- tremities of the Into the fore part of the cuneiform pro- To bend the head forwards. (N) When either this or the preceding muscle acts with its fellow, the epiglottis is drawn directly downwards upon the glottis. 197 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. transverse processes of thefive lower- most cervical ver- tebræ. cess of the os occi- pitis. 2. Rectus capitis in ternus minor From the anterior and upper part of the first cervical vertebra. Near the basis of the condyloid process of the os occipitis. To assist the last de- scribed muscle. 3. Rectus capitis la- teralis. From the anterior and upper part of the transverse pro- cess of the first cer- vical vertebra Into the os occipi- tis, opposite to the stylo-mastoid fora- men. To move the head to one side. 4. Longus colli. Within the thorax, laterally from the bodies of the three uppermost dorsal vertebræ ; from the basis and fore part of the transverse processes of the first and second dorsal Into the second cer- vical vertebra an- teriorly. To pull the neck to one side (O). (O) When both muscles act, the neck is drawn directly forwards. 198 PART II. A TABLE OF THE MUSCLES. Names. Origin. Insertion. Use. vertebræ, and of the last cervical ver- tebra ; and, lastly, from the anterior extremities of the transverse processes of the 6th, 5th, 4th, and 3d cervical vertebræ. —at the fore part of the abdo- men 1. Obliquus externus. From the lower edges of the eight infe- rior ribs, near their cartilages. Into the linea alba (P), ossa pubis (Q), and spine of the ilium (R). To compress and sup- port the viscera, as- sist in evacuating the fæces and urine, (P) The linea alba is that tendinous expansion which reaches from the cartilago ensiformis to the os pubis. It is formed by the interlacement of the tendinous fibres of the oblique and transverse muscles, and on this account some anatomists have considered these as three digastric muscles. (Q) A little above the pubis the tendinous fibres of this muscle separate from each other, so as to form an opening call- ed the ring of the obliquus externus, and commonly, though improperly, the ring of the abdominal muscles, there being no such aperture either in transversalis or obliquus internus. This ring in the male subject affords a passage to the sper- matic vessels, and in the female to the round ligament of the uterus. (R) From the anterior and upper spinous process of the ilium, this muscle is stretched tendinous to the os pubis, and thus forms what is called by some Fallopius's, and by others Poupart's ligament. The blood-vessels pass under it to the thigh. 199 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. draw down the ribs and bend the trunk forwards, or ob- liquely to one side. 2. Obliquus internus. From the spinous pro- cess of the three lowermost lumbar vertebræ, the back part of the os sa- crum, the spine of the ilium, and back part of Fallopius's ligament (T). Into the cartilages of all the false ribs, linea alba (S), and fore part of pubis. To assist the obliquus externus. 2. Transversalis. From the cartilages of the seven inferi- or ribs ; the trans- verse processes of the last dorsal, and four upper lumbar vertebræ ; the in- Into the linea alba and cartilago ensi- formis. To compress the ab- dominal viscera. (S) The tendon formed by the upper par part of this, muscle in its way to the linea alba is divided into two layers. The posterior layer runs under, and the anterior one over, the rectus muscle. (T) From this part it detaches some fibres which extends downwards upon the spermatic chord, and form what is de- scribed as the cremaster muscle. 200 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. ner part of Fallo- pius's ligament and the spine of the i- lium. 4. Rectus abdominis. From the upper edge of the pubis and the symphysis pu- bis. Into the cartilages of the 5th, 6th, and 7th ribs, and the edge of the carti- lago ensiformis (U). To compress the fore part of the abdo- men, and to bend the trunk forwards. 5. Pyramidalis (V). From the anterior and upper part of the pubis. Into the linea alba and inner edge of the rectus, com- monly about two inches above the pubis. To assist the lower portion of the rect- us. MUSCLES at the fore part of the thorax 1. Pectoralis Major. From the cartilagi- Into the upper and To draw the arm for- (U) The fibres of the rectus are generally divided by three tendinous intersections. The two upper thirds of this muscle passing between the tendinous layers of the obliquus internus, are inclosed as it were in a sheath; but at its lower part we find it im- mediately contiguous to the peritonæum, the inferior portion of the tendon of the transversalis passing over tne rectus, and adhering to the anterior layer of the obliquus internus. (V) This muscle is sometimes wanting. 201 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. nous ends of the 5th and 6th ribs; anterior part of the clavicle. inner part of the os humeri (W). the sternum, and wards or oblique- ly forwards. 2. Subclavius. From the cartilage of the first rib. Into the under sur- face of the cla- vicle. To move the clavicle forward sand down- wards and to assist in raising the first rib. 3. Pectoralis minor (X). From the upper edges of the 3d 4th and 5th ribs. Into the coracoid pro- cess of the sca- pula. To move the scapula for wards and down. wards or to elevate the ribs. 4. Serratus Magnus. From the eight supe- rior ribs. Into the basis of the scapula. To bring the scapula forwards. MUSCLES that con- cur in forming the thorax, 1. Diaphragma (Y). (W) The fibres of this muscle pass towards the axilla in a folding manner, and with those of the latissimus dorsi form the arm-pit. (X) This and some other muscles derive their name serratus, from their arising by a number of tendinous or fleshy digitations, resembling the teeth of a saw (serra). (Y) For a description of the diaphragm, see Part IV. Sect. IV. 202 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 2. Levatores costa- rum. From the transverse processes of the last cervical and the eleven upper dor- sal vertebræ. Into the upper side of each rib, near its tuberosity. To move the ribs up- wards and outwards. 3. Intercostales exter- ni. From the lower edge of each upper rib. Into the superior edge of each lower rib. To elevate the ribs. 4. Intercostales in- terni (A). 5. Sterno-costales (B). From the cartilagoen- Into the cartilages of To depress the carti- (A) The origin, insertion, and use of the internal intercostals, are similar to those of the external. The reader, how- ever, will be pleased to observe, that the intercostales externi occupy the spaces between the ribs only from the spine to their cartilages ; from thence to the sternum, there being only a thin membrane, which is spread over the intercostales in- terni; and the latter, on the contrary, extend only from the sternum to the angles of each rib. The fibres of the external muscles run obliquely forwards; those of the internal obliquely backward. The difference in the direction of their fibres induced Galen to suppose that they were intended for different uses; that the external in- tercostals, for instance, serve to elevate, and the internal ones to depress the ribs. Fallopius seems to have been the first who ventured to dispute the truth of this doctrine, which has since been revived by Boyle, and more lately still by Ham- berger, whose theoretical arguments on this subject have been clearly refuted by the experiments of Haller. (B) These consist of four, and sometimes five distinct muscles on each side. Vesalius, and after him Douglas, and Al- binus, consider them as forming a single muscle, which, on account of its shape, they name triangularis. Verheyen, Winslow, and Haller, more properly describe them as so many separate muscles, which, on account of their origin and insertion, they name sterno-costales. 203 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. siformis,and lower and middle part of the sternum. the 2d, 3d, 4th, 5th, and 6th ribs. lages of the ribs. —at the back part of the neck and trunk. 1. Trapezius (C), or cucullaris. From the middle of Into the os occipitis, and the spinous pro- cesses of the two inferior cervical, and of all the dor- sal, vertebræ (D). Into the posterior half of the clavicle, part of the acro- mion, and the spine of the scapula. To move the scapula. 2. Rhomboideus (E). From the spinous pro- cesses of the three lowermost cervical, and of all the dor- sal vertebræ. Into the basis of the scapula. To move the scapula. upwards and back- wards. 3. Latissimus dorsi. From part of the spine of the os i- Into the os humeri, at the inner edge of To draw the os hu- meri downwards (C) So named by Riolanus, from Τραπεζα, on account of its quadrilateral shape. Columbus and others gave it the name of cucullaris, from its resemblance to a monk's hood. (D) The tendinous fibres of this muscle, united with those of its fellow in the nape of the neck, form what is called, the ligamentum colli. (E) This muscle consists of two distinct portions, which are described as separate muscles by Albinus, under the names of rhomboideus minor and rhombiodeus major. 204 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. lium, the spinous processess of the os sacrum and lumbar vertebræ, and of six or eight of the dor- sal vertebræ; also from the four infe- rior false ribs near their cartilages. the groove for lod- ging the long head of the biceps muscle. and backwards, and to roll it upon its axis. 4. Serratus inferior posticus. From the spinous pro- cessess of the two lowermost dorsal, and of three of the lumbar vertebræ. Into the lower edges of the three or four lowermost ribs near their cartilages. To draw the ribs out- wards, downwards, and backwards. 5. Levator scapulæ. From the transverse processess of the four uppermost vert- bræ colli. Into the upper angle of the scapula. To move the scapula forwards, and up- wards. 6. Serratus superior posticus. From the lower part of the ligamentum colli, the spinous process of the low- ermost cervical ver- tebra, and of the Into the 2d, 3d, and 4th ribs. To expand the tho- rax. 205 PART II. A TABLE OF THE MUSCLES. Name. Origin Insertion. Use. two superior dorsal vertebræ. 7. Splenius (F). From the spinous pro- cesses of the four or five uppermost vertebræ of the back, and of the lowermost cervical vertebra. Into the transverse processes of the two first cervical verte- bræ, the upper and back part of the mastoid process, and a ridge on the os occipitis. To move the head backwards. 8. Complexus (G). From the transverse processes of the four or five uppermost dorsal, and of the fix lowermost cer- vical vertebræ. Into the os occipitis. To draw the backwards. 9. Trachelo-mastoi- deus (H). From the transverse processes of the first dorsal vertebra, and Into the mastoid pro- cess. To draw the head backwards. (F) According to some writers, this muscle has gotten its name from its resemblance to the spleen; others derive it from spenium splint. (G) So named on account of its complicated structure. (H) So named from its origin from the neck (Τραιχηλοις) and its insertion into the mastoid process. 206 PART II. A TABLE OF THE MUSCLES. Name. Origin. Inserion. Use. four or five of the lowermost, cervi- cal vertebræ. 10. Rectus capitis po- sticus major. From the spinous pro- cess of the second cervical vertebra. Into the os occipitis. To extend the head and draw it back- wards. 11. Rectus capitis po- sticus minor. From the first verte- bra of the neck. Into the os occipitis. To assist the rectus major. 12. Obliquus superior capitis. From the transverse process of the first cervical vertebra. Into the os occipitis. To draw the head backwards. 13. Obliquus inferior capitis. From the spinous pro- cess of the second cervical vertebra. Into the transverse process of the first cervical vertebra. To draw the face to- wards the shoulder, and to move the first vertebra upon the second. 14. Sacro-lumbalis (I) From the back of the os sacrum, spine of the ilium, Into the lower edge of each rib. To draw the ribs downwards, move the body upon its as (I) Several thin fasciculi of fleshy fibres arise from the lower ribs, and terminate in the inner side of this muscle. Steno names them masculi ad sacro lumbalem accessorii. The sacro-lumbalis likewise sends off a fleshy slip from its upper part, which by Douglas and Albinus is described as a distinct muscle, under the name of cervicalis descendens. Morgagni has very properly considered it as a part of the sacro-lumbalis. 207 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. spinous processes and roots of the transverse processes of the vertebræ of the loins. axis, assist in erect- ing the trunk and turn the neck back- wards, or to one side. 15. Longissimus dor- si (K). The same as that of the sacro-lumbalis. Into the transverse processes of the dor- sal vertebræ. To stretch the verte- bræ of the back, and keep the trunk erect. 16. Spinalis dorsi. From the spinous pro- cesses of the upper- most lumbar and lowermost dorsal vertebræ. Into the spinous pro- cesses of the nine superior dorsal ver- tebræ To extend the verte- bræ. 17. Semi-spinalis dor- si. From the transverse processes of the 7th, 8th, 9th, and 10th vertebræ of the back. Into the spinous pro- cesses of the four uppermost dorsal, and lowermost of the cervical vertebræ. To extend the spine obliquely back- wards. (K) At the upper part of this muscle a broad thin layer of fleshy fibres is found crossing, and intimately adhering to it. This portion, which is described by Albinus, under the name of transversalis cervicis, may very properly be considered as an appendage to the longissimus dorsi. It arises from the transverse processes of the five or six superior dorsal vertebræ, and is inserted into the transverse processes of the six inferior cervical vertebræ. By means of this appendage the longissi mus dorsi may serve to move the neck to one side, or obliquely backwards. 208 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 18. Multisidus Spi- From the os facrum, næ (L). From the os sacrum, ilium, oblique and transverse processes of the lumbar ver- tebræ, transverse processes of the dor- sal, and four of the cervical vertebræ. Into the spinous pro- cesses of the lum- bar, dorsal, and six of the cervical ver- tebræ. To extend the back and draw it back- wards, or to one side. 19. Semi spinalis col- li. From the transverse processes of the five or six uppermost dorsal vertebræ. Into the spinous pro- cesses of the 2d, 3d, 4th, 5th, and 6th cervical ver- tebræ. To stretch the neck obliquely back- wards. 20. Scalenus (M). From the transverse processes of the five inferior cervical vertebræ. Into the upper and outer part of the first and second ribs. To move the neck forwards, or to one side. (L) Anatomists in general have unnecessarily multiplied the muscles of the spine. Albinus has the merit of having in- troduced greater simplicity into this part of myology. Under the name of multifidus spinæ, he has very properly included those portions of muscular flesh intermixed with tendinous fibres, situated close to the back part of the spine, and which described by Douglas under the names of transversales colli, dorsi, & lumborum. (M) The ancients gave it this name from its resemblance to an irregular triangle (σχαληνος). It consists of three fleshy portions. The anterior one affords a passage to the axillary artery, and between this and the middle portion we find the nerves going to the upper extremities. The middle is in part covered by the posterior portion, which is the longest and thinnest of the three. 209 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 21. Inter-spinalis (N). From the upper part of each of the spi- nous processes of the six inferior cer- vical vertebræ. Into the under part of each of the spinous processes of the ver- tebræ above. To draw the spinous processes towards each other. 22.Inter-transversa- les (O). From the upper part of each of the trans- verse processes of the vertebræ. Into the under part of each of the trans- verse processes of the vertebræ above. To draw the trans- verse processes to- wards each other. MUSCLES within the cavity of the abdo- men, on the ante- rior and lateral parts of the spine, 1. Psoas parvus (P). From the sides and transverse processes of the uppermost lumbar vertebra, and sometimes of Into the brim of the pelvis, at the junc- tion of the os pubis with the ilium. To bend the loins for- wards. (N) In generality of anatomical books we find these muscles divided into inter-spinalis cervicis, dorsi, and lumborum, but we do not find any such muscles either in the loins or back. (O) These muscles are to be found only in the neck and loins; which have been described, as the inner-transversales dorsi being rather small tendons than muscles. (R) This and the following pair of muscles derive their name of psoas from το α, lumbus, on account of their situation at the anterior part of the loins. 210 PART II. A TABLE of the MUSCLES. Name. Origin. Insertion. Use. the lowermost dor- sal vertebra. 2. Psoas magnus. From the bodies and transverse processes of the last dorsal, and all the lumbar vertebræ. Into the os femoris, a little below the tro- chanter minor. To bend the thigh forwards. 3. Iliacus internus. From the inner lip, hollow part, and edge of the os ilium. In common with the psoas magnus. To assist the psoas magnus. 4. Quadratus lumbo- rum (Q). From the posterior part of the spine of the ilium. Into the transverse processes of the four uppermost lumbar vertebræ, the infe- rior edge of the last rib, and the side of the lowermost dor- sal vertebra. To support the spine, or to draw it one side. 5. Coccygæus. From the posterior and inner edge of Into the lower part of the os sacrum, To draw the os coc- cygis forwards and (Q) So called from its shape, which is that of an irregular square. 211 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. the spine of ischium. and almost the whole length of the os coccygis late- rally. inwards (R). MUSCLES on the sca- pula and upper part of the os humeri, 1. Deltoides (S). From the clavicle, processus acromion, and spine of the scapula. Into the anterior and middle part of the os humeri. To raise the arm. 2. Supra-spinatus. From the basis, spine, and upper costa of the scapula. Into a large tuberosi- ty at the head of the os humeri. To raise the arm. 3. Infra-spinatus. From the base and spine of the scapu- la. Into the upper and middle part of the tuberosity. 4. Teres minor (T). From the inferior co- sta of the scapula. Into the lower part of the tuberosity. To assist the infra spi- natus. 5. Teres major. From the inferior Into the ridge at the To assist in the rota- (R) Some of the fibres of this muscle are united with those of the levator ani, so that it assists in closing the lower part of the pelvis. (S) So named from its supposed resemblance to the Greek Δ reversed. (T) This and the following pair are called teres, from their being of a long and round shape. O2 212 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. angle, and inferior costa of the scapu- la. inner side of the groove formed for the long head of the biceps. tory motion of the arm. 6. Subscapularis. From the basis, supe- rior and Inferior co- sta of the scapula. Into the upper part of a small tuberosity at the head of the os humeri. To roll the arm in- wards. 7. Coraco-brachia- lis (U) From the coracoid process on the sca- pula. Into the middle and inner side of the os humeri. To roll the arm for- wards and upwards. MUSCLES on the os humeri, 1. Biceps flexor cubi- ti. By two heads, one from the coracoid process, and the o- ther, or long head, from the upper and outer edge of the glenoid cavity of the scapula. 2. Brachialis internus From the os humeri, below, and at each Into a small tuberosity at the fore part of To assist in bending the fore-arm. (U) This muscle affords a passage to the musculo-cutaneous nerve. 213 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. side of the tendon the coronoid pro- cess of the ulna. 3. Triceps extensor cubiti. By three heads: the first, from the infe- rior costa of the scapula; the second from the upper and outer part of the os humeri; and the third, from the back part of that bone. Into the upper and outer part of the olecranon. To extend the fore- arm. MUSCLES on the fore- arm, 1. Supinator longus. From the outer ridge and anterior surface of the os humeri, a little above its out- er condyle. Into the radios near its styloid process. To assist in turning the palm of the hand upwards. 2. Extensor carpi ra- dialis longus. Immediately below the origin of the supinator longus. Into the upper part of the metacarpal bone of the fore-finger. To extend the wrist. 3. Extensor capri ra- dialis brevis. From the outer and lower part of the Into the upper part of the metacarpal To assist the extensor longus. 214 PART II. A TABLE OF THE MUSCLES. outer condoyle of the os humeri, and the upper part of the radius. bone of the middle finger. 4. Extensor digito- run communis. From the outer con- dyle of the os hu- meri. Into the back part of all the bones of the fore finger. To extend the fingers. 5. Extensor minimi digiti. From the outer con- dyle of the os hu- meri. Into the bones of the little finger. To extend the little finger. 6. Extensor carpi ul- naris. From the outer con- dyle of the os hu- meri. Into the metacarpal bone of the little finer. To assist in extending the wrist. 7. Anconæus (V). From the outer con- dyle of the os hu- meri. Into the outer edge of the ulna. To extend the fore- arm. 8. Flexor carpi ulna- ris. From the inner con- dyle of the os hu- meri, and anterior edge of the olecra- non (W). Into the os pisiforme. To assist in bending the hand. 9. Palmaris longus. From the inner con- Into the internal an- To bend the hand. (V) So called αΓχων, cubitus. (W) Between the two origins of this muscle we find the ulnor nerve going to the fore-arm. 215 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. dyle of the os hu- meri. nular ligament, and aponeurosis palma- ris (X). 10. Flexor carpi ra- dialis. From the inner con- dyle of the os hu- meri. Into the metacarpal bone of the fore finger. To bend the hand. 11. Pronator radii teres. From the outer con- dyle of the os hu- meri, and coronoid process of the ulna. Into the anterior and convex edge of the radius near its middle. To roll the hand in- wards. 12. Flexor sublimis perforatus (Y). From the inner con- dyle of the os hu- meri, inner edge of the coronoid pro- cess of the ulna, and upper and an- terior part of the radius. Into the second bone of each finger. To bend the second joint of the finger. (X) The aponeurosis palmaris is a tendinous membrane that extends over the palm of the hand. Some anatomists have supposed it to be a production of the tendon of this muscle, but without sufficient grounds; for in some subjects we find. the palmaris longus inserted wholly into the annular ligament, so as to be perfectly distinct from its aponeurosis; and it now and then happens, that no palmaris longus is to be found, whereas this expansion is never deficient. (Y) This muscle is named perforatus, on account of the four tendons in which it terminates, being perforated by those of another muscle, the perforans. 216 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 13. Supinator radii brevis. From the outer con- dyle of the os hu- meri, and posterior surface and outer edge of the ulna. Into the anterior, in- ner, and upper part of the radius. To roll the radius outwards. 14. Abductor polli- cis longus. From the middle and back part of the ulna, interosseous ligament, and ra- dius. By two tendons into the os trapezium, and first bone of the thumb. To stretch the first bone of the thumb outwards. 15. Extensor minor pollicis. From the back part of the ulna, and in- terosseous ligament and radius. Into the convex part of the second bone of the thumb. To extend the second bone of the thumb obliquely outwards. 16. Extensor major pollicis. From the back of the ulna and interosse- ous ligament. Into the third and last bone of the thumb. To stretch the thumb obliquely back- wards. 17. Indicator. From the middle of the ulna. Into the metacarpal bone of the fore- finger. To extend the fore- finger. 18. Flexor profundus perforans. From the upper and fore part of the ulna, and interos- seous ligament. Into the fore part of the last bone of each of the fingers. To bend the last joint of the fingers. 217 Part II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 19. Flexor longus pollicis. From the upper and fore part of the radius. Into the last joint of the thumb. To bend the last joint of the thumb. 20. Pronator radii quadratus. From the inner and lower part of the ulna. Into the radius, op- posite to its origin. To roll the radius in- wards and of course to assist in the pro- nation of the hand. MUSCLES on the hand, I. Lumbricales (Z). From the tendons of the perforans. Into the tendons of the extensor digi- torum communis. To bend the first, and to extend the two last joints of the fingers (A). 2. Abductor brevis pollicis. From the fore part of the internal an- nular ligament, os scaphoides, and one of the tendons of the abductor lon- gus pollicis. Into the outer side of the 2d bone of the thumb, near its root. To move the thumb from the fingers. 3. Opponens pollicis. From the inner and anterior part of the internal annular li- Into the first bone of the thumb. To move the thumb inwards and to turn it upon its axis. (Z) So named from their being shaped somewhat like the lumbricus or earth-worm. (A) Fallopius was the first who remarked the two opposite uses of this muscle. Their extending power is owing to their connection with the extensor communis. 218 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. gament, and from the os scaphoides. 4. Flexor brevis pol- licis. From the os trapezoi- des, internal annu- lar ligament, os magnum, and os unciforme. Into the ossa sesamoi- dea and second bone of the thumb. To bend the second joint of the thumb. 5. Adductor pollicis. From the metacarpal bone of the middle finger. Into the basis of the second bone of the thumb. To move the thumb towards the fingers 6. Adductor indices. From the inner side of the first bone of the thumb, and from the os trape- zium. Into the first bone of the fore finger po- steriorly. To move the fore-fin - ger towards the thumb. 7. Palmaris brevis. From the internal an- nular ligament,and aponeurosis palma- ris. Into the os pisiforme, and the skin cover- ing the abductor minimi digiti. To contract the palm of the hand. 8. Adductor minimi digiti. From the internal an- nular ligament and os pisiforme. Into the side of the first bone of the little finger. To draw the little finger from the rest. 9. Flexor parvus mi- nimi digiti. From the os uncifor- me and internal an- nular ligament. Into the first bone of the little finger. To bend the little fin- ger. 219 PART II. A TABLE OF THE MUSCLES. Name. Use. Insertion. Use. 10. Adductor meta- carpi minimi digiti. From the os uncifor- me and internal an- nular ligament. Into the metacarpal bone of the little finger. To move that bone towards the rest. 11. Interossei interni. Situated between the metacarpal bones. Into the roots of the fingers. To extend the fingers and move them to- wards the thumb (B). 12. Interossei externi. Situated between the matacarpal bones on the back of the hand. Into the roots of the fingers. To extend the fin- gers; but the first draws the middle finger inwards, the second draws it out- wards, and the third draws the ring fin- ger inwards. MUSCLES at the back part of the pelvis, and upper part of the thigh, 1. Glutæus (C) max- imus. From the spine of the illium, posterior sa- cro ischiatic liga- ments, os sacrum, and os coccygis. Into the upper part of the linea aspera of the os femoris. To extend the thigh and draw it out- wards. (B) The third interosseus internus (for there are four of the externi and three of the interni) differs from the rest in drawing the middle finger from the thumb. (C) From γλδτος, nates. 220 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 2. Glutæus medius. From the spine and superior surface of the ilium. Into the outer and back part of the great trochanter of the os femoris. To draw the thigh outwards and a lit- tle backwards, and when it is bended, to roll it. 3. Glutæus minimus. From the outer sur- face of the ilium and the border of its great niche. Into the upper and anterior part of the great trochanter. To assist the former. 4. Pyriformis (D). From the anterior part of the os sa- crum. Into a cavity at the root of the trochan- ter major. To roll the thigh out- wards. 5. Gemini (E). By two portions, one from the outer sur- face of the spine of the ischium; the other from the tuberosity of the ischium and poste- rior sacro-ischiatic ligament. Into the same cavity as the pyriformis. To roll the thigh out- wards, and likewise to confine the ten- don of the obtura- tor internus, when the latter is in ac- tion. (D) So named from its pear-like shape. (E) The two portions of this muscle having been described as two distinct muscles by some anatomists, have occasioned it to be named gemini. The tendon of the obturator internus runs between these two portions. 221 PART II. A TABLE OF THE MUSCLES. Names. Origin. Insertion. Use. 6. Obturator internus. From the superior half of the inner border of the fo- ramen thyroideum. Into the same cavity with the former. To roll the thigh out- wards. 7- Quadratus (F) fe- moris. From the tuberosity of the ischium. Into a ridge between the trochanter ma- jor and trochanter minor. To move the thigh outwards. MUSCLES on the thigh (G), 1. Biceps flexor cru- ris. By two heads; one from tuberosi- ty of the ischium, the other from the linea aspera near the insertion of the glutæus maximus. Into the upper and back part of the fi- bula (H). To bend the leg. 2. Semi-tendinosus. From the tuberosity of the ischium. Into the upper and inner part of the tibia. To bend and draw the leg inwards. (F) This muscle is not of the square shape its name would seem to indicate. (G) The muscles of the leg and thigh are covered by a broad tendinous membrane called fascia lata, that surrounds them in the manner of a sheath. It is sent off from the tendons of the glutæi and other muscles, and dipping down be- tween the muscles it covers, adheres to the linea aspera, and spreading over the joint of the knee, gradually disappears on the leg. It is thickest on the inside of the thigh. (H) The tendon of this muscle forms the outer-ham-string. 222 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 3. Semi-membrano- sus (I). From the tuberosity of the ischium. Into the upper and back part of the head of the tibia. To bend the leg. 4. Tensor vaginæ fe- moris. From the superior and anterior spinous pro- cess of the ilium. Into the inner side of the fascia lata, which covers the outside of the thigh. To stretch the fascia. 5. Sartorius. From the superior and anterior spinous process of the ili- um. Into the upper and in- ner part of the ti- bia. To bend the leg in- wards (K). 6. Rectus. By two tendons; one from the anterior and inferior spi- nous process of the ilium ; the other from the posterior edge of the coty- loid cavity. Into the upper and fore-part of the pa- tella. To extend the leg. 7. Gracilis. From the fore-part of the ischium and pubis. Into the upper and inner part of the tibia. To bend the leg. (I) So named on account of its origin, which is by a broad flat tendon three inches long. (K) Spigelius was the first who gave this the name sartorius, or the taylor's muscle, its use in crossing the legs. 223 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 8. Vastusexternus (L) From the anterior and lower part of the great trochanter, and the outer edge of the linea as- pera. To the upper and outer part of patella. To extend the leg. 9. Vastus internus. From the inner edge of the linea aspera, beginning between the fore-part of the os femoris and the root of the lesser trochanter. Into the upper and inner part of the patella. To extend the leg. 10. Cruræus (M) From the outer and anterior part of the lesser trochan- ter. Into the upper part of the patella. To extend the leg. 11. Pectinalis. From the anterior edge of the os pu- bis, or pectinis, as it is sometimes called. Into the upper and fore part of the li- nea aspera. To draw the thigh inwards, upwards, and to roll it a little outwards. (L) The vastus externus, vastus internus, and cruræus, are so intimately connected with each other, that some anato- mists have been induced to consider them as a triceps, or single muscle with three heads. (M) Under the cruræus we sometimes meet with two small muscles, to which Albinus has given the name of sub-cruræi. They terminate on each side of the patella, and prevent the capsular ligament from being pinched. When they are want- ing, which is very often the case, some of the fibres of the cruræus are found adhering to the capsula. 224 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 12. Adductor longus Femoris (N). From the upper and fore part of the os pubis. Near the middle and back part of the linea aspera. To draw the thigh inwards, upwards, and to roll it a little outwards. 13. Adductor brevis femoris. From the fore part of the ramus of the os pubis. Into the inner and upper part of the linea aspera. To draw the thigh inwards, upwards, and to roll it a little outwards. 14. Adductor mag- nus femoris. From the lower and fore part of the ra- mus of the os pu- bis. Into the whole length of the li- nea aspera. To draw the thigh inwards, upwards, and to roll it a little outwards. 15. Obturator exter- nus. From part of the ob- turator ligament, and the inner half of the circumfe- rence of the fora- men thyroideum. Into the os femoris near the root of the great trochan- ter. To move the thigh outwards in an ob- lique direction, and likewise to bend and draw it in- wards. MUSCLES on the leg, 1. Gastrocnemius (O) externus. By two heads; one from the inner con- dyle,the other from the outer condyle of the os femoris. By a great round ten- don, common to this and the follow- ing muscle. To extend the foot (N) This and the two following muscles have been usually, but improperly, considered as forming a single muscle with three heads, and on that account named triceps femoris. (O) sαsροχγημια sura, the calf of the leg." 3 225 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 2. Gastrocnemius (P) internus. By two heads; one from the back part of the head of the fibula, the other from the upper and back part of the By a large tendon (the tendo achillis) common to this and the former muscle, into the lower and back part of the os calcis. To extend the foot. 3. Plantaris (Q) From the upper and posterior part of the outer condyle of the os femoris. Into the inside of the back part of the os calcis. To assist in extending the foot. 4. Popliteus (R) From the outer con- dyle of the thigh. Into the upper and inner part of the tibia. To assist in bending the leg and rolling it inwards. 5. Flexor longus digi- torum pedis (S) From the upper and inner part of the tibia. By four tendons, which, after passing through the perfo- rations in those of the flexor digito- To bend the last joint of the toe. (P) This muscle is by some anatomists named soleus, on account of its being shaped like the sole-fish. (Q) This muscle has gotten the name of plantaris, from its being supposed to furnish the aponeurosis that covers the sole of the foot; but it does not in the least contribute to the formation of that tendinous, expansion. (R) So called on account of its situation at the ham (poples). (S) This muscle, about the middle of the foot, unites with a fleshy mass, which, from its having first been described by Sylvius, is usually called massa carnea JACOBI SYLVII. P 226 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. rum brevis, are in- serted into the last bone of all the toes except the great toe. 6. Flexor longus pol- licis pedis. From the back part, and a little below the head of the fi- bula. Into the last bone of the great toe. To bend the great toe. 7. Tibialis posticus. From the back part and outer edge of the tibia, and like- wise from the in- terosseous ligament and adjacent part of the fibula. 8. Peroneus longus. From the outer side of the head of the tibia, and also from the upper, anterior and outer part of the perone or fibu- la, to which it ad- heres for a consi- derable way down. Into the metatarsal bone of the great toe. To move the foot out- wards. 227 PART II. A TABLE OF THE MUSCLES. Name. Origin. Inseation. Use. 9. Peroneus brevis. From the outer and fore-part of the fi- bula. Into the metatarsal bone of the little toe. To assist the last de- scribed muscle. 10. Extensor longus digitorum pedis. From the upper, out- er, and fore part of the tibia, inter- osseous ligament, and inner edge of the fibula. By four tendons into the first joint of the smaller toes. To extend the toes. 11. Peroneus tertius. From the fore-part bone of the little toe. Into the metatarsal of the lower half of the fibula, and from the interos- seous ligament. To bend the foot. 12. Tibialis amicus. From the upper and fore part of the ti- bia. Into the os cunei- forme internum. To bend the foot. 13. Extensor proprius pollicis pedis. From the upper and fore part of the ti- bia. Into the convex sur- face of the bones of the great toe. To extend the great toe. MUSCLES on the foot, 1. Extensor brevis di- gitorum pedis. From the upper and anterior part of the os calcis. By four tendons; one of which joins the tendon of the ex- ternus longus polli- To extend the toes. P2 228 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. cis, and the other three the tendons of the extensor di- gitorum longus. 2. Flexor brevis digi- torum pedis. From the lower part of the os calcis. By four tendons, which, after af- fording a passage to those of the flex- or longus, are in- serted into the se- cond phalanx of each of the small toes. To bend the second joint of the toes. 3. Abductor pollicis pedis. From the inner and lower part of the os calcis. Into the first joint of the great toe. To move the great toe from the other toes. 4. Abductor minimi digiti. From the outer tuber- cle of the os calcis, the root of the me- tatarsal bone of the little toe, and also from the aponeuro- sis plantaris. Into the outer side of the first joint of the little toe. To draw the little toe outwards. 229 PART II. A TABLE OF THE MUSCLES. Name. Origin. Insertion. Use. 5. Lumbricalis pedis. From the tendons of the flexor longus digitorum pedis. Into the tendinous expansion at the upper part of the toes. To draw the toes in- wards. 6. Flexor brevis pol- licis pedis. From the inferior and anterior part of the os calcis, and also from the inferior part of the os cu- neiforme externum By two tendons into the first joint of the great toe. To bend the first joint of the great toe. 7. Adductor pollicis pedis. From near the roots of the metatarsal bones of the 2d, 3d, and 4th toes. Into the outer os se- samoideum, or first joint of the great toe. To draw the great toe nearer to the rest, and also to bend it. 8. Transversales pe- dis. From the outer and under part of the interior end of the metatarsal bone of the little toe. Into the inner os se- samoideum, and an- terior end of the metatarsal bone of the great toe. To contract the foot. 9. Flexor brevis mi- nimi digiti pedis. From the basis of the metatarsal bones of the little toe. Into the first jont of the little toe. To bend the little toe. 230 PART II. A TABLE OF THE MUSCLES. 10. Interossei pedis interni ( T ). Situated between the metatarsal bones. —exter- ni (U). (T) The interossei interni are three in number ; their use is to draw the smaller toes towards the great toe. (U) the interossei externi are four in number ; the first serves to move the fore-toe towards the great toe: and the rest move the toes outwards. All the interossei assist in extending the toes. 231 Part II. OF THE MUSCLES. EXPLANATION of PLATES XXIII and XXIV. PLATE XXIII. FIG. 1. The MUSCLES immediately under the common teguments on the anterior part of the body are represented on the right side; and on the left side the MUSCLES are seen which come in view when the exterior ones are taken away. A, The frontal muscle. B, The tendinous aponeurosis which joins it to the occipital; hence both named occipito-frontalis. C, Attolens au- rem. D, The ear. E, Anterior auris. FF, Orbicularis palpebrarum. G. Levator labii su- perioris alæque nasi. H, Levator anguli oris. I, Zygomaticus minor. K, Zygomaticus ma- jor. L, Massater. M, Orbicularis oris. N, Depressor labii inferioris. O, Depressor anguli oris. P, Buccinator. QQ, Platysma myoides. R R, Sterno-cleido-mastoidæus. S, Part of the trapezius. T, Part of the scaleni. SUPERIOR EXTREMITY.—U, Deltoides. V, Pectoralis major. W, Part of the latissimus dor- si. X X, Biceps flexor cubiti. YY, Part of the brachialis externus. ZZ, The beginning of the tendinous aponeurosis (from the biceps), which is spread over the muscles of the fore-arm. a a, Its strong tendon inserted into the tubercle of the radius, b b, Part of the brachialis inter- nus. c, Pronator radii teres, d. Flexor carpi radialis. 232 Part II. OF THE MUSCLES. radialis. e, Part of the flexor carpi ulnaris. f, Palmaris longus. g, Aponeurosis palmaris. 3, Palmaris brevis. 1, Ligamentum carpi annulare. 2 2, Abductor minimi digiti. h, Supinator radii longus. i, The tendons of the thumb. k, Ab- ductor pollicis. l, Flexor pollicis longus. mm, The tendons of the flexor sublimis perforatus, profundus perforans, and lumabricales.—The sheaths are entire in the right hand,—in the left cut open to show the tendons of the flexor pro- fundus perforating the sublimis. MUSCLES not referred to—in the left superior extremity.—n, Pectoralis minor, seu serratus an- ticus minor. o, The two heads of (xx) the biceps. p, Coraco-brachialis. qq, The long head of the triceps extensor cubiti. rr, Teres major. ff, Subscapularis. tt, Extenfores ra- diales, u, Supinator brevis. v, The cut extre- mity of the pronator teres. w, Flexor sublimis perforatus. x, Part of the flexor profundus. y, Flexor pollicis longus. z, Part of the flexor pol- licis brevis. 4, Abductor minimi digiti. 5, The four lumbricales. TRUNK.—6, Serrated extremities of the ser- ratus anticus major, 77, Obliquus externus ab- dominis. 88, The linea alba. 9, The umbi- licus. 10, Pyramidalis. 1111, The sperma- tic cord. On the left side it is covered by the cremaster. 1212, Rectus abdominis. 13, Qbliquus internus. 1414, &c. Intercostal muscles. INFERIOR 233 PART II. OF THE MUSCLES. INFERIOR EXTREMITIES.—aa, The gra- cilis. b b, Parts of the triceps, cc, Pectialis. dd, Psoas magnus. ee. Iliacus internus. f, Part of the gluteus medius. g, Part of the glu- teus minimus. h, Cut extremity of the rectus cruris. ii, Vastus externus. k, Tendon of the rectus cruris. ll, Vastus internus. * Sartorius muscle. ** Fleshy origin of the tensor vaginæ fæmoris or membranosus. Its tendinous apo- neurosis covers (i) the vastus externus in the right side. mm, Patella. nn, Ligament or tendon from it to the tibia. o, Rectus cruris. p, Cruræus. qq, The tibia, rr, Part of the Gemellus or gastrocnemius externus.* sss, Part of the soleus or gastrocnemius internus. t, Tibi- alis anticus. u, Tibialis posticus. vv, Peronæi muscles. ww, Extensor longus digitorum pe- dis, xx, Extensor longus pollicis pedis. y, Ab- ductor pollicis pedis. FIG 2. The MUSCLES, GLANDS, &c. of the Left Side of the face and neck, after the com- mon Teguments and Platysma myoides have been taken off. a, The frontal muscle. b, Temporalis and temporal artery. c, Orbicularis palpebrarum. d, Levator labii superioris alæque nasi. e, Le- vator anguli oris. f, Zygomaticus. g, Depressor labii inferioris. h, Depressor anguli oris. i, Buc- cinator, k, Masseter. ll, Parotid gland. m, Its duct. 234 Part II. OF THE MUSCLES. duct. n, Sterno-cleido-mastoidæus. o, Part of the trapezius. p, Sterno-hyoidæus. q, Sterno thyroidæus. r, Omo-hyoidæus. s, Levator scapulæ. tt , Scaleni. u, Part of the splenius. FIG. 3. The MUSCLES of the Face and Neck in view after the exterior ones are taken away. aa, Corrugator supercilii. b, Temporalis. c, Tendon of the levator palpebræ superioris. d, Tendon of the orbicularis palpebrarum. e, Mas- seter. f, Buccinator. g, Levator anguli oris. h, Depressor labii superioris alæque nasi. i, Or- bicularis oris. k, Depressor anguli oris. l, Mus- cles of the os hyoides. m, Sterno-cleido-masto- idæus. FIG. 4. Some of the MUSCLES of the Os Hy- oides and Submaxillary Gland. a, Part of the masseter muscle. b, Posterior head of the digrastic. c, Its anterior head. dd, Sterno-hyoidæus. e, Omo-hyoidæus. f, Stylo- hyoidæus. g, Submaxillary gland in situ. FIG. 5. The submaxillary Gland and Duct. a, Musculus mylo-hyoidæus. b. Hyo-glos- sus. c, Submaxillary gland extra situ. d, Its duct. PLATE 235 Part II. OF THE MUSCLES. PLATE XXIV. FIG. 1 The muscles immediately under the common teguments on the posterior part of the body, are represented in the right side; and on the left side the MUSCLES are seen which come in view when the exterior ones are taken away. HEAD.—AA, Occipito-frontalis. B, At- tollens aurem. C, Part of the orbicularis pal- pebrarum. D, Masseter. E, Pterygoidæus in- ternus. TRUNK.—Right side. FFF, Trapezius seu cucullaris. GGGG, Latissimus dorsi. H, Part of the obliquus externus abdominis. TRUNK.—Left side. I, Splenius. K, Part of the complexus. L, Levator scapulæ. M, Rhomboides. NN, Serratus posticus inferior. O, Part of the longissimus dorsi. P, Part of the sacro-lumbalis. Q, Part of the semi-spinalis dorsi. R, Part of the serratus anticus major. S, Part of the obliquus internus abdominis. SUPERIOR EXTREMITY.—Right side. T, Deltoides. U, Triceps extensor cubiti. V, Su- pinator longus. WW, Extensores carpi radia- lis longior and brevior. XX, Extensor carpi ulnaris. YY, Extensor digitorum communis. Z, Abductor indicis. 123, Extensores pol- licis. SUPERIOR EXTREMITY.—Left side. a, Su- pra spinatus. b, Infra-spinatus. c, Teres mi- nor. 236 PART II. OF THE MUSCLES. nor. d, Teres major. e, Triceps extensor cu- biti. ff, Extensores carpi radiales. g, Supi- nator brevis. h, Indicator. 123, Extensores pollicis. i, Abductor minimi digiti. k, Inter- ossei. INFERIOR EXTREMITY.—Right side. l, Glutæus maximus. m, Part of the Glutæus medius. n, Tensor vaginæ femoris. o, Graci- lis. p p, Abductor femoris magnus. q, Part of the vastus internus. r, Semimembranosus. s, Semitendinosus. t, Long head of the biceps flexor cruris. u u, Gastrocnemius externus seu gemellus. v, Tendo Achiilis. w, Solens seu gastrocnemius interims. xx, Peronæus longus and brevis. y, Tendons of the flexor longus digi- torum pedis;—and under them * flexor brevis di- gitorum pedis. z, Abductor minimi digiti pedis. INFERIOR EXTREMITY. Left side. m, n, o, p, q, r, s, t, v, ww, xx, y, z, Point the same parts as in the right side. a, Pyriformis. bb, Gemini. cc, Obturator internus. d, Quadra- tus femoris. e, Coccygaæus. f, The short head of the biceps flexor cruris. gg, Plantaris. b, Poplitæus. i, Flexor longus pollicis pedis. FIG. 2. The Palm of the Left Hand after the common Teguments are removed, to show the MUSCLES of the Fingers. a, Tendon of the flexor carpi radialis. b, Tendon of the flexor carpi ulnaris. c, Tendons of the flexor sublimis perforatus, profundus per- forans     237 PART II. OF THE MUSCLES. forans and lumbricales. d, Abductor pollicis. e e, Flexor pollicis longus. f, Flexor pollicis brevis. g, Palmaris brevis. h, Abductor mi- nimi digiti. i, Ligamentum carpi annulare, k, A probe put under the tendons of the flexor di- gitorum sublimis; which are perforated by l, the flexor digitorum profundus, mmmm, Lum- bricales. n, Abductor pollicis. FIG. 3. A Fore-view of the foot and Tendons of the Flexores Digitorum. a, Cut extremity of the tendo Achillis. b, Upper part of the astragalus. c, Os calcis. d, Tendon of the tibialis anticus. e, Tendon of the extensor pollicis longus. f, Tendon of the peronæus brevis. g, Tendons of the flexor di- gitorum longus, with the nonus Vesalii. h h, The whole of the flexor digitorum brevis. FIG. 4. Muscles of the Anus. a a, An out line of the buttocks, and upper part of the thighs. b, The testes contained in the scrotum. cc, Sphincter ani. d, Anus, e, Levator ani. ff, Erector penis. gg, Accele- rator urinæ. h, Corpus cavernosum urethræ. FIG. 5. Muscles of the Penis. aa, b, d, ee, ff, h, point the same as in fig. 4. c, Sphincter ani. gg, Transversalis penis. PART 238 PART III. OF THE ABDOMEN. PART III. OF THE ABDOMEN, OR LOWER BELLY. THE abdomen, or lower belly, extends from the lower extremity of the sternum, or the hollow, usually called the pit of the stomach, and more properly scrobiculus cordis, to the low- er part of the trunk. It is distinguished into three divisions called regions; of these the upper one, which is called the epigastric region, begins immediately under the sternum, and extends to within two fingers breadth of the navel, where the middle or um- bilical region begins, and reaches to the same dis- tance below the navel. The third, which is called the hypogastric, includes the rest of the abdomen, as far as the os pubis. Each of these regions is subdivided into three others; two of which compose the sides, and the other the middle part of each region. The middle part of the upper region is called epigastrium, and its two sides hypochondria. The middle part of the next region is the umbilical region, properly so called, and its two sides are the flanks, or iliac regions. Lastly, the middle part of the lower region retains the name of hy- pogastrium, and its sides are called inguina or groins. The back part of the abdomen bears the name of lumbar region. These 239 PART III. OF THE ABDOMEN. These are the divisions of the lower belly, which are necessary to be held in remembrance, as they frequently occur in surgical and anato- mical writing. We will now proceed to exa- mine the contents of the abdomen; and after having pointed out the names and arrangement of the several viscera contained in it, describe each of them separately. After having removed the skin, adipose mem- brane, and abdominal muscles, we discover the peritonæum or membrane that envelopes all the viscera of the lower belly. This being opened, the first part that presents itself is the omentum or cawl, floating on the surface of the intestines, which are likewise seen every where loose and moist, and making a great number of circum- volutions through the whole cavity of the abdo- men. The stomach is placed in the epigastrium, and under the stomach is the pancreas. The liver fills the right hypochondrium, and the spleen is situated in the left. The kidneys are seen about the middle of the lumbar region, and the urinary bladder and parts of generation are are seated in the lower division of the belly. SECT. 1. Of the Peritonæum. The peritonæum is a strong simple mem- brane, by which all the viscera of the abdomen are surrounded, and in some measure supported. Many anatomical writers, particularly Winslow, have 240 PART III. OF THE ABDOMEN. have described it as being composed of two dis- tinct membranous laminæ; but their description seems to be erroneous. What perhaps appeared to be a second lamina, being found to be simply a cellular coat, which fends off productions to the blood-vessels passing out of the abdominal cavity. The aorta and vena cava likewise de- rive a covering from the same membrane, which seems to be a part of the cellular membrane we have already described. The peritonæum, by its productions and re- duplications, envelopes the greatest part of the abdominal viscera. It is soft, and capable of considerable extension; and is kept smooth and moist by a vapour, which is constantly exhaling from its inner surface, and is returned again into the circulation by the absorbents. This moisture not only contributes to the soft- ness of the peritonæum, but prevents the attri- tion, and other ill effects which would other- wise probably be occasioned, by the motion of the viscera upon each other. When this fluid is supplied in too great a quantity, or the absorbents become incapable of carrying it off, it accumulates, and constitutes an ascites or dropsy of the belly; and when by any means the exhalation is discontinued, the peritonæum thickens, becomes diseased, and the viscera are sometimes found adhering to each other. The peritonæum is not a very vascular mem- 3 brane, 241 PART III. OF THE ABDOMEN. brane. In a sound state it seems to be endued with little or no feeling, and the nerves that pass through it appear to belong to the abdomi- nal muscles. SECT. II. Of the Omentum. The omentum, epiploon, or cawl, is a dou- ble membrane, produced from the peritonæum. It is interlarded with fat, and adheres to the sto- mach, spleen, duodenum, and colon; from thence hanging down loose and floating on the surface of the intestines. Its size is different in different subjects. In some it descends as low as the pel- vis, and it is commonly longer at the left side than the right. This part, the situation of which we have just now described, was the only one known to the ancients under the name of epiploon; but at pre- sent we distinguish three omenta, viz. omentum magnum colico gastricum, omentum parvum hepa- tico gastricum, and omentum colicum. They all agree in being formed of two very delicate la- minæ, separated by a thin layer of cellular mem- brane. The omentum magnum colico gastricum, of which we have already spoken, derives its ar- teries from the splenetic and hepatic. Its veins terminate in the vena portæ. Its nerves, which are very few, come from the splenetic and he- patic plexus. Q The 242 PART III. OF THE ABDOMEN. The omentum parvum hepatico gastricum, abounds less with fat than the great epiploon. It begins at the upper part of the duodenum, ex- tends along the lesser curvature of the stomach as far as the œsophagus, and terminates about the neck of the gall-bladder, and behind the left ligament of the liver, so that it covers the lesser lobe; near the beginning of which we may observe a small opening, first described by Win- slow, through which the whole pouch may ea- sily be distended with air (X). The vessels of the omentum parvum are derived chiefly from the coronary stomachic arteries and veins. The omentum olicum begins at the fore part of the cœcum and right side of the colon. It ap- pears as a hollow conical appendage to these intestines, and usually terminates at the back of the omentum magnum. It seems to be no- thing more than a membranous coat of the cœcum and colon, assuming a conical shape with air. The uses of the omentum are not yet satisfac- torily determined. Perhaps by its softness and looseness it may serve to prevent those adhesions of the abdominal viscera, which have been found to take place when the fat of the omentum has been much wasted. Some authors have sup- posed (X) This membranous bag, though exceedingly thin and transparent, is found capable of supporting mercury, thrown into it by the same channel. 243 PART III. OF THE ABDOMEN. posed, that it assists in the preparation of bile; but this idea is founded merely on conjecture. SECT. III. Of the Stomach. The stomach is a membranous and muscular bag, in shape not unlike a bagpipe, lying across the upper part of the abdomen, and inclining ra- ther more to the left than the right side. It has two orifices, one of which receives the end of the œsophagus, and is called the cardia, and sometimes the left and upper orifice of the stomach; though its situation is not much higher than the other, which is styled the right and in- ferior orifice, and more commonly the pylorus: both these openings are more elevated than the body of the stomach. The aliment passes down the œsophagus into the stomach through the cardia, and after hav- ing undergone the necessary digestion, passes out at the pylorus where the intestinal canal commences. The stomach is composed of four tunics or coats, which are so intimately connected toge- ther that it requires no little dexterity in the ana- tomist to demonstrate them. The exterior one is membranous, being derived from the perito- næum.—The second is a muscular tunic, com- posed of fleshy fibres which are in the greatest number about the two orifices.—The third is called the nervous coat, and within this is the Q2 villous 244 OF THE ABDOMEN. villous or velvet-like coat which composes the inside of the stomach. The two last coats being more extensive than the two first, form the folds, which are observ- ed every where in the cavity of this viscus, and more particularly about the pylorus; where they seem to impede the too hasty exclusion of the aliment, making a considerable plait, called val- vula pylori. The inner coat is constantly moistened by a mucus, which approaches to the nature of the saliva, and is called the gastric juice; this liquor has been supposed to be secreted by certain mi- nute glands (Y) seated in the nervous tunic, whose excretory ducts open to the surface of the villous coat. The arteries of the stomach called the gas- tric arteries are principally derived from the cæ- liac; some of its veins pass to the splentic, and others to the vena portæ; and its nerves are chiefly from the eight pair or par vagum. The account given of the tunics of the sto- mach may be applied to the whole alimentary canal; (Y) Heister, speaking of these glands, very properly says, "in porcis facile, in homine raro observantur;" for although many anatomical writers have described their appearance and figure, yet they do not seem to have been hitherto sa- tisfactorily demonstrated in the human stomach; and the gastric juice is now more generally believed to be derived from the exhalent arteries of the stomach. 145 PART III. OF THE ABDOMEN. canal; for both the œsophagus and intestines are, like this viscus, composed of four coats. Before we describe the course of the aliment and the uses of the stomach, it will be necessary to speak of other parts which assist in the pro- cess of digestion. SECT. IV. Of the Oesophagus. The œsophagus or gullet is a membranous and muscular canal, extending from the bottom of the mouth to the upper orifice of the stomach. —Its upper part where the aliment is received is shaped somewhat like a funnel, and is called the pharynx. From hence it runs down close to the bodies of the vertebræ as far as the diaphragm, in which there is an opening through which it passes, and then terminates in the stomach about the ele- venth or twelfth vertebra of the back. The œsophagus is plentifully supplied with arteries from the external carotid, bronchial, and superior intercostal arteries; its veins empty themselves into the vena azygos, internal jugu- lar, and mammary veins, &c. Its nerves are derived chiefly from the eighth pair. We likewise meet with a mucus in the œsopha- gus, which every where lubricates its inner surface, and tends to assist in deglutition.—This mucus seems to be secreted by very minute glands, like the 246 PART III. OF THE ABDOMFN. the mucus in other parts of the alimentary ca- nal. SECT. V. Of the Intestines. THE intestines form a canal, which is usually six times longer than the body to which it be- longs. This canal extends from the pylorus, or inferior orifice of the stomach, to the anus. It will be easily understood, that a part of such great length must necessarily make many circum- volutions, to be confined with so many other viscera within the cavity of the lower belly. Although the intestines are in fact, as we have observed, only one long and extensive canal, yet different parts have been distinguished by diffe- rent names. The intestines are first distinguished into two parts, one of which begins at the stomach, and is called the thin or small intestines, from the small size of the canal, when compared with the other part, which is called the large intestines, and includes the lower portion of the canal down to the anus. Each of these parts has it its subdivisions.— The small intestines being distinguished into duodenum, jejunum, and ilium, and the larger portion into cœcum, colon, and rec- tum. The small intestines fill the middle and fore parts of the belly, while the large intestines fill the 247 PART III. OF THE ABDOMEN. the sides and both the upper and lower parts of the cavity. The duodenum, which is the first of the small intestines, is so called, because it is about 12 inches long. It begins at the pylorus and terminates in the jejunum, which is a part of the canal observed to be usually more empty than the other intestines.—This appearance gives it its name, and likewise serves to point out where it begins. The next division is the ilium, which of it- self exceeds the united length of the duodenum and jejunum, and has received its name from its numerous circumvolutions. The large circum- volution of the ilium covers the first of the large intestines called the cœcum (X) which seems pro- perly to belong to the colon, being a kind of pouch of about four fingers in width, and near- ly of the same length, having anteriorly a lit- tle appendix, called appendix cœci. The cœcum is placed in the cavity of the os ilium on the right side, and terminates in the colon, which is the largest of all the intestines. This intestine ascends by the right kidney to which it is attached, passes under the hollow part (X) Anatomists have differed with respect to this division of the intestines.—The method here followed is now gene- rally adopted; but there are authors who allow the name of cœcum only to the little appendix, which has likewise been called the vermiform appendix, from its resemblance to a worm in size and length. 248 PART III. OF THE ABDOMEN. part of the liver, and the bottom of the stomach, to the spleen, to which it is likewise secured, as it is also to the left kidney; and from thence passes down towards the os sacrum, where, from its straight course, the canal begins to take the name of rectum. There are three ligamentous bands extending through the whole length of the colon, which, by being shorter than its two inner coats, serve to increase the plaits on the inner surface of this gut. The anus which terminates the intestinum rectum, is furnished with three muscles; one of these is composed of circular fibres, and from its use in shutting the passage of the anus is called sphincter ani. The other two are the levatores ani, so called, because they elevate the anus after dejection. When these by palsy, or any other disease, lose the power of contracting, the anus prolapses; and when the sphincter is affected by similar causes, the fæces are voided involuntarily. It has been already observed, that the intestinal canal is composed of four tunics; but it remains to be remarked, that here, as in the stomach, the two inner tunics being more extensive than the other two, form the plaits which are to be seen in the inner surface of the intestines, and are called valvulæ conniventes. Some authors have considered these plaits as tending to retard the motion of the fæces, in order 249 PART III. OF THE ABDOMEN. order to afford more time for the separation of the chyle; but there are others who attribute to them a different use: they contend, that these valves, by being naturally inclined downwards, can- not impede the descent of the fæces, but that they are intended to prevent their return up- wards. They are probably destined for both these uses; for although these folds incline to their lower side, yet the inequalities they occasion in the ca- nal are sufficient to retard, in some measure, the progressive motion of the fæces, and to afford a greater surface of the absorption of chyle, and their natural position seems to oppose itself to the return of the aliment. Besides these valvulæ conniventes, there is one more considerable than the rest, called the valve of the colon; which is found at that part of the canal where the intestinum ilium is joined to the colon. This valve permits the alimentary pulp to pass downwards, but serves to prevent its return upwards; and it is by this valve, that glysters are prevented from passing into the small intestines (Y). Of the little vermiform appendix of the cœ- cum, it will be sufficient to say, that its uses have never (Y) This is not invariably the case, for the contents of a glyster have been found not only to reach the small intes- tines, but to be voided at the mouth. Such instances, how- ever, are not common. 250 PART III. OF THE ABDOMEN. never yet been ascertained. In birds we meet with two of these appendices. The intestines are lubricated by a constant supply of mucus, which is probably secreted by very minute folicles (Z). This mucus promotes the descent of the alimentary pulp, and in some measure defends the inner surface of the intes- tines from the irritation to which it would, per- haps, otherwise be continually exposed from the aliment; and which, when in a certain degree, excites a painful disorder called colic, a name given to the disease, because its most usual seat is in the intestinum colon. The intestines are likewise frequently distend- ed with air, and this distention sometimes occa- sions pain, and constitutes the flatulent colic. The arteries of the intestines are continuati- ons of the mesenteric arteries, which are deriv- ed in two considerable branches from the aorta. —The redundant blood is carried back into the vena portarum. In (Z) Some writers have distinguished these glands into mi- liary, lenticular, &c.—Brunner and Peyer were the first ana- tomists who described the glands of the intestines, and their descriptions were chiefly taken from animals, these glandular appearances not seeming to have been hitherto satisfactorily pointed out in the human subject.—It is now pretty general- ly believed, that the mucus which every where lubricates the alimentary canal, is exhaled from the minute ends of ar- teries, and that these extremities first open into a hollow ve- sicle, from whence the deposited juice of several branches flows out through one common orifice. 251 PART III. OF THE ABDOMEN. In the rectum the veins are called hemorrhoi- dal, and are there distinguished into internal and external; the first are branches of the inferior mesenteric vein, but the latter pass into other veins. Sometimes these veins are distended with blood from obstructions, from weakness of their coats, or from other causes, and what we call the hæmorrhoids takes place. In this disease they are sometimes ruptured; and the discharge of blood which consequently follows, has probably occasioned them to be called hæmorrhoidal veins. The nerves of the intestines are derived from the eight pair. SECT. VI. Of the Mesentery. The name of the mesentery implies the situa- tion amidst the intestines. It is in fact a part of the peritonæum, being a reduplication (A) of that membrane (A) He who only reads of the reduplication of mem- branes, will perhaps not easily understand how the perito- næum and pleura are reflected, over the viscera in their several cavities; for one of these serves the same purposes in the tho- rax that the other does in the abdomen. This disposition, for the discovery of which we are indebted to modern anato- mists, constitutes a curious part of anatomical knowledge: but the student, unaided by experience, and assisted only by what the limits of this work would permit us to say on the occasion, would probably imbibe only confused ideas of the matter; and it will perfectly answer the present purpose, if he considers the mesentery as a membrane attached by one of its sides to the lumbar vertebræ, and by the other to the intestines. 252 PART III. OF THE ABDOMEN. membrane from each side of the lumbar verte- bræ, to which it is firmly attached, so that it is formed of two laminæ, connected to each other by cellular membrane. The intestines, in their different circumvolu- tions, form a great number of arches, and the mesentry accompanies them through all these turns; but by being attached only to the hollow part of each arch; it is found to have only a third of the extent of the intestines. That part of this membrane which accompa- nies the small intestines is the mesentery, properly so called; but those parts of it which are attach- ed to the colon and rectum are distinguished by the names of meso-colon and meso-rectum. There are many conglobate glands dispersed through this double membrane, through which the lacteals and lymphatics pass in their way to the thoracic duct. The blood-vessels of the mesentery were described in speaking of the in- testines. This membrane, by its attachment to the vertebræ, serves to keep the intestines in their na- tural situation. The idea usually formed of the colic called miserere, is perfectly erroneous; it being impossible that the intestines can be twisted, as many suppose they are, in that disease, their attachment to the mesentery effectually prevent- ing such an accident—but a disarrangement sometimes takes place in the intestinal canal it- self, which is productive of disagreeable and some- 253 PART III. OF THE ABDOMEN. sometimes fatal consequences.—This is by an in- trosusception of the intestine, an idea of which may be easily formed, by taking the finger of a glove, and involving one part of it within the other. If inflammation takes place, the stricture in this case is increased, and the peristaltic motion of the intestines (by which is meant the pro- gressive motion of the fæces downwards) is in- verted, and what is called the iliac passion takes place. The same effect may be occasioned by a descent of the intestine, or of the omentum either with it or by itself, and thus constituting what is called an hernia or rupture; a term by which in general is meant the falling down or protrusion of any part of the intestine or omen- tum, which ought naturally to be contained with- in the cavity of the belly. To convey an idea of the manner in which such a descent takes place, it will be necessary to observe, that the lower edge of the tendon of of the musculus obliquus externus, is stretched from the fore-part of the os ilium or haunch- bone of the os pubis, and constitutes what is called Poupart's or Fallopius's ligament, forming an opening, through which pass the great crural artery and vein. Near the os pubis the same tendinous fibres are separated from each other, and form an opening on each side, called the abdominal ring, through which the spermatic vessels pass in men, and the ligamenta uteri in women. 254 PART III. OF THE ABDOMEN. women. In consequence of violent efforts, or perhaps of natural causes, the intestines are found sometimes to pass through these openings: but the peritonæum which incloses them when in their natural cavity, still continues to surround them even in their natural cavity, still continues to surround them even in their descent. This membrane does not become torn or lacerated by the violence, as might be easily imagined; but its dilatibility enables it to pass out with the vis- cus, which it incloses as it were in a bag, and thus forms what is called the hernial sac. If the hernia be under Poupart's ligament, it is called femoral; if in the groin, inguinal (B), and scrotal, if in the scrotum. Different names are likewise given to the hernia as the contents of the sac differ, whether of omentum only or intestine, or both;—but these definitions more properly belong to the province of surgery. SECT. VII Of the Pancreas. THE pancreas is a conglomerate gland placed behind the bottom of the stomach, towards the first vertebra of the loins; shaped like a dog's tongue with its point stretched out towards the spleen, and its other end extending towards the duodenum. It is about eight fingers breadth in length, (B) The hernia congenita will be considered with the male organs of generation, with which it is intimately connected. 255 PART III. OF THE ABDOMEN. length, two or three in width, and one in thick- ness. This viscus, which is of a yellowish colour, somewhat inclined to red, is covered with a mem- brane which it derives from the peritonæum. Its arteries, which are rather numerous than large, are derived chiefly from the splenic and hepatic, and its veins pass into the veins of the same name.—Its nerves are derived from the in- tercostal. The many little glands of which it has been observed the pancreas is composed, all serve to secrete a liquor called the pancreatic juice, which in its colour, consistence, and other properties, does not seem to differ from the saliva. Each of these glands sends out a little excretory duct, which, uniting with others, help to form larger ducts; and all these at last terminate in one com- mon excretory duct (first discovered by Virtsun- gus in 1642), which runs through the middle of the gland, and is now usually called ductus pancreaticus Virtsungi. This canal opens into the intestinum duodenum, sometimes by the same orifice with the biliary duct, and sometimes by a distinct opening. The liquor it discharges be- ing of a mild and insipid nature, serves to dilute the alimentary pulp, and to incorporate it more easily with the bile. SECT. 256 PART III. OF THE ABDOMEN. SECT. VIII. Of the Liver. The liver is a viscus of considerable size, and of a reddish colour; convex superiority and an- teriorly where it is placed under the ribs and di- aphragm, and of an unequal surface posteriorly. It is chiefly situated in the right hypochondrium, and under the false ribs; but it likewise extends into the epigastric region, where it borders upon the stomach. It is covered by a production of the peritonæum, which serves to attach it by three of its reduplications to the false ribs. These reduplications are called ligaments, though very different in their texture from what are called by the same name in other parts of the body. The umbilical cord, too, which in the fœtus is per- vious, gradually becomes a simple ligament after birth; and, by passing to the liver, serves like- wise to secure it in its situation. At the posterior part of this organ where the umbilical vessels enter, it is found divided into two lobes. Of these, the largest is placed in the right hypochondrium; the other, which covers part of the stomach, is called the little lobe. All the vessels which go to the liver pass in at the fissure we have mentioned; and the production of the peritonæum, which invests the liver, was described by Glisson, an English anatomist, as accompanying them in their passage, and sur- rounding them like a glove; hence this produc- 2 tion 257 PART III. OF THE ABDOMEN. tion has been commonly known by the name of capsula of Glisson: but it appears to be chiefly a continuation of the cellular membrane which co- vers the vena porta ventralis. The liver was considered by the ancients as an organ destined to prepare and perfect the blood; but later discoveries have proved, that this opinion was wrong, and that the liver is a glandular substance formed for the secretion of the bile. The blood is conveyed to the liver by the he- patic artery and the vena porta. This is contra- ry to the mode of circulation in other parts, where veins only serve to carry off the redundant blood: but in this viscus the hepatic artery, which is derived from the cæliac, is principally destined for its nourishment; and the vena por- ta, which is formed by the union of the veins from most of the abdominal viscera, furnishes the blood from which the bile is chiefly to be separat- ed; so that these two series of vessels serve very distinct purposes. The vena porta, as it is rami- fied through the liver, performs the office both of a vein and an artery; for like the former it returns the blood from the extremities of arte- ries, while as the latter it prepares it for secre- tion. The nerves of the liver are branches of the intercostal and par vagum. The bile, after be- ing separated from the mass of blood, in a man- ner of which mention will be made in another R place 258 PART III. OF THE ABDOMEN. place, is conveyed out of this organ by very mi- nute excretory ducts, called poribiliarii; these uniting together like the excretory ducts in the pancreas, gradually form larger ones, which at length terminate in a considerable canal called ductus hepaticus. SECT. IX. Of the Gall-bladder. The gall-bladder is a little membranous bag, shaped like a pear, and attached to the posterior and almost inferior part of the great lobe of the liver. It has two tunics; of which the exterior one is a production of the peritonæum. The inte- rior, or villous coat, is supplied with a mucus that defends it from the acrimony of the bile. These two coverings are intimately connected by means of cellular membrane, which from its firm glistening appearance has generally been spoken of as a muscular tunic. The gall-bladder is supplied with blood-vessels from the hepatic arteries. These branches are called the cystic arteries, and the cystic veins car- ry back the blood. Its nerves are derived from the same origin as those of the liver. The neck of the gall-bladder is continued in the form of a canal called ductus cysticus, which soon unites with the ductus hepaticus we describ- ed as the excretory duct of the liver; and form- ing 259 PART III. OF THE ABDOMEN. ing one common canal, takes the name of ductus choledochus communis, through which both the cystic and hepatic bile are discharged into the du- odenum. This canal opens into the intestine in an oblique direction, first passing through the exterior tunic, and then piercing the other coats after running between each of them a very little way. This œconomy serves two useful pur- poses;—to promote the discharge of bile and to prevent its return. The bile may be defined to be a natural liquid soap, somewhat unctuous and bitter, and of a yellowish colour, which easily mixes with water, oil, and vinous spirits, and is capable of dissolv- ing resinous substances. From some late experi- ments made by M. Cadet*, it appears to be formed of an animal oil, combined with the al- kaline base of sea-salt, a salt of the nature of milk, and a calcareous earth which is slightly fer- ruginous. Its definition seems sufficiently to point out the uses for which it is intended (C). It blends the alimentary mass, by dividing and attenuating it; corrects the too great disposition to acescency, which the aliment acquires in the stomach; and, finally, by its acrimony, tends to excite the pe- ristaltic motion of the intestines. R2 After * Mem. de l' Acad. des Sciences. 1767. (C) The ancients, who were not acquainted with the real use of the liver, considered the bile as an excrementitious and useless fluid. 260 PART III. OF THE ABDOMEN. After what has been said, it will be conceived that there are two sorts of bile; one of which is derived immediately from the liver through the hepatic duct, and the other from the gall-blad- der. These two biles, however, do not essenti- ally differ from each other. The hepatic bile in- deed is milder, and more liquid than the cystic, which is constantly thicker and yellower; and by being bitterer, seems to possess greater activity than the other. Every body knows the source of the hepatic bile, that it is secreted from the mass of blood by the liver; but the origin of the cystic bile has occasioned no little controversy amongst anato- mical writers. There are some who contend, that it is separated in the substance of the liver, from whence it passes into the gall-bladder through particular vessels. In deer, and in some other quadrupeds, as well as in several birds and fishes, there is an evident communication, by means of particular vessels, between the liver and the gall-bladder. Bianchi, Winslow, and others, have asserted the existence of such vessels in the human subject, and named them hepati- cystic ducts; but it is certain that no such ducts exist.—In obstructions of the cystic duct, the gall-bladder has been found shrivelled and emp- ty: so that we may consider the gall-bladder as a reservoir of hepatic bile; and that it is an es- tablished fact, that the whole of the bile con- tained in the gall-bladder is derived from the li- ver; 261 PART III. OF THE ABDOMEN. ver; that it passes from the hepatic to the cystic dud, and from that to the gall-bladder. The difference in the colour, consistence, and taste of the bile, is merely the consequence of stagnation and absorption. When the stomach is distend- ed with aliment, this reservoir undergoes a cer- tain degree of compression, and the bile passes out into the intestinal canal; and in the efforts to vomit, the gall-bladder seems to be constantly af- fected, and at such times discharges itself of its contents. Sometimes the bile concretes in the gall-blad- der, so as to form what are called gall-stones (D). When these concretions pass into the cystic duct, they sometimes occasion exquisite pain, by dis- tending the canal in their way to the duodenum; and by lodging in the ductus choledochus com- munis, and obstructing the course of the bile, this fluid will be absorbed, and by being carried back into the circulation occasion a temporary jaundice. SECT. X. Of the Spleen. The spleen is a soft and spongy viscus, of a bluish (D) These concretions sometimes remain in the gall-blad- der without causing any uneasiness. Dr. Heberden relates, that a gall-stone weighing two drams was found in the gall- bladder of the late Lord Bath, though he had never com- plained of the jaundice, nor of any disorder which he could attribute to that cause. Med. Trans. Vol. ii. 262 PART III. OF THE ABDOMEN. bluish colour, and about five or six fingers breadth in length, and three in width, situ- ated in the left hypochondrium, between the stomach and the false ribs. That side of it which is placed on the side of the ribs is convex; and the other, which is turned toward the stomach, is concave. The splenic artery, which is a branch from the cæliac, supplies this viscus with blood, and a vein of the same name carries it back into the vena porta. Its nerves are derived from a particular plexus called the splenic, which is formed by branches of the intercostal nerve, and by the eight pair, or par vagum. The ancients, who supposed two sorts of bile, considered the spleen as the receptacle of what they called atra bilis. Havers, who wrote pro- fessedly on the bones, determined its use to be that of secreting the synovia; and the late Mr. Hewson imagined, that it concurred with the thymus and lymphatic glands of the body in forming the red globules of the blood. All these opinions seem to be equally fanciful. The want of an excretory duct has occasioned the real use of this viscus to be still doubtful. Perhaps the blood undergoes some change in it, which may assist in the preparation of the bile. This is the opinion of the generality of modern physiolo- gists; and the great quantity of blood with which it is supplied, together with the course of its 263 PART III. OF THE ABDOMEN. its veins into the vena portæ, seem to render this notion probable. SECT. XI Of the Glandulæ Renales, Kidneys, and Ureters. The glandulæ renales, which were by the ancients supposed to secrete the atra bilis, and by them named capsulæ atrabilares, are two flat bo- dies of an irregular figure, one on each side be- tween the kidney and the aorta. In the fœtus they are as large as the kidneys: but they do not increase afterwards in proportion to those parts; and in adults and old people they are generally found shrivelled, and much wasted, They have their arteries and veins. Their ar- teries usually arise from the splenic or the emul- gent, and sometimes from the aorta; and their veins go to the neighbouring veins, or to the vena cava. Their nerves are branches of the intercostal. The use of these parts is not yet perfectly known. In the fetus the secretion of urine must be in a very small quantity, and a part of the blood may perhaps then pass through these chan- nels, which in the adult is carried to the kidneys to supply the matter of urine. The kidneys are two in number, situated one on the right and the other on the left side in the lumbar region, between the last false rib and the os ilium, by the sides of the vertebræ. Each kidney 264 PART III. OF THE ABDOMEN. kidney in its figure resembles a sort of bean, which from its shape is called kidney-bean. The concave part of each kidney is turned towards the aorta and vena cava ascendens. They are surrounded by a good deal of fat, and receive a coat from the peritonæum; and when this is removed, a very fine membrane is found invest- ing their substance and the vessels which ramify through them. Each kidney has a considerable artery and vein, which are called the emulgent. The arte- ry is a branch from the aorta, and the vein passes into the vena cava. Their nerves, which every where accompany the blood-vessels, arise from a considerable plexus, which is derived from the intercostal. In each kidney, which in the adult is of a pretty firm texture, there are three substances to be distinguished (E). The outer part is glandu- lar or cortical, beyond this is the vascular or tu- bular substance, and the inner part is papillary or membranous. It is in the cortical part of the kidney that the secretion is carried on; the urine being here re- ceived from the minute extremities of the capilla- ry arteries, is conveyed out of this cortical sub- stance by an infinite number of very small cylin- drical canals or excretory vessels, which consti- tute (E) The kidneys in the fœtus are distinctly lobulated; but in the adult they become perfectly firm, smooth, and re- gular. 265 PART III. OF THE ABDOMEN. tute the tubular part. These tubes, as they ap- proach the inner substance of the kidney, gradu- ally unite together; and thus forming larger ca- nals, at length terminate in ten or twelve little protuberances called papillæ, the orifices of which may be seen without the assistance of glasses. Thee papillæ open into a small cavity or refer- voir called the pelvis of the kidney, and formed by a distinct membranous bag which embraces the papillæ. From this pelvis the urine is con- veyed through a membranous canal which passes out from the hollow side of the kidney, a little below the blood vessels, and is called ureter. The ureters are each about as large as a com- mon writing-pen. They are somewhat curved in their course from the kidneys, like the letter s, and at length terminate in the posterior and almost inferior part of the bladder, at some dis- tance from each other. They pass into the blad- der in the same manner as the ductus choledoc- dus communis passes into the intestinum duode- num, not by a direct passage, but by an oblique course between the two coats; so that the dis- charge of urine into the bladder is promoted, whilst its return is prevented. Nor does this mode of structure prevent the passage of fluids only from the bladder into the ureters, but like- wise air:—for air thrown into the bladder in- flates it, and it continues to be distended if a ligature is passed round its neck; which seems to 266 PART III. OF THE ABDOMEN. to prove sufficiently that it cannot pass into the ureters. SECT. XII. Of the Urinary Bladder. The urinary bladder is a membranous and muscular bag of an oblong roundish shape, situ- ated in the pelvis, between the os pubis and in- testinum rectum in men, and between the os pu- bis and uterus in women. Its upper and widest part is usually called the bottom, its narrow part the neck of the bladder; the former only is co- vered by the peritonæum. The bladder is formed of three coats, connect- ed together by means of cellular membrane. The external or peritonæal, is only a partial one, covering the upper and back part of the bladder. The middle, or muscular coat, is composed of irritable, and of course muscular fibres, which are most collected around the neck of the blad- der, but not so as to form a distinct muscle, or sphincter, as the generality of anatomists have hitherto supposed. The inner coat, though much smoother, has been said to resemble the villous tunic of the in- intestines, and like that is provided with a mu- cus, which defends it against the acrimony of the urine. It will be easily conceived from what has been said, that the kidneys are two glandular bo- dies, through which a saline and excrementitious fluid 267 PART III. OF THE ABDOMEN. fluid called urine is constantly filtering from the mass of blood. While only a small quantity of urine is col- lected in the bladder, it excites no kind of un- easiness; but when a greater quantity is accu- mulated, so that the bladder is distended in a cer- tain degree, it excites in us a certain sensation, which brings on as it were a voluntary contrac- tion of the bladder to promote its discharge.— But this contraction is not effected by the mus- cular fibres of the bladder alone: for all the ab- dominal muscles contract in obedience to our will, and press downwards all the viscera of the lower belly; and these powers being united, at length overcome the resistance of the fibres sur- rounding the neck of the bladder, which dilates and affords a passage to the urine through the urethra. The frequency of this evacuation depends on the quantity of urine secreted; on the degree of acrimony it possesses; on the size of the bladder, and on its degree of sensibility. The urine varies much in its colour and con- tents. These varieties depend, on age, sex, climate, diet, and other circumstances. In infants it is ge- nerally a clear watery fluid, without smell or taste. As we advance in life, it acquires more colour and smell, and becomes more impregnat- ed with salts. In old people it becomes still more acrid and fetid. In 268 PART III. OF THE ABDOMEN. In a healthy state it is nearly of a straw co- lour.—After being kept for some time, it depo- sites a tartarous matter, which is found to be composed chiefly of earth and salt, and soon in- crusts the sides of the vessel in which it is con- tained. While this separation is taking place, appearances like minute fibres or threads of a whitish colour, may be seen in the middle of the urine, and an oily scum observed floating on its surface. So that the most common appearances of the urine are sufficient to ascertain that it is a watery substance, impregnated with earthy, sa- line, and oily particles. The urine is not always voided of the same colour and consistence; for these are found to depend on the proportion of its watery part to that of its other constituent principles.—Its co- lour and degree of fluidity seem to depend on the quantity of saline and inflammable particles contained in it: so that an increased proportion of those parts will constantly give the urine a higher colour, and add to the quantity of sedi- ment. The variety in the appearance of the urine, depends on the nature and quantity of solid and fluid aliment we take in; and it is likewise occa- sioned by the different state of the urinary vessels, by which we mean the channels through which it is separated from the blood, and conveyed through the pelvis into the ureters. The causes of calculous concretions in the urinary passages, are 269 PART. III OF THE ABDOMEN. are to be looked for in the natural constitution of the body, mode of life, &c. It having been observed, that after drinking any light wine or Spa water, it very soon passed off by urine, it has been supposed by some, that the urine is not altogether conveyed to the blad- der by the ordinary course of circulation, but that there must certainly exist some other shorter means of communication, perhaps by certain vessels between the stomach and the bladder, or by a retrograde motion in the lymphatics. But it is certain, that if we open the belly of a dog, press out the urine from the bladder, pass a liga- ture round the emulgent arteries, and then sew up the abdomen, and give him even the most diuretic liquor to drink, the stomach and other channels will be distended with it, but not a drop of urine will be found to have passed into the bladder; or the same thing happens when a liga- ture is thrown round the two ureters. This ex- periment then seems to be a sufficient proof, that all the urine we evacuate, is conveyed to the kidneys through the emulgent arteries, in the manner we have described.—It is true, that wine and other liquors promote a speedy evacuation of urine: but the discharge seems to be merely the effect of the stimulus they occasion; by which the bladder and urinary parts are solicited to a more co- pious discharge of the urine, which was before in the body, and not immediately of that which was last drank; and this increased discharge, if the supply 270 PART III. OF THE ABDOMEN. supply kept up, will continue: nor will this appear wonderful, if we confider the great capa- city of the vessels that go to the kidneys; the constant supply of fresh blood that is essential to health; and the rapidity with which it is inces- santly circulated through the heart to all parts of the body. SECT. XIII. Of Digestion. We are now proceeding to speak of digestion, which seems to be introduced in this place with propriety, after a description of the abdominal viscera, the greater part of which contribute to this function. By digestion is to be understood, the changes the aliment undergoes for the forma- tion of chyle:—these changes are effected in the mouth, stomach, and small intestines. The mouth, of which every body has a ge- neral knowledge, is the cavity between the two jaws, formed anteriorly and laterally by the lips, teeth; and cheeks, and terminating posteriorly in the throat. The lips and cheeks are made up of fat and muscles, covered by the cuticle, which is conti- nued over the whole inner surface of the mouth, like a fine and delicate membrane.—Beside this membrane, the inside of the mouth is furnished with a spongy and very vascular substance called the gums, by means of which the teeth are se- cured in their sockets. A similar substance covers the roof of the mouth, and forms what is called the 271 PART III. OF THE ABDOMEN. the velum pendulum palati, which is fixed to the extremity of the arch formed by the ossa max- illaria and ossa palati, and terminates in a soft, small, and conical body, named uvula; which appears, as it were, suspended from the middle of the arch over the basis of the tongue. The velum pendulum palati performs the of- fice of a valve between the cavity of the mouth and the pharynx, being moved by several mus- cles (F). The tongue is composed of several muscles (G) which enable it to perform a variety of mo- tions for the articulation of the voice; for the purposes of mastication; and for conveying the aliment into the pharynx. Its upper part is co- vered with papillæ, which constitute the organ of taste, and are easily to be distinguished; it is covered by the same membrane that lines the in- side of the mouth, and which makes at its infe- rior part towards its basis a reduplication called frænum. Posteriorly, under the veium palati, and at the basis of the tongue, is the pharynx: which is the beginning of the œsophagus, stretched out every way, so as to resemble the top of a fun- 3 nel, (F) These are the circumflexus palati, levator palati mol- lis, palato-pharyngæus constrictor isthmi saucium and azygos uvulæ. (G) These are, the genio-glossus, hyo-glossus, lingualis, and stylo-glossus. 272 PART III. OF THE ABDOMEN. nel, through which the aliment passes into the stomach. The mouth has a communication with the nostrils at its posterior and upper part; with the ears, by the Eustachian tubes; with the lungs, by means of the larynx; and with the stomach, by means of the œsophagus. The pharynx is constantly moistened by a fluid, secreted by two considerable glands called the tonsils, one on each side of the velum palati. These glands, from their supposed resemblance to almonds, have likewise been called amyg- dalis. The mouth is moistened by a considerable quantity of saliva. This fluid is derived from the parotid glands; a name which by its ety- mology points out their situation to be near the ears. They are two in number, one on each side under the os malæ: and they are of the con- glomerate kind; being formed of many smaller glands, each of which sends out a very small ex- cretory duct, which unites with the rest, to form one common channel, that runs over the cheek, and piercing the buccinator muscle, opens into the mouth on each side, by an orifice into which a bristle may be easily introduced.—Besides these, the maxillary glands, which are placed near the inner surface of the angle of the lower jaw on each side; the sublingual glands, which are situated at the root of the tongue ; the glands of the palate, which are seated in the velum pa- lati; 273 PART III. OF THE ABDOMEN. lati; and those of the cheeks, lips, &c. together with many other less considerable ones,—pour the saliva into the mouth through their several excretory ducts. The saliva, like all the other humours of the body, is found to be different in different peo- ple: but in general, it is a limpid and insipid fluid, without smell in healthy subjects; and these properties would seem to prove that it contains very few saline or inflammable particles. The uses of the saliva seem to be to moisten and lubricate the mouth, and to assist in reducing the aliment into a soft pulp before it is conveyed into the stomach. The variety of functions which are constant- ly performed by the living body, must necessa- rily occasion a continual waste and dissipation of its several parts. A great quantity is every day thrown off by the insensible perspiration and other discharges; and were not these losses constantly recruited by a fresh supply of chyle, the body would soon effect its own disso- lution. But nature has very wisely favour- ed us with organs fitted to produce such a sup- ply; and has at the same time endued us with the sensations of hunger and thirst, that our at- tention may not be diverted from the necessary business of nutrition. The sensation of hunger is universally known; but it would perhaps be difficult to describe it perfectly in words. It may, however, be defined to be a certain uneasy sensation S in 274 PART III. OF THE ABDOMEN. in the stomach, which induces us to wish for so- lid food; and which likewise serves to point out the proper quantity, and time for taking it. In describing the stomach, mention was made of the gastric juice, as every where lubricating its inner coat. This humour mixes itself with the aliment in the stomach, and helps to prepare it for its pas- sage into the intestines; but when the stomach is perfectly empty, this same fluid irritates the coats of the stomach itself, and produces the sensation of hunger. A certain proportion of liquid aliment is required to assist in the process of digestion, and to afford that moisture to the body, of which there is such a constant dissipation.—Thirst in- duces us to take this necessary supply of drink; and the seat of this sensation is in the tongue, fauces, and œsophagus, which from their great sensibility are required to be kept moist : for though the fauces are naturally moistened by the mucus and salival juices; yet the blood, when deprived of its watery part or rendered acrimo- nious by any natural causes, never fails particu- larly to affect these parts, and the whole alimen- tary canal, and to occasion thirst.—This is the common effect of fevers and of hard labour, by both which too much of the watery part of the blood is dissipated. Mastication and deglutition. It has been ob- served, that the aliment undergoes some prepa- tion in the mouth before it passes into the sto- mach; 275 PART III. OF THE ABDOMEN. mach; and this preparation is the effect of mas- tication. In treating of the upper and lower jaws, mention was made of the number and ar- rangement of the teeth. The upper jaw was described as being immoveable; but the lower jaw was spoken of as being capable of eleva- tion and depression, and of a grinding motion. The aliment, when first carried into the mouth, is pressed between the teeth of the two jaws by a very strong and frequent motion of the lower jaw; and the tongue and the cheeks assisting in this process, continue to replace the food be- tween the teeth till it is perfectly divided, and reduced to the consistence of pulp. The inci- sores and canini divide it first into smaller pieces, but it is between the surfaces of the dentes mo- lares by the grinding motion of the jaw that the mastication is completed. During this process, the salival glands being gently compressed by the contraction of the mus- cles that move the lower jaw, pour out their sa- liva: this helps to divide and break down the food, which at length becomes a kind of pulp, and is then carried over the basis of the tongue into the fauces. But to effect this passage into the œsophagus, it is necessary that the other openings which were mentioned as having a communication with the mouth as well as the pharynx, should be closed; that none of the ali- ment, whether solid or liquid, may pass into them, whilst the pharynx alone is dilated to re- S2 ceive 276 PART III. OF THE ABDOMEN. ceive it:—And such a disposition actually takes place in a manner we will endeavour to de- scribe. The trachea arteria, or windpipe, through which the air is conveyed to the lungs, is plac- ed before the œsophagus—in the act of swallow- ing, therefore, if the larynx (for so the upper part of the trachea is called) is not closed, the aliment will pass into it in its way to the œso- phagus. But this is prevented by a small and very elastic cartilage, called epiglottis, which is attached only to the fore-part of the larynx; so that the food in its passage to the œsophagus presses down this cartilage, which then covers the glot- tis or opening of the larynx; and at the same time the velum palati being capable of some de- gree of motion, is drawn backwards by its mus- cles, and closes the openings into the nose and the Eustachian tubes.—This, however, is not all. The larynx, which being composed of car- tilaginous rings, cannot fail in its ordinary state to compress the membranous canal of the œso- phagus, is in the act of deglutition carried for- wards and upwards by muscles destined for that purpose; and consequently drawing the fore- part of the pharynx with it, that opening is ful- ly dilated. When the aliment has reached the pharynx, its descent is promoted by its own pro- per weight, and by the muscular fibres of the œsophagus, which continue to contract from above downwards, until the aliment has reach- ed 277 PART III. OF THE ABDOMEN. ed the stomach. That these fibres have no in- considerable share in deglutition, any person may experience, by swallowing with his head down- wards, when the descent of the aliment cannot possibly be effected by its weight. It is necessary that the nostrils and the lungs should communicate with the mouth, for the purposes of speech and respiration: but if the most minute part of our food happens to be in- troduced into the trachea, it never fails to pro- duce a violent cough, and sometimes the most alarming symptoms. This is liable to happen when we laugh or speak in the act of degluti- tion: the food is then said to have passed the wrong way. And indeed this is not improper- ly expressed: for death would soon follow, if the quantity of aliment introduced into the trachea should be sufficient to obstruct the respi- ration only during a very short time; or if the irritating particles of food should not be thrown up again by means of the cough, which in these cases very seasonably increases in proportion to the degree of irritation. If the velum palati did not close the passage to the nostrils, deglutition would be performed with difficulty, and perhaps not at all; for the ali- ment would return through the nose, as is some- times the case in drinking. Children, from a deficiency in this velum palati, have been seen to die a few hours after birth; and they who from 278 PART III. OF THE ABDOMEN. from disease or any other causes have not this part perfect, swallow with difficulty. The aliment after having been sufficiently di- vided by the action of the teeth, and attenuated by the saliva, is received into the stomach, where it is destined to undergo a more considerable change. The properties of the aliment not being much altered at its first entrance into the stomach, and before it is thoroughly blended with the gastric juice, is capable of irritating the inner coat of the stomach to a certain degree, and occasions a contraction of its two orifices.—In this mem- branous bag, surrounded by the abdominal vis- cera, and with a certain degree of natural heat, the aliment undergoes a constant agitation by means of abdominal muscles and of the dia- phragm, and likewise by a certain contraction or expansion of the muscular fibres of the stomach itself. By this motion, every part of the food is exposed to the action of the gastric juice, which gradually divides and attenuates it, and prepares it for its passage into the intestines. Some observations lately published by Mr. Hunter in the Philosophical Transactions, tend to throw considerable light on the principles of digestion. There are few dead bodies in which the stomach, at its great end, is not found to be in some degree digested (H). Animals, or parts of (H) The Abbe Spallanzani, who has lately written upon di- gestion, finds, from a variety of experiments, made upon quadru- 279 PART III. OF THE ABDOMEN. of animals, possessed of the living principle, when taken into the stomach, are not in the least affected by the action of that viscus; but the moment they lose the living principle, they become subject to its digestive powers. This seems to be the case with the stomach, which is enabled to resist the action of its juices in the living body: but when deprived of the living principle, it is then no longer able to resist the powers of that menstruum, which it had itself formed for the digestion of its contents; the pro- cess of digestion appearing to be continued after death. This is confirmed by what happens in the stomachs of fishes: they frequently swal- low, without mastication, fish which are larger than the digesting parts of their stomach can contain; quadrupeds, birds, and fishes, that digestion goes on for some time after death, though far less considerable than in living animals, or at least promotes it in a much greater de- gree. He found also, that when the stomach was cut out of the body, it had somewhat of the power of digestion, though this was trifling when compared with that which took place when the stomach was left in the body. In not one of the animals was the great curvature of the stomach dissolved, or much eroded after death. There was often a little ero- sion, especially in different fishes; in which, when he had cleared the stomach of its contents, the internal coat was wanting. In other animals there was only a slight excoria- tion; and the injury in all of them was at the inferior part, or great curvature. The coats of the stomach suffer less af- ter death than flesh, or part of the stomach of similar animals put into it: the author assigns as a reason for this, that these bodies are invested on all sides by the gastric fluid, whereas it only acts on the internal surface of the stomach. 280 PART III. OF THE ABDOMEN. contain; and in such cases, that part which is taken into the stomach is more or less dissolved, while that part which remains in the œso- phagus is perfectly sound; and here, as well as in the human body, the digesting part of the stomach is often reduced to the same state as the digested part of the food. These appearances tend to prove, that digestion is not effected by a mechanical power, by contractions of the stomach, or by heat; but by a fluid se- creted in the coats of the stomach, which is poured into its cavity, and there animalizes the food, or assimilates it to the nature of the blood. From some late experiments by M. Sage*, it appears, that inflammable air has the property of destroying and dissolving the animal texture: and as we swallow with the substances which serve us for food a great quantity of atmosphe- rical air, M. Sage thinks it possible, that de- plogisticated, which is its principle, may be con- verted in the stomach into inflammable air, or may modify into inflammable air a portion of the oily substance which is the principle of ali- ments. In this case, would not the inflammable air (he asks), by dissolving our food, facilitate its conversion into chyle? Be this as it may, the food, after having re- mained one, two, or three hours in the stomach, is * Hist. de l'Academie royale des Sciences, &c. pour 1784. mem. 15. 281 PART III. OF THE ABDOMEN. is converted into a greyish pulp, which is usual- ly called chymus, a word of Greek etymology, signifying juice, and some few milky or chylous particles begin to appear.—But the term of its residence in this bag is proportioned to the na- ture of the aliment, and to the state of the sto- mach and its juices. The thinner and more perfectly digested parts of the food pass by a lit- tle at a time into the duodenum, through the py- lorus, the fibres of which relax to afford it a pas- sage; and the grosser and less digested particles remain in the stomach, till they acquire a suffi- cient fluidity to pass into the intestines, where the nature of the chymus is perfectly changed. The bile and pancreatic juice which flow into the duodenum, and the mucus, which is every where distilled from the surface of the intestines, mix themselves with the alimentary pulp; which they still farther attenuate and dissolve, and in- to which they seem to infuse new properties. Two matters very different from each other in their nature and destination, are the result of this combination.—One of these, which is com- posed of the liquid parts of the aliment, and of some of its more solid particles, extremely di- vided and mixed with the juices we have de- scribed, constitutes a very mild, sweet, and whitish fluid, resembling milk, and distinguish- ed by the name of chyle. This fluid is absorb- ed by the lacteal veins, which convey it into the circulation, where, by being assimilated into the 282 PART III. OF THE ABDOMEN. the nature of blood, it affords that supply of nu- trition which the continual waste of the body is found to require.—The other is the remains of the alimentary mass deprived of all its nu- tritious particles, and containing only such parts as were rejected by the absorbing mouths of the lacteals. This grosser part, called the fæces, passes on through the course of the intestines, to be voided at the anus, as will be explained here- after; for this process in the œconomy cannot be well understood till the motion of respiration has been explained. But the structure of the intestines is a subject which may be properly described in this place, and deserves to be at- tended to. It has been already observed, that the intesti- nal canal is five or six times as long as the body, and that it forms many circumvolutions in the cavity of the abdomen, which it traverses from the right to the left, and again from the left to the right; in one place descending, and in ano- ther extending itself upwards. It was noticed likewise, that the inner coat of the intestines, by being more capacious than their exterior tunics, formed a multitude of plaits placed at a certain distance from each other, and called valvulæ con- niventes. Now this disposition will be found to afford a farther proof of that divine wisdom, which the anatomist and physiologist cannot fail to dis- cover in all their pursuits.—For if the intesti- nal canal was much shorter than it naturally is; if instead of the present circumvolutions it pass- ed 283 PART III. OF THE ABDOMEN. ed in a direct course from the stomach; and if its inner surface was smooth and destitute of valves; the aliment would consequently pass with great rapidity to the anus, and sufficient time would be wanting to assimilate the chyle, and for the necessary absorption of it into the lacteals: so that the body would be deprived of the supply of nutrition, which is so essential to life and health; but the length and circumvo- lutions of the intestines, the inequality of their internal surface, and the course of the aliment through them, all concur to perfect the separa- tion of the chyle from the fæces, and to afford the necessary nourishment to the body. SECT. XIV. Of the Course of the Chyle, and of the Lymphatic System. An infinite number of very minute vessels, called the lacteal veins, arise like net-work from the inner surface of the intestines, (but princi- pally from the jejunum and ilium), which are distended to imbibe the nutritious fluid or chyle. These vessels, which were discovered by Asel- lius in 1622 (I), pass obliquely through the coats (I) We are informed by Galen, that the lacteals had been seen in kids by Erasistratus, who considered them as arteries carrying a milky fluid: but from the remote time in which he lived, they do not seem to have been noticed till they were discovered in a living dog by Asellius, who denominated them lacteals, and considered them as serving to convey the chyle from the intestines to the liver; for before the discovery of the thoracic duct, the use of the liver was universally sup- posed 284 PART III. OF THE ABCOMEN. coats of the intestine, and running along the mesentery, unite as they advance, and form larger branches, all of which pals through the mesenteric or conglobate glands, which are very numerous in the human subject. As they run between the intestines and these glands, they are styled venæ lacteæ primis generis: but after leaving these glands, they are found to be less numerous, and being increased in size, are then called vene lacteæ secundi generis, which go to deposit their contents in the thoracic duct, through which the chyle is conveyed into the blood. This thoracic duct begins about the lower part of the first vertebra lumborum, from whence it passes up by the side of the aorta, between that and the vena azygos, close to the vertebræ, be- ing covered by the pleura. Sometimes it is found divided into branches; but they usually unite again into one canal, which opens into the left subclavian vein, after having run a little way in an oblique course between its coats. The sub- clavian vein communicates with the vena cava, which passes to the right auricle of the heart. The posed to be that of converting the chyle into blood. But the discovery of the thoracic duct by Pecquet, not long after, corrected this error. Pecquet very candidly confesses that this discovery accidentally arose from his observing a white fluid, mixed with the blood, flowing out of the vena cava, after he had cut off the heart of a living dog; which he suspected to be chyle, and afterwards traced to its source from the thoracic duct: this duct had been seen near an hundred years before in a horse by Eustachius, who speaks of it as a vein of a particular structure, but without know- ing any thing of its termination or use. 285 PART III. OF THE ABDOMEN. The lower part of this duct being usually larger than any other part of it, has been nam- ed receptaculum chyli, or Pecquet's receptacle, in honour of the anatomist who first discovered it in 1651. In some quadrupeds, in turtle and in fish, this enlargement * is more considerable in proportion to the size of the dud, than it usu- ally is in the human subject, where it is not com- monly found large enough to merit the name of receptaculum. Opportunities of observing the lacteals in the human subject do not often occur; but they may be easily demonstrated in a dog or any other quadruped that is killed two or three hours af- ter feeding upon milk; for then they appear fill- ed with white chyle. But these lacteals which we have described, as passing from the intestines through the mesen- tery to the thoracic duct, compose only a part of a system of vessels which perform the office of absorption, and which constitute, with their common trunk, the thoracic duct, and the conglobate glands that are dispersed through the body, what may be styled the lymphatic system. So that what is said of the structure of one of these series of vessels may very properly be ap- plied to that of the other. The lymphatics veins (K) are minute pellu- cid * Hewson's Exp. Inq. Part II. (K) The arteries in their course through the body becom- ing gradually too minute to admit the red globules of the blood, 286 PART III. OF THE ABDOMEN. cid tubes, which like the lacteals, direct their course towards the centre of the body, where they pour a colourless fluid into the thoracic duct. The lymphatics from all the lower parts of the body gradually unite as they approach this duct, into which they enter by three or four very large trunks, that seem to form the lower extremity of this canal, or receptaculum chyli, which may be considered as the great trunk of the lympha- tic system. The lacteals open into it near the same place; and the lymphatics, from a large share of the upper parts of the body, pour their lymph into different parts of this duct as it runs upwards, to terminate in the left subclavian vein. The lymphatics from the right side of the neck, thorax, and right arm, &c. terminate in the right subclavian vein. As the lymphatics commonly lie close to the large blood-vessels, a ligature passed round the crural blood, have then been styled capillary or lymphatic arteries. The vessels which are here described as constituting the lym- phatic system, were at first supposed to be continued from those arteries, and to convey back the lymph, either into the red veins or the thoracic duct; the office of absorption hav- ing been attributed to the red veins. But we know that the lymphatic veins are not continuations of the lymphatic arteries, but that they constitute the absorbent system. There are still, however, some very respectable names among the anatomists of the present age, who contend, that the red veins act like- wise as absorbents:—but it seems to have been clearly prov- ed, that the red veins do absorb no where but in the caver- nous cells of the penis, the erection of which is occasioned by a distention of those cells with arterial blood. 287 PART III. OF THE ABDOMEN. crural artery in a living animal, by including the lymphatics, will occasion a distention of these vessels below the ligature, so as to demonstrate them with ease; and a ligature passed round the thoracic duct, instantly after killing an animal, will, by stopping the course of its contents into the subclavian vein, distend not only the lacteals, but also the lymphatics in the abdomen and lower extremities, with their natural fluids (L). The coats of these vessels are too thin to be separated from each other; but the mercury they are capable of sustaining, proves them to be ve- ry strong; and their great power of contraction, after undergoing considerable distention, toge- ther with the irritability with which Baron Hal- ler found them to be endued *, seems to render it probable, that like the blood-vessels, they have a muscular coat. The lymphatics are nourished after the same manner as all the other parts of the body. For even the most minute of these vessels are pro- bably supplied with still more minute arteries and veins. This seems to be proved by the in- flammation of which they are susceptible; and the (L) In the dead body they may be easily demonstrated by opening the artery ramifying through any viscus, as in the spleen, for instance, and then throwing in air; by which the lymphatics will be distended. One of them may be punctured, and mercury introduced into it through a blow- pipe. * Sur le movement du fang Ex. 295, 298. 288 PART III. OF THE ABDOMEN. the painful swellings which sometimes take place in lymphatic vessels, prove that they have nerves as well as blood-vessels. Both the lacteals, lymphatics, and thoracic duct, are furnished with valves, which are much more common in these vessels than in the red veins. These valves are usually in pairs, and serve to promote the course of the chyle and lymph towards the thoracic duct, and to pre- vent its return. Mention has been made of the glands, through which the lacteals pass in their course through the mesentery; and it is to be ob- served, that the lymphatics pass through similar glands in their way to the thoracic duct. These glands are all of a conglobate kind, but the changes which the chyle and lymph undergo in their passage through them, have not yet been ascertained. The lymphatic vessels begin from surfaces and cavities in all parts of the body as absorbents. This is a fact now universally allowed; but how the fluids they absorb are poured into those cavities, is a subject of controversy. The con- tents of the abdomen, for instance, were de- scribed as being constantly moistened by a very thin watery fluid. The same thing takes place in the pericardium, pleura, and all the other ca- vities of the body, and this watery fluid is the lymph. But whether it is exhaled into those cavities through the minute ends of arteries, or transuded through their coats, are the points in 2 dispute. 289 PART III. OF THE ABDOMEN. dispute. We cannot here be permitted to re- late the many ingenious arguments that have been advanced in favour of each these opinions; nor is it perhaps of consequence to our present purpose to enter into the dispute. It will be suffi- cient if the reader can form an idea of what the lymph is, and of the manner in which it is ab- sorbed. The lymph, from its transparency and want of colour, would seem to be nothing but water; and hence the first discoverers of these vessels stiled them ductus aquosi but experiments prove, that the lymph of an healthy animal coagulates by being exposed to the air, or a certain degree of heat, and likewise by being suffered to rest ; seem- ing to agree in this property with that part of the blood called the coagulable lymph.—This proper- ty of the lymph leads to determine its use, in moistening and lubricating the several cavities of the body in which it is found; and for which, by its gelatinous principle, it seems to be much better calculated than a pure and watery fluid would be, for such it has been supposed to be by some anatomists. The mouths of the lymphatics and lacteals, by acting as capillary tubes, seem to absorb the lymph and chyle somewhat in the same manner as a capillary tube of glass, when put into a bason of water, is enabled to attract the wa- ter into it to a certain height, but it is pro- bable that they likewise possess a living power, T which 290 PART III. OF THE ABDOMEN. which assists in performing this office. In the human body the lymph or the chyle, is pro- bably conveyed upon this principle as far as the first pair of valves, which seem to be placed not far from the orifice of the absorbing vessel, whe- ther lymphatic or lacteal: and the fluid will then be propelled forwards, by a continuation of the absorption at the orifice. But this does not seem to be the only inducement to its progress to- wards the thoracic duct; these vessels have pro- bably a muscular coat, which may serve to press the fluid forwards from one pair of valves to ano- ther; and as the large lymphatic vessels and the thoracic duct are placed close to the large arteries, which have a considerable pulsa- tion, it is reasonable to suppose, that they de- rive some advantage from this situation. SECT. XV. Of the Generative Organs; of Con- ception, &c. § 1. The Male Organs. The male organs of generation have been usually divided into the parts which serve to pre- pare the semen from the blood, and those which are distended to convey it into the womb. But it seems to be more proper to distinguish them into the preparing, the containing, and the ex- pelling parts, which are the different offices of the testes, the vesiculæ seminales, and the penis; and this 251 PART III. OF THE ABDOMEN. this is the order in which we propose to describe them. The testes are two glandular bodies, serving to secrete the semen from the blood. They are originally formed and lodged within the cavity of the abdomen; and it is not till after the child is born, or very near that time, that they begin to pass into the groin, and from thence into the scrotum (M). By this disposition they are very wisely protected from the injuries to which they would be liable to be exposed, from the different positions of the child at the time of parturition. The testicles in this state are loosely attached to the psoæ muscles, by means of the peritonæ- um by which they are covered; and they are at this time of life connected in a very particular T2 manner (M) It sometimes happens in differing ruptures, that the intestine is found in the same sac, and in contact with the testis. This appearance was at first attributed to a supposed lacera- tion of the peritonæum; but later observations, by pointing out the situation of the testicles in the fœtus, have led to prove, the testis, as it descends into the scrotum, carries with it a portion or elongation of the peritonæum, which becomes its tunica vaginalis, or a kind of sac, in which the testicle is lodged, as will be explained in the course of this section. The communication between this sac and the cavity of the abdomen, is usually soon cut off; but in some subjects it con- tinues open during life; and when an hernia or descent of the intestine takes place in a such subject, it does not push down a portion of the peritonæum before it, as it must otherwise necessarily do, but passes at once through this opening, and comes in contact with the naked testicle, constituting that particular species of rupture called hernia congenita. 292 PART III. OF THE ABDOMEN. manner to the parietes of the abdomen, and like- wise to the scrotum, by means of a substance which Mr. Hunter calls the ligament or guberna- culum testis, because it connects the testis with the scrotum, and directs its course in its descent. This gubernaculum is of a pyramidal form, with its bulbous head fixed to the lower end of the testis and epididymis, and loses its lower and slender extremity in the cellular membrane of the scrotum. It is difficult to ascertain what the structure and composition of this gubernaculum is, but it is certainly vascular and fibrous; and, from certain circumstances, it would seem to be in part composed of the cremaster muscle, run- ning upwards to join the lower end of the testis. We are not to suppose that the testicle, when descended into the scrotum, is to be seen loose as a piece of gut or omentum would be in a com- mon hernial sac. We have already observed, that during its residence in the cavity of the ab- domen it is attached to the peritonæum, which descends with it; so that when the sac is com- pleted in the scrotum, the testicle is at first at- tached only to the posterior part of it, while the fore part of it lies loose, and for some time af- fords a communication with the abdomen. The spermatic chord, which is made up of the sper- matic artery and vein, and of the vas deferens or excretory duct of the testis, is closely attached behind to the posterior part of this elongation of the peritonæum. But the fore part of the peri- toneal 293 PART III. OF THE ABDOMEN. toneal sac, which is at first loose and not attached to the testicle, closes after a certain time, and be- comes united the posterior part, and thus per- fectly surrounds the testicle as it were in a purse. The testicles of the fœtus differ only in their size and situation from those of the adult. In their passage from the abdomen they descend through the abdominal rings into the scrotum, where they are supported and defended by vari- ous integuments. What the immediate cause of this descent is, has not yet been satisfactorily determined. It has been ascribed to the effects of respiration, but the testicles have sometimes been found in the scrotum before the child has breathed; and it does not seem to be occasioned by the action of the cremaster muscle, because the same effect would be liable to happen to the hedge-hog, and some other quadrupeds, whose testicles remain in the abdomen during life. The scrotum, which is the external or com- mon covering of both testicles, is a kind of sac formed by the common integuments, and exter- nally divided into two equal parts by a promi- nent line called raphe. In the inner part of the scrotum we meet with a cellular coat called dartos (N), which by its duplicature (N) The dartos has usually been considered as a muscle, and is described as such both by Douglas and Winslow. But there 294 PART III.OF THE ABDOMEN. duplicature divides the scrotum into two equal parts, and forms what is called septum scroti, which corresponds with the raphe. The collap- sion which is so often observed to take place in the scrotum of the healthy subject, when excit- ed by cold or by the stimulus of venery, seems to be very properly attributed to the contrac- tile motion of the skin, and not to any muscular fibres, as is the cafe in dogs and some other quad- rupeds. The scrotum, then, by means of its septum, is found to make two distinct bags, in which the testicles, invested by their proper tunics, are se- cured lodged and separated from each other. These coats are the cremaster, the tunica vagina- lis, and the tunica albuginea. The first of these is composed of muscular fibres, and is to be con- sidered only as a partial covering of the testis; for it surrounds only the spermatic chord, and terminates upon the upper and external parts of the tunica vaginalis testis, serving to draw up and suspend the testicle (O). The tunica vaginalis testis has already been described as being a thin production there being no part of the scrotum of the human subject which can be said to consist of muscular fibres, Albinus and Haller have very properly omitted to describe the dartos as a muscle, and consider it merely as a cellular coat. (O) The cremaster muscle is composed of a few fibres from the obliquus internus abdominis, which uniting with a few from the transversalis, descend upon the spermatic chord, and are insensibly lost upon the tunica vaginalis of the testi- cle. It serves to suspend and draw up the testicle. 295 PART III. OF THE ABDOMEN. production of the peritonæum, loosely adhering every where to the testicle, which it includes as it were in a bag. The tunica albuginea is a firm, white, and very compact membrane of a glis- tening appearance, which immediately invests the body of the testis and the epididymus; serv- ing in some measure to connect them to each other, but without extending itself at all to the spermatic chord. This tunica albuginea serves to confine the growth of the testis and epididy- mus within certain limits, and by giving them a due degree of firmness, enables them to per- form their proper functions. Having removed this last tunic, we discover the substance of the testicle itself, which appears to be made up of an infinite number of very elastic filaments, which may be best distinguish- ed after macerating the testicle in water. Each testicle is made up of the spermatic artery and vein, and the excretory vessels or tubuli semini- feri. There are likewise a great number of ab- sorbent vessels, and some branches of nerves to be met with in the testicles. The spermatic arteries arise one on each side from the aorta, generally about an inch below the emulgents. The right spermatic vein com- monly passes into the vena cava; but the left spermatic vein usually empties itself into the emulgent on that side; and it is supposed to take this course into the emulgent, that it may avoid passing 296 PART III. OF THE ABDOMEN. passing over the aorta, which it would be oblig- ed to do in its way to the vena cava. The blood is circulated very slowly through the spermatic artery, which makes an infinite number of circumvolutions in the substance of the testicle, where it deposites the semen, which passes through the tubuli seminiferi. These tu- buli seminiferi are seen running in short waves from the tunica albuginea to the axis of the testi- cle ; and are divided into distinct portions by certain thin membranous productions, which originate from the tunica albuginea. They at length unite, and by an infinite number of con- volutions form a sort of appendix to the testis called epididymis (P), which is a vascular body of an oblong shape, situated upon the superior part of each testicle. These tubuli of the epididymis at length form an excretory duct called vas de- ferens, which ascends towards the abdominal rings, with the other parts that make up the spermatic chord, and then a separation takes place; the nerves and blood-vessels passing on to their several terminations, and the vas deferens going to deposit its semen in the vesiculæ semi- nales, which are two soft bodies of a white and convoluted appearance externally, situated ob- liquely between the rectum and the lower part of (P) The testicles were named didymi by the ancients, and the name of this part was given to it on account of its situa- tion upon the testicle. 297 PART III. OF THE ABDOMEN. of the bladder, arid uniting together at the lower extremity. From these reservoirs (Q), which are (Q) That the bags called vesiculæ seminales are reservoirs of semen, is a circumstance which has been by anatomists universally believed. Mr. J. Hunter, however, from several circumstances, has been induced to think this opinion erro- neous. He has examined these vesiculæ in people who have died suddenly, and he found their contents to be different in their properties from the semen. In those who had lost one of the testicles, or the use of one of them, by disease, both the vesiculæ were full, and their contents similar. And in a lusus naturæ, where there was no communication between the vasa deferentia and vesiculæ, nor between the vesiculæ and penis, the same thing took place. From these observations, he thinks we have a presumptive proof, That the semen can be absorbed in the body of the testicle and in the epididymis, and that the vesiculæ secrete a mucus which they are capable of absorbing when it cannot be made use of: That the semen is not retained in reservoirs after it is secreted, and kept there till it is used; but that it is secreted at the time, in consequence of certain affections of them, and stimulating the testicles to this action. He corroborates his observations by the appearance on dissection in other animals; and here he finds, That the shape and contents of the vesiculæ vary much in different animals, while the semen in most of them he has examined is nearly the same: That the vasa deferentia in many animals do not communicate with the vesiculæ: That the contents of the vesiculæ of castrated and perfect animals are similar, and nearly equal in quantity, in no way resembling the semen as emitted from the animal in coitu, or what is found in the vas deferens after death. He observes likewise, that the bulb of the urethra of perfect males is considerably larger than in castrated animals. From the whole, he thinks the following inferences may be 298 PART III. OF THE ABDOMEN. are plentifully supplied with blood-vessels and nerves, the semen is occasionally discharged through two short passages, which open into the urethra be fairly drawn: That the bags called vesiculæ seminales are not seminal reservoirs, but glands secreting a peculiar mucus; and that the bulb of the urethra is properly speaking the receptacle of the semen, in which it is accumulated previous to ejection. But although he has endeavoured to prove that the vesi- culæ do not contain the semen, he has not been able to as- certain their particular use. He thinks, however, we may be allowed upon the whole to conclude, that they are, toge- ther with other parts, subservient to the purposes of gene- ration. Although the author has treated this subject very ably, and made many ingenious observations, some things may be objected to what he has advanced; of which the following are a few: That those animals who have bags called vesiculæ seminales perform copulation quickly; whereas others that want them, as in the dog kind, are tedious in copulation: That in the human body, at least, there is a free communica- tion between the vasa deferentia and vesiculæ; and in ani- mals where the author has observed no communication be- tween the vasa deferentia and vesiculæ, there may be a com- munication by vessels not yet discovered, and which may be compared to the hepato-cystic ducts in fowls and fishes: That the fluid in the end of the vasa deferentia and the vesi- culæ seminales are similar, according to the author's own ob- servation: That the vesiculæ in some animals increase and decrease with the testicle at particular seasons: That in birds and certain fishes, there is a dilatation of the ends of the vasa deferentia, which the author himself allows to be a reser- voir for the semen. With respect to the circumstance of the bulb of the ure- thra answering the purpose of a reservoir, the author has mention no facts which tend to establish this opinion. See Observations on certain Parts of the Animal Oeconomy. 299 PART III. OF THE ABDOMEN. urethra close to a little eminence called verumon- tanum. Near this eminence we meet with the pros- tate, which is situated at the neck of the bladder, and is described as being of a glandu- lar structure. It is shaped somewhat like a heart with its small end foremost, and inverts the ori- gin of the urethra. Internally it appears to be of a firm substance, and composed of several fol- licles, secreting a whitish viscid fluid, that is dis- charged by ten or twelve excretory ducts into the urethra, on each side of the openings of the vesiculæ seminales at the same time, and from the same causes that the semen is expelled. As this latter fluid is found to be exceedingly limpid in the vesiculæ seminales of the dead subject, it probably owes its whiteness and viscidity to this liquor of the prostate. The penis, which is to be considered as the vehicle or active organ of procreation, is com- posed of two columns, the corpora cavernosa, corpus spongiosum. The corpora cavernosa, which constitute the greatest part of the penis, may be described as two cylindrical ligamentous tubes, each of which is composed of an infinite number of minute cells of a spongy texture, which communicate with each other. These two bodies are of a very pliant texture, and ca- pable of considerable distention: and being unit- ed laterally to each other, occasion by this union a space above and another below. The upper- and 300 PART III. OF THE ABDOMEN. most of these spaces is filled by the blood vessels, and the lower one, which is larger than the other, by the urethra and its corpus spongiosum. These two cavernous bodies are at first only se- parated by a partition of tendinous fibres, which allow them to communicate with each other; but they afterwards divaricate from each other like the branches of the letter Y, and diminishing gradually in size, are attached, one on each side, by means of the ligamentum suspensorium penis to the ramus ischii, and to the inferior portion of the os pubis. The corpus spongiosum penis, or corpus spon- giosum urethræ, as it is styled by some authors, begins as soon as the urethra has passed the pros- tate, with a thick origin almost like a heart, first under the urethra, and afterwards above it, be- coming gradually thinner, and surrounding the whole canal of the urethra, till it terminates in a considerable expansion, and constitutes what is called the glans penis, which is exceedingly vas- cular, and covered with papillæ like the tongue. The cuticle which lines the inner surface of the urethra, is continued over the glans in the same manner as it is spread over the lips. The penis is invested by the common integu- ments, but the cutis is reflected back every where from the glans as it is in the eye-lids; so that it covers this part, when the penis is in a relaxed state, as it were with a hood, and from this use is called prepuce. The 301 PART III. OF THE ABDOMEN. The prepuce is tied down to the under part of the glans by a small ligament called frænum, which is in fact only a continuation of the cuticle and cutis. There are many simple sebaceous follicles called glandulæ odoriferæ, placed round the basis of the glans; and the fluid they secrete serves to preserve the exquisite sensibility of this part of the penis, and to prevent the ill effects of attrition from the prepuce. The urethra may be defined to be a membran- ous canal, passing from the bladder through the whole extent of the penis. Several very small openings, called lacunæ, communicate with this canal, through which a mucus is discharged into it; and besides these, there are two glands, first described by Cowper, as secreting a fluid for lu- bricating the urethra, and called Cowper's glands (R); and Littre * speaks of a gland situated near the prostate, as being destined for the same use. The urethra being continued from the neck of the bladder, is to be considered as making part of the urinary passage; and it likewise affords a conveyance to the semen, which we have observ- ed is occasionally discharged into it from the ve- siculæ seminales. The direction of this canal being first under and then before the pubis, oc- casions (R) Both Heister and Morgagni observe, that they have sometimes not been able to find these glands; so that they do not seem to exist in all subjects. * Memories de l' Acad. Royale des Sciences, 1700. 302 PART III. OF THE ABDOMEN. casions a winding in its course from the blad- der to the penis not unlike the turns of the let- ter S. The penis has three pair of muscles, the erec- tores, acceleratores, and transversales. They push the blood from the crura to the fore part of the corpora cavernosa. The first originate from the tuberosity of the ischium, and terminate in the corpora cavernosa. The acceleratores arise from the sphincter, and by their insertion serve to compress the bulbous part of the urethra; and the transversales are destined to afford a pas- sage to the semen, by dilating the canal of the urethra. The arteries of the penis are chiefly derived from the internal iliacs. Some of them are sup- posed to terminate by pabulous orifices within the corpora cavernosa and corpus spongiosum; and others terminate in veins, which at last make up the vena magna dorsi penis, and other small- er veins, which are in general distributed in like order with the arteries. Its nerves are large and numerous. They rise from the great sciatic nerve, and accom- pany the arteries in their course through the penis. We have now described the anatomy of this organ; and there only remains to be explained, how it is enabled to attain that degree of firmness and distension which is essential to the great work of generation. The 303 PART III. OF THE ABDOMEN. The greatest part of the penis has been spoken of as being of a spongy and cellular texture, plentifully supplied with blood-vessels and nerves, and as having muscles to move it in different di- rections. Now, the blood is constantly passing into its cells through the small branches of the arteries which which open into them, and is from thence as constantly returned by the veins, so long as the corpora cavernosa and corpus spon- giosum continue to be in a relaxed and pliant state. But when, from any nervous influence, or other means, which it is not necessary here to define or explain, the erectores penis, ejacu- latores seminis, levatores ani, &c. are induced to contract, the veins undergo a certain degree of compression, and the passage of the blood through them is so much impeded, that it col- lects in them in a greater proportion than they are enabled to carry off, so that the penis gradu- ally enlarges; and being more and more forcibly drawn up against the os pubis, the vena magna itself is at length compressed, and the penis be- comes fully distended. But as the causes which first occasioned this distension subside, the penis gradually returns to its state of relaxation. § 2. Female Organs of Generation. Anatomical writers usually divide the fe- male organs of generation into external and inter- nal. In the first division they include the mons 3 veneris, 304 PART III. OF THE ABDOMEN. veneris, labia pudendi, perinæum, clitoris, nymphæ, and canunculæ myrtiformes; and in the latter, the vagina; with the uterus and its appendages. The mons veneris, which is placed on the upper part of the symphysis pubis, is internally com- posed of adipose membranes, which makes it soft and prominent: it divides into two parts called labia pudendi, which descending towards the rectum, from which they are divided by the perinæeum, form what is called the fourchette. The perinæum is that fleshy space which extends about an inch and an half from the fourchette to the anus, and from, thence about two inches to the coccyx. The labia pudendi being separated, we observe a sulcus called fossa magni; in the upper part of which is placed the clitoris, a small round spongy body, in some measure resembling the male pe- nis, but impervious, composed of two corpora cavernosa arising from the tuberosities of the ossa ischii; furnished with two pair of muscles, the erectores clitoridis, and the sphincter or con- strictor ostii vaginæ; and terminating in a glans, which is covered with its prepuce. From the lower part, on each side of the fossa, pass the nymphæ, two membranous and spongy folds which seem destined for useful purposes in partu- rition, by tending to enlarge the volume of the vagina as the child's head passes through it. Be- tween these, about the middle of the fossa mag- na, we perceive the orifice of the vagina or os ex- 2 ternum, 305 PART III. OF THE ABDOMEN. ternum, closed by folds and wrinkles; and about half an inch above this, and about an inch be- low the clitoris, appears the meatus urinarius or orifice of the urethra, much shorter, though somewhat larger, than in men, with a little pro- minence at its lower edge, which facilitates the introduction of the catheter. The os externum is surrounded internally by several membranous folds called carunculæ myr- tiformes, which are partly the remains of a thin membrane called hymen, that covers the vagina in children. In general the hymen is sufficiently open to admit the passage of the menses, if it exists at the time of their appearance; some- times, however, it has been found perfectly closed. The vagina, situated between the urethra and the rectum, is a membranous cavity, surround- ed especially at its external extremity with a spongy and vascular substance, which is covered by the sphincter ostii vaginæ. It terminates in the uterus, about half an inch above the os tin- cæ, and is wider and shorter in women who have had children than in virgins. All these parts are plentifully supplied with blood-vessels and nerves. Around the nymphæ there are sebaceous follicles, which pour out a fluid to lubricate the inner surface of the vagina; and the meatus urinarius, like the urethra in the male subject, is constantly moistened by a mu- U cus, 306 PART III. OF THE ABDOMEN. cus, which defends it against the acrimony of the urine. The uterus is a hollow viscus, situated in the hypogastric region, between the rectum and blad- der. It is destined to receive the first rudiments of the fœtus, and to assist in the developement of all its parts, till it arrives at a state of per- fection, and is fitted to enter into the world, at the time appointed by the wise Author of nature. The uterus, in its unimpregnated state, resem- bles a pear in shape, somewhat flattened, with its fundus or bottom part turned towards the ab- domen, and its cervix or neck surrounded by the vagina. The entrance into its cavity forms a little protuberance, which has been compared to the mouth of a tench, and is therefore called os tinæ. The substance of the uterus, which, is of a considerable thickness, appears to be composed of muscular and small ligamentous fibres, small branches of nerves, some lymphatics, and with arteries and veins innumerable. Its nerves are chiefly derived from the intercostal, and its arte- ries and veins from the hypogastric and spermatic. The membrane which lines its cervix, is a con- tinuation of the inner membrane of the vagina; but the outer surface of the body of the uterus is covered with the peritonæum, which is re- flected over it, and descends from thence to the intestinum rectum. This duplicature of the peritonæum, 307 PART III. OF THE ABDOMEN. peritonæum, by passing off from the sides of the uterus to the sides of the pelvis, is there firmly connected, and forms what are called ligamenta uteri lata; which not only serve to support the uterus, but to convey nerves and blood-vessels to it. The ligamenta uteri rotunda arise from the sides of the fundus uteri, and passing along with- in the fore-part of the ligamenta lata, descend through the abdominal rings, and terminate in the substance of the mons veneris. The sub- stance of these ligaments is vascular, and al- though both they and the ligamenta lata admit the uterus in the virgin state, to move only about an inch up and down, yet in the course of preg- nancy they admit of considerable distension, and after parturition return nearly to their original state with surprising quickness. On each side of the inner surface of the ute- rus, in the angle near the fundus, a small orifice is to be discovered, which is the beginning of one of the tubæ fallopianæ. Each of these tubes, which are two in number, passing through the substance of the uterus, is extended along the broad ligaments, till it reaches the edge of the pelvis, from whence it reflects back; and turn- ing over behind the ligaments, about an inch of its extremity is seen hanging loose in the pelvis, near the ovarium. These extremities, having a jagged appearance, are called fimbriæ, or morsus diaboli. Each tuba Fallopiana is usually about U2 three 308 PART III. OF THE ABDOMEN. three or four inches long. Their cavities are at first very small, but become gradually larger, like a trumpet, as they approach the fimbriæ. Near the fimbriæ of each tuba Fallopiana, about an inch from the uterus, is situated an oval body called ovarium, of about half the size of the male testicle. Each of these ovaria is cover- ed by a production of the peritonæum, and hangs loose in the pelvis. They are of a flat and angular form, and appear to be composed of a white and cellular substance, in which we are able to discover several minute vesicles filled with a coagulable lymph, of an uncertain num- ber, commonly exceeding 12 in each ovary. In the female of riper years, these vesicles become exceedingly turgid, and a kind of yellow coa- gulum is gradually formed within one of them, which increases for a certain time. In concep- tion, one of these mature ova is supposed to be impregnated with the male semen, and to be squeezed out of its nidus into the Fallopian tube; after which the ruptured part forms a substance which in some animals is of a yel- low colour, and is therefore called corpus lute- um; and it is observable, that the number of these tears or fissures in the ovarium, constantly corresponds with the number of fœtuses exclud- ed by the mother. § 3. Of 309 PART III. OF THE ABDOMEN. § 3. Of Conception. Man, being ever curious and inquisitive, has naturally been led to enquire after the origin of his existence; and the subject of generation has employed the philosophical world in all ages: but in following nature up to her minute re- cesses, the philosopher soon finds himself bewil- dered, and his imagination often supples that which he so eagerly wishes to discover, but which is destined perhaps never to be revealed to him. Of the many theories which have been formed on this subject, that of the ancient philosophers seems to have been the most simple: they con- sidered the male semen as alone capable of form- ing the fœtus, and believed that the female only afforded it a lodging in the womb, and supplied it with a nourishment after it was perfectly form- ed. This opinion, however, soon gave place to another, in which the female was allowed a more considerable share in conception. This second system considered the fœtus as being formed by the mixture of the seminal li- quor of both sexes, by a certain arrangement of its several particles in the uterus. But in the 16th century, vesicles or eggs were discovered in the ovaria or female testicles; the fœtus had been found sometimes in the abdomen, and sometimes in the Fallopian tubes; and the two former opinions were exploded in favour of a new 210 PART III. OF THE ABDOMEN. new doctrine. The ovaria were compared to a bunch of grapes, being supposed to consist of ve- sicles, each of which had a stalk; so that it might be disengaged without hurting the rest, or spill- ing the liquor it contained. Each vesicle was said to include a little animal, almost complete in all its parts; and the vapour of the male se- men being conveyed to the ovarium, was sup- posed to produce a fermentation in the vesicle, which approached the nearest to maturity; and inducing it to disengage itself from the ovarium, it passed into the tuba Fallopiana, through which it was conveyed to the uterus. Here it was supposed to take root like a vegetable seed, and to form, with the vessels originating from the uterus, what is called the placenta; by means of which the circulation is carried on between the mother and the fœtus. This opinion, with all its absurdities, conti- nued to be almost universally adopted till the close of the same century, when Lieuwenhoeck, by means of his glasses, discovered certain opake particles, which he described as so many animalcula, floating in the seminal fluid of the male. This discovery introduced a new schism among the philosophers of that time, and gave rise to a system which is not yet entirely exploded. According to this theory the male semen passing into the tubæ Fallopianæ, one of the animalcula penetrates into the substance of the ovarium, and 211 Part III. OF THE ABDOMEN. and enters into one of its vesicles or ova. This impregnated ovum is then squeezed from its husk, through the coats of the ovarium, and be- ing seized by the fimbriæ, is conducted through the tube to the uterus, where it is nourished till it arrives at a state of perfection. In this sys- tem there is much ingenuity; but there are cer- tain circumstances supposed to take place, which have been hitherto inexplicable. A celebrated modern writer, M. Buffon, endeavours to re- store, in some measure, the most ancient opi- nion, by allowing the female semen a share in this office; asserting, that animalcula or organic particles are to be discovered in the seminal li- quor of both sexes: he derives the female semen from the ovaria, and he contends that no ovum exists in those parts. But in this idea he is evi- dently mistaken; and the opinion now most ge- nerally adopted is, that an impregnation of the ovum, by the influence of the male semen, is essential to conception (S). That the ovum is to be impregnated, there can be no doubt; but as the manner in which such an impregnation is supposed to take place, and the means by which the ovum afterwards get into the Fallopian tube, and (S) The learned Abbe Spallanzani has thrown much light on this curious subject, and has proved by a variety of ex- periments than the animalcule exists entire in the female ovum, and that the male seed is only necessary to vivify and put it in motion.—His experiments and observations are worthy of the attentive perusal of every physiologist. 212 PART III. OF THE ABDOMEN. and from thence into the uterus, are still found- ed chiefly on hypothesis, we will not attempt to extend farther the investigation of a subject con- cerning which so little can be advanced with cer- tainty. § 4. Of the Fœtus in Utero. Opportunities of dissecting the human gravid uterus occurring but seldom, the state of the embryo (T) immediately after conception cannot be perfectly known. When the ovum descends into the uterus, it is supposed to be very minute; and it is not till a considerable time after conception that the rudiments of the embryo begin to be ascer- tained. About the third or fourth week the eye may discover the first lineaments of the fœtus; but these lineaments are as yet very imperfect, it be- ing only about the size of a house-fly. Two little vessels appear in an almost transparent jel- ly; the largest of which is destined to become the head of the fœtus, and the other smaller one is reserved for the trunk. But at this period no extremities are to be seen; the umbilical cord appears only as a very minute thread, and the placenta (T) The rudiments of the child are usually distinguished by this name till the human figure can be distinctly ascertained, and then it has the appellation of fœtus. 213 PART III. OF THE ABDOMEN. placenta does not yet absorb the red particles of the blood. At six weeks, not only the head but the features of the face begin to be developed. The nose appears like a small prominent line, and we are able to discover another line under it, which is destined for the separation of the lips. Two black points appear in the place of eyes, and two minute holes mark the ears. At the sides of the trunk, both above and below, we see four minute protuberances, which are the the rudiments of the arms and legs. At the end of eight weeks the body of the fœtus is upwards of an inch in length, and both the hands and feet are to be distinguished. The upper extre- mities are found to increase faster than the lower ones, and the separation of the fingers is accom- plished sooner than that of the toes. At this period the human form may be deci- sively ascertained;—all the parts of the face may be distinguished, the shape of the body is clearly marked out, the haunches and the abdomen are elevated, the fingers and toes are separated from each other, and the intestines appear like minute threads. At the end of the third month, the fœtus mea- sures about three inches; at the end of the fourth month, five inches; in the fifth month, six or seven inches; in the sixth month, eight or nine inches; in the seventh month, eleven or twelve inches; in the eighth month, fourteen or fif- teen inches; and at the end of the ninth month, or 214 PART III. OF THE ABDOMEN. or full time, from eighteen to twenty-two in- ches. But as we have not an opportunity of ex- amining the same fœtus at different periods of pregnancy, and as their size and length may be influenced by the constitution and mode of life of the mother, calculations of this kind must be very uncertain. The fœtus during all this time assumes an oval figure, which corresponds with the shape of the uterus. Its chin is found reclining on its breast with its knees drawn up towards its chin, and its arms folded over them. But it seems likely, that the posture of some of these parts is varied in the latter months of pregnancy, so as to cause those painful twitches which its mother usually feels from time to time. In natural cases, its head is probably placed towards the os tincæ from the time of conception to that of its birth; though formerly it was considered as being plac- ed towards the fundis uteri till about the eighth or ninth mouth, when the head, by becom- ing specifically heavier than the other parts of the body, was supposed to be turned down- wards. The capacity of the uterus increases in pro- portion to the growth of the fœtus, but without becoming thinner in its substance, as might na- turally be expected. The nourishment of the fœtus, during all this time, seems to be derived from the placenta, which appears to be original- ly formed by that part of the ovum which is next 215 Part III. OF THE ABDOMEN. next the fundis uteri. The remaing part of the ovum is covered by a membrane called spongy chorion (U); within which is another called true chorion, which includes a third termed amnios (V): this contains a watery fluid, which is the liquor amnii (W), in which the fœtus floats till the time of its birth. On the side next the fœtus, the placenta is covered by the amnios and true cho- rion; on the fide next the mother it has a pro- duction continued from the spongy chorion. The (U) Dr. Hunter has described this as a lamella from the inner surface of the uterus. In the latter months of preg- nancy it becomes gradually thinner and more connected with the chorion : he has named it menbrana caduca, or decidua, as it is cast off with the placenta. Signior Scarpa, with more probability, considers it as being composed of an in- spissated coagulable lymph. (V) In some quadrupeds, the urine appears to be con- veyed from the bladder through a canal called urachus, to the alantois, which is a reservoir, resembling a large and blind gut, situated between the chorion and amnios. The human fœtus seems to have no such reservoir, though some writers hare supposed that it does exist. From the top of the bladder a few longituddinal fibres are extended to the umbilical chord; and these fibres have been considered as the urachus, though without having been ever found pervious. (W) The liquor amnii coagulates like the lymph. It has been supposed to pass into the œsophagus, and to afford nou- rishment to the fœtus; but this does not seem probable. Children have come into the world without an œsophagus, or any communication between the stomach and the mouth; but there has been no well attested instance of a child's hav- ing been born without a placenta; and it does not seem like- ly, that any of the fluid can be absorbed through the pores of the skin, the skin in the fœtus being every where covered with a great quantity of mucus. 216 PART III. OF THE ABDOMEN. The amnios and chorion are remarkably thin and transparent, having no blood-vessels entering in- to their composition. The spongy chorion is opake and vascular. In the first months of pregnancy, the involu- cra bear a large proportion to their contents; but this proportion is afterwards reversed, as the fœ- tus increases in bulk. The placenta, which is the medium through which the blood is conveyed from the mother to the fœtus, and the manner in which this conveyance takes place, deserve next to be con- sidered. The placenta is a broad, flat, and spongy sub- stance, like a cake, closely adhering to the inner surface of the womb, usually near the fundus, and appearing to be chiefly made up of the rami- fications of the umbilical arteries and vein, and partly of the extremities of the uterine vessels. The arteries of the uterus discharge their con- tents into the substance of this cake; and the veins of the placenta, receiving the blood either by a direct communication of vessels, or by ab- sorption, at length form the umbilical vein, which passes on to the sinus of the vena porta, and from thence to the vena cava, by means of the canalis venosus, a communication that is closed in the adult. But the circulation of the blood through the heart is not conducted in the fœtus as in the adult: in the latter the blood is carried from the right auricle of the heart through the 217 PART III. OF THE ABDOMEN. the pulmonary artery, and is returned to the left auricle by the pulmonary vein; but a dilatation of the lungs is essential to the passage of the blood through the pulmonary vessels, and this dilatation cannot take place till after the child is born and has respired. This deficiency, howe- ver, is supplied in the fœtus by the immediate communication between the right and left auri- cle, through an oval opening, in the septum which divides the two auricles, called foramen ovale. The blood is likewise transmitted from the pulmonary artery to the aorta, by means of a duct called canalis arteriosus, which, like the canalis venosus, and foramen ovale, gradually closes after birth. The blood is returned again from the fœtus through two arteries called the umbilical arteries, which arise from the iliacs. These two vessels taking a winding course with the vein, form with that, and the membranes by which they are surrounded, what is called the umbilical chord. These arteries, after ramifying through the sub- stance of the placenta, discharge their blood into the veins of the uterus; in the same manner as the uterine arteries discharged their blood into the branches of the umbilical vein. So that the blood is constantly passing in at one side of the placenta and out at the other; but in what par- ticular manner it gets through the placenta is a point not yet determined. 2 EXPLA- 218 PART III. OF THE ABDOMEN. EXPLANATION of PLATES XXV. XXVI. AND XXVII. PLATE XXV. FIG. 1. Shows the Contents of the Thorax and Abdomen in situ. 1. Top of the trachea, or wind-pipe. 22, The internal jugular veins. 33, The subcla- vian veins. 4, The vena cava descendens. 5, The right auricle of the heart. 6, The right ventricle. 7, Part of the left ventricle. 8, The aorta descendens. 9, The pulmonary artery. 10, The right lung, part of which is cut off to show the great blood-vessels. 11, The left lung, entire. 1212, The anterior edge of the dia- phragm. 1313, The two great lobes of the liver. 14, The ligamentum rotundum. 15, The gall-bladder. 16, The stomach. 1717, The jejunum and ilium. 18, The spleen. FIG. 2. Shows the Organs subservient to the Chylopoietic Viscera,—with those of the U- rine and Generation. 11, The under side of the two great lobes of the liver. a, Lobulus Spigelii. 2, The li- gamentum rotundum. 3, The gall-bladder. 4, The pancreas, 5, The spleen. 66, The kidneys. 7, The aorta descendens. 8, Vena cava 319 PART III. OF THE ABDOMEN. cava ascendens. 99, The renal veins covering the arteries. 10, A probe under the spermatic vessels and a bit of the inferior mesenteric artery, and over the ureters. 1111, The ureters. 12 12, The iliac arteries and veins. 13, The rectum intestinum. 14, The bladder of urine. FIG 3. Shows the Chylopoietic Viscera, and Organs subservient to them, taken out of the Body entire. AA the under side of the two great lobes of the liver. B, Ligamentum rotundum. C, The gall-bladder. D, Ductus cysticus. E, Ductus hepaticus. F, Ductis communis choledochus. G, Vena portarum. H, Arteria hepatica. II, The stomach. KK, Venæ & arteriæ gastro- epiploicæ, dextræ & sinistræ. LL, Venæ & arteriæ coronariæ ventriculi. M, The spleen. N N, Mesocolon, with its vessels. OOO, In- testinum colon. P, One of the ligaments of the colon, which is a bundle of longitudinal muscular fibres. QQQQ, Jejunum and ilium. R R, Sigmoid flexure of the colon, with the ligament continued, and over S, The rec- tum intestinum. TT, Levatores ani. U, Sphincter ani. V, The place to which the pro- state glandis connected. W, The anus. FIG. 4. 320 PART III. OF THE ABDOMEN. FIG 4. Shows the heart of a Fœtus at the full time, with the Right Auricle cut open to show the Foramen Ovale, or passage between both Auricles. a, The right ventricle, b, The left ventricle. cc, The outer side of the right auricle stretched out. d d, The posterior side, which, forms the anterior side of the septum. e, The foramen ovale, which the membrane or valve which co- vers the left side. f, Vena cava inferior passing through g, A portion of the diaphragm. FIG. 5. Shows the Heart and Large Vessels of a Fœtus at the full time. a, The left ventricle. b, The right ventri- cle. c, A part of the right auricle. d, Left auricle, ee, The right branch of the pulmona- ry artery. f, Arteria pulmonalis. g g, The left branch of the pulmonary artery, with a num- ber of its largest branches dissected; from the lungs. h, The canalis arteriosus. i, The arch of the aorta. kk, The aorta descendens. l, The left subclavian artery. m, The left carotid ar- tery. n, The right carotid artery. o. The right subclavian artery. p, The origin of the right carotid and right subclavian arteries in one common trunk. q, The vena cava superior or descendens. r, The right common subclavian vein. S, The left common subclavian vein. N. B. All the parts described in this figure are to be found in the adult, except the canalis arteriosus. PLATE ANATOMY PLATE XXV  321 PART III. OF THE ABDOMEN. PLATE XXVI. FIG. 1. Exhibits the more superficial Lympha- tic Vessels of the Lower Extremity. A, The spine of the os ilium. B, The os pubis. C, The iliac artery. D, The knee, EEF, Branches of the crural artery. G, The muscu- lus gastrocnemius. H, The tibia. I, The ten- don of the musculus tibialis anticus. On the out-lines, a, A lymphatic vessel belonging to the top of the foot. b, Its first division into branches. c,c,c, Other divisions of the same lymphatic vessel. d, A small lymphatic gland. c, The lymphatic vessels which lie between the skin and the muscles of the thigh, ff, Two lymphatic glands at the upper part of the thigh below the groin. g g, Other glands. h, A lymphatic vessel which passes by the side of those glands without communicating with them ; and, bending towards the inside of the groin at (i), opens into the lymphatic gland (k). l, l, Lym- phatic glands in the groin, which are common to the lymphatic vessels of the genitals and those of the lower extremity, m, n, A plexus of lymphatic vessels passing on the inside of the iliac artery. X FIG. 2. 322 PART III. OF THE ABDOMEN. FIG. 2. Exhibits a Back View of the Lower Ex- tremity, dissected so as to show the deeper- seated Lymphatic Vessels which accompany the arteries. A, The os pubis. B, The tuberosity of the ischium. C, That part of the os ilium which was articulated with the os sacrum. D, The extremity of the iliac artery appearing above the groin. E, The knee. FF, The two cut surfaces of the triceps muscle, which was divid- ed to show the lymphatic vessels that pass through its perforation along with the crural artery. G, The edge of the musculus gracilis. H, The gas- troncnemius and soleus, much shrunk by being dried, and by the soleus being separated from the tibia to expose the vessels. I, The heel. K, The sole of the foot. L, The superficial lymphatic vessels passing over the knee, to get to the thigh. On the out-lines; M, The poste- rior tibial artery. a, A lymphatic vessel accom- panying the posterior tibial artery, b, The same vessel crossing the artery, c, A small lymphatic gland, through which this deep-seated lympha- tic vessel passes. d, The lymphatic vessel pas- sing under a small part of the soleus, which is left attached to the bone, the rest being re- moved. e, The lymphatic vessel crossing the popliteal artery. f, g, h, Lymphatic glands in the ham, through which the lymphatic vessel passes. i, The lymphatic vessel passing with the 323 PART III. OF THE ABDOMEN. the crural artery, through the perforation of the triceps muscle. k, The lymphatic vessel, af- ter it had passed the perforation of the triceps, dividing into branches which embrace the ar- tery (l). m, A lymphatic gland belonging to the deep-seated lymphatic vessel. At this place those vessels pass to the fore-part of the groin, where they communicate with the super- ficial lymphatic vessel appearing on the brim of the pelvis. FIG. 3. Exhibits the Trunk of the Human Sub- ject prepared to show the Lymphatic Vessels and the Ductus Thoracicus. A, The neck. B B, The two jugular veins. C, The vena cava superior. DDDD, The subclavian veins. E, The beginning of the aorta, pulled to the left side by means of a liga- ture, in order to show the thoracic duct behind it. F, The branches arising from the curvature of the aorta. GG, The two carotid arteries. HH, the first ribs, II, The trachea. KK, The spine. L, The vena azygos. MM, The descending aorta. N, The cœliac artery, dividing into three branches. O, The superior mesenteric artery. P, The right crus diaphrag- matis. QQ, The two kidneys. R, The right emulgent artery. SS, The external iliac arte- ries, g d, The musculi psoæ. T, The inter- nal iliac artery. U, The cavity of the pelvis. X2 XX, The 324 PART III. OF THE ABDOMEN. XX, The spine of the os ilium. YY, The groins. a, A lymphatic gland in the groin, in- to which lymphatic vessels from the lower ex- tremity are seen to enter. b b, The lymphatic vessels of the lower extremities passing under Poupart's ligament. cc, A plexus of the lym- phatic vessels lying on each side of the pelvis. d, The psoas muscle with lymphatic vessels lying upon its inside. e, A plexus of lymphatics, which having passed over the brim of the pel- vis at (c), having entered the cavity of the pel- vis, and received the lymphatic vessels belong- ing to the viscera contained in that cavity, next ascends, and passes behind the iliac artery to (g). f, some lymphatic vessels of the left side pas- ssing over the upper part of the os sacrum, to meet those of the right side. g, The right psoas, with a large plexus of lymphatics lying on its inside. hh, The plexus lying on each side of the spine. iii, Spaces occupied by the lym- phatic glands. k, The trunk of the lacteals, lying on the under side of the superior mesente- ric artery. l, The same dividing into two branches, one of which passes on each side of the aorta ; that of the right side being seen to enter the thoracic duct at (m). m, The thora- cic beginning from the large lymphatics. n, The duct passing under the lower part of the crus diaphragmatis, and under the right emulgent ar- tery. o, The thoracic duct penetrating the thorax. p, Some lymphatic vessels joining that duct 325 PART III. OF THE ABDOMEN. duct in the thorax. q, The thoracic duct pas- sing under the curvature of the aorta to get to the left subclavian vein. The aorta being drawn aside to show the duct. r, A plexus of lym- phatic vessels passing upon the trachea from the thyroid gland to the thoracic duct. PLATE XXVII. FIG 1. Represents the Under and Posterior Side of the Bladder of Urine, &c. a, The bladder. b b, The insertion of the uterers, cc, The vasa deferentia, which con- vey the semen from the testicles to d d, The ve- siculæ seminales,—and pass through e, The pro- state gland, to discharge themselves into f, The beginning of the urethra. FIG 2. A transverse Section of the Penis. g g, Corpora carvernosa penis. h, Corpus cavernosum urethræ. i, Urethra. k, Septum penis. ll, The septum between the corpus ca- vernosum urethræ and that of the penis. FIG. 3. A longitudinal Section of the Penis mm, The corpora carvernosa penis, divided by o, The septum penis, n, The corpus ca- vernosum glandis, which is the continuation of that of the urethra. FIG. 4. 326 PART III. OF THE ABDOMEN. FIG. 4. Represents the Female Organs of Ge- neration. a, That side of the uterus which is next the os sacrum. 1, Its fundus. 2, Its cervix. b b, The Fallopian or uterine tubes, which open in- to the cavity of the uterus;—but the other end is open within the pelvis, and surrounded by cc, The fimbriæ. d d, The ovaria. e, The os internum uteri, or mouth of the womb. ff, The ligamenta rotunda, which passes without the belly, and is fixed to the labia pudendi. gg, The cut edges of the ligamenta lata which connects the uterus to the pelvis. h, The inside of the vagina. i, The orifice of the urethra. k, The clitoris surrounded by (l,) The præputium. mm, The labia pudendi. nn, The nymphæ. FIG. 5. Shows the Spermatic Ducts of the Tes- ticle filled with Mercury. A, The vas deferens. B, Its beginning, which forms the posterior part of the epididy- mis. C, The middle of the epididymis, com- posed of serpentine ducts. D, The head or an- terior part of the epididymis unravelled, eeee, The whole ducts which compose the head of the epiddiymis unravelled. ff, The vasa defe- rentia. gg, Rete testis. hh, Some rectilineal ducts ANATOMY Plate XXV  ANATOMY Plate XXVII  327 PART III. OF THE ABDOMEN. ducts which send off the vasa deferentia. ii, The substance of the testicle. FIG. 6. The right Testicle entire, and the Epi- didymis filled with Mercury. A, The beginning of the vas deferens. B, The vas deferens ascending towards the abdomen. C, The posterior part of the epididymis, named globus minor. D, The spermatic vessels inclosed in cellular substance. E, The body of the epidy- dimis. F, Its head, named globus major. G, Its beginnning from the testicle. H, The body of the testicle, inclosed in the tunica albuginea. PART IV. 328 PART IV. OF THE THORAX. PART IV. OF THE THORAX. THE thorax, or chest, is that cavity of the trunk which extends from the cla- vicles, or the lower part of the neck, to the dia- phragm, and includes the vital organs, which are the heart and lungs; and likewise the tra- chea and œsophagus.—This cavity is formed by the ribs and vertebræ of the back, covered by a great number of muscles, and by the common integuments, and anteriorly by two glandular bodies called the breasts. The spaces between the ribs are filled by the muscular fibres, which from their situation are called intercostal muscles. SECT. I. Of the Breasts. The breasts may be defined to be two large conglomerate glands, mixed with a good deal of adipose membrane. The glandular part is com- posed of an infinite number of minute arteries, veins, and nerves. The arteries are derived from two different trunks; one of which is called the internal, and the other the external, mammary artery. The first of these arises from the subclavian, and the latter from the axillary, The 329 Part IV. OF THE THORAX. The veins every where accompany the arte- ries, and are distinguished by the same name. The nerves are chiefly from the vertebral pairs. Like all other conglomerate glands, the breasts are made up of a great many small distinct glands, in which the milk is secreted from the ultimate branches of arteries. The excretory ducts of these several glands gradually uniting as they ap- proach the nipple, form the tubuli lactiferi, which are usually more than a dozen in number, and open at its apex, but have little or no com- munication, as has been supposed, at the root of the nipple. These ducts, in their course from the glands, are surrounded by a ligamentary elastic substance, which terminates with them in the nipple. Both this substance, and the ducts which it contains, are capable of considerable extension and contraction; but in their natural state are moderately corrugated, so as to prevent an involuntary flow of milk, unless the distend- ing force be very great from the accumulation of too great a quantity. The whole substance of the nipple is very spongy and elastic: its external surface is une- ven, and full of small tubercles. The nipple is surrounded with a disk or circle of a different colour, called the areola; and on the inside of the skin, under the areola, are many sebaceous glands, which pour out a mucus to defend the areola and nipple: for the skin upon these parts is 330 PART IV. OF THE THORAX. is very thin; and the nervous papillæ lying very bare, are much exposed to irritation. The breasts are formed for the secretion of milk, which is destined for the nourishment of the child for some time after its birth. This se- cretion begins to take place soon after delivery, and continues to flow for many months in very large quantities, if the woman suckles her child. The operation of suction depends on the prin- ciples of the air-pump, and the flow of milk through the lactiferous tubes is facilitated by their being stretched out. The milk, examined chemically, appears to be composed of oil, mucilage, and water, and of a considerable quantity of sugar. The ge- nerality of physiologists have supposed that, like the chyle, it frequently retains the properties of the aliment and medicines taken into the sto- mach; but from some late experiments *, this supposition appears to be ill-founded. SECT. II. Of the Pleura. THE cavity of the thorax is every where lin- ed by a great membrane of a firm texture call- ed pleura. It is composed of two distinct porti- ons or bags, which, by being applied to each other laterally, form a septum called mediastinum; which * Journ. de Med. 1781. 331 PART IV. OF THE THORAX. which divides the cavity into two parts, and is attached posteriorly to the vertebræ of the back, and anteriorly to the sternum. But the two la- minæ of which this septum is formed, do not every where adhere to each other; for at the lower part of the thorax they are separated, to afford a lodgement to the heart; and at the upper part of the cavity, they receive between them the thymus. The pleura is plentifully supplied with arte- ries and veins from the internal mammary and the intercostals. Its nerves, which are very in- considerable, are derived chiefly from the dorsal and intercostal nerves. The surface of the pleura, like that of the pe- ritonæum and other membranes lining cavities, is constantly bedewed with a serous moisture (W) which prevents adhesion of the viscera. The mediastinum, by dividing the breast into two cavities, obviates many inconveniences, to which we should otherwise be liable. It pre- vents the two lobes of the lungs from compres- sing each other when we lie on one side; and consequently contributes to the freedom of res- piration, which is disturbed by the least pressure on the lungs. If the point of a sword pene- trates between the ribs into the cavity of the tho- rax, the lungs on that side cease to perform their office; (W) When this fluid is exhaled in too great a quantity, or is not properly carried off, it accumulates and constitutes the hydrops pectoris. 352 PART IV.OF THE THORAX. office; because the air being admitted through the wound, prevents the dilatation of that lobe; while the other lobe, which is separated from it by the mediastinum, remains unhurt, and con- tinues to perform its function as usual. SECT. II. Of the Thymus. THE thymus is a glandular substance, the use of which is not perfectly ascertained, its excre- tory duct not having yet been discovered. It is of an oblong figure, and is larger in the fœtus and in young children than in adults, being some- times nearly effaced in very old subjects. It is placed in the upper part of the thorax, between the two laminæ of the mediastinum; but at first is not altogether contained within the cavity of the chest, being found to border upon the upper extremity of the sternum. SECT. IV. Of the Diaphragm. THE cavity of the thorax is separated from that of the addomen, by a fleshy and membranous substance called the diaphragm or midriff. The greatest part of it is composed of muscular fibres; and on this account systematic writers usually place very properly among the muscles. Its mid- dle part is tendinous, and it is covered by the pleura above, and by the peritonæum below. It seems to have been improperly named septum transversum, 333 PART IV. OF THE THORAX. trasversum, as it does not make a plane trans- verse division of the two cavities, but forms a kind of vault, the fore-part of which is attach- ed to the sternum. Laterally it is fixed to the last of the true ribs, and to all the false ribs; and its lower and posterior part is attached to the vertebræ lumborum, where it may be said to be divided in two portions or crura (X). The principal arteries of the diaphragm are derived from the aorta, and its veins pass into the vena cava. It nerves are chiefly derived from the cervical pairs. It affords a passage to the vena cava through its tendinous part, and to the œsophagus through its fleshy portion. The aorta passes down behind it between its cura. The diaphragm not only serves to divide the thorax from the abdomen, but by its muscular structure is is rendered one of the chief agents in respiration. When its fibres contract, its convex side, which is turned towards the tho- rax, becomes gradually flat, and by increasing the cavity of the breast, affords room for a com- plete dilatation of the lungs, by means of the air which is then drawn into them by the act of inspiration. The fibres of the diaphragm then relax; and as it resumes its former state, the ca- vity (X) Anatomical writers have usually described the dia- phragm as being made up of two muscles united by a middle tendon; and these two portions or crura form what they speak of as the inferior muscle, arising from the sides and fore part of the vertebræ. 334 PART IV. OF THE THORAX. vity of thorax becomes gradually diminished, and the air is driven out again from the lungs by a motion contrary to the former one, called ex- spiration. It is in some measure, by means of the dia- phragm, that we void the fæces at the anus, and empty the urinary bladder. Besides these offices, the acts of coughing, sneezing, speaking, laugh- ing, gaping, and sighing, could not take place without its assistance; and the gentle pressure which all the abdominal viscera receive from its constant and regular motion, cannot fail to assist in the performance of the several functions which were ascribed to those viscera. SECT. V. Of the Trachea. THE trachea or windpipe, is a cartilaginous and membranous canal, through which the air passes into the lungs. Its upper part, which is called the larynx, is composed of five cartilages. The uppermost of these cartilages is placed over the glottis or mouth of the larynx, and is called epiglottis, which has been before spoken of, as closing the passage to the lungs in the act of swallowing. At the sides of the glottis are placed the two arytenoide cartilages, which are of a very complex figure, not easy to be de- scribed. The anterior and larger part of the larynx is made up of two cartilages; one of which is called thyroides or scutiformis, from its being 335 PART IV. OF THE THORAX. being shaped like a buckler; and the other cri- coides or annularis, from its resembling a ring. Both these cartilages may be felt immediately under the skin, at the fore-part of the throat, and the thyroides, by its convexity, forms an eminence called pomum adami, which is usually more considerable in the male than in the female subject. All these cartilages are united to each other by means of very elastic, ligamentous fibres; and are enabled, by the assistance of their several muscles, to dilate or contract the passage of the larynx, and to perform that variety of mo- tion which seems to point out the larynx as the principal organ of the voice; for when the air passes out through a wound in the trachea, it pro- duces no sound. These cartilages are moistened by a mucus, which seems to be secreted by minute glands si- tuated near them. The upper part of the trachea is covered anteriorly and laterally by a consider- able body, which is supposed to be of a glandu- lar structure, and from its situation near the thy- roid cartilage is called the thyroid gland; though its excretory duct has not yet been discovered, or its use ascertained. The glottis is interiorly covered by a very fine membrane, which is moistened by a constant supply of a watery fluid. From the larynx the canal begins to take the name of trachea or as- pera arteria, and extends from thence as far down 336 PART IV. OF THE THORAX. as the third or fourth vertebra of the back, where it divides into two branches, which are the right and left bronchial tube. Each of these bronchi (Y), ramifies through the substance of that lobe of the lungs to which it is distributed, by in- finite number of branches, which are formed of cartilages separated from each other like those of the trachea, by an intervening membranous and ligamentary substance. Each of these cartilages is of an angular figure; and as they become gra- dually less and less, in their diameter, the lower ones are in some measure received into those above them, when the lungs, after being inflat- ed, gradually collapse by the air being pushed out from them in expiration. As the branches of the bronchi become more minute, their car- tilages become more and more angular and mem- branous, till at length they are found to be per- fectly membranous, and at last become invisible. The trachea is furnished with fleshy or mus- cular fibres; some of which pass through its whole extent longitudinally, while the others are carried round, it in a circular direction; so that by the contraction or relaxation of these fi- 2 bres; (Y) The right bronchial tube is usually found to be some- what shorter and thicker than the left; and M. Portal, who has published a memoir on the action of the lungs on the aorta in respiration, observes, that the left bronchial tube is closely connected by the aorta; and from some experiments he is induced to conclude, that in the first respirations, the air only enters into the right lobe of the lungs. Memoires de l' Academie Royale des Sciences, 1769. 337 PART IV. OF THE THORAX. bres, it is enabled to shorten or lengthen itself, and likewise to dilate or contract the diameter of its passage. The trachea and its branches, in all their ra- mifications, are furnished with a great number of small glands which are lodged in their cellular substance, and discharge a mucous fluid on the inner surface of these tubes. The cartilages of the trachea, by keeping it constantly open, afford a free passage to the air, which we are obliged to be incessantly respiring; and its membranous part, by being capable of contraction and dilatation, enables us to receive and expel the air in a greater or less quantity, and with more or less velocity, as may be requir- ed in singing or in declamation. This membran- ous structure of the trachea posteriorly, seems likewise to assist in the descent of the food, by preventing that impediment to its passage down the œsophagus, which might be expected if the cartilages were complete rings. The trachea receives its arteries from the ca- rotid and subclavian arteries, and its veins pass into the jugulars. Its nerves arise from the re- current branch of the eighth pair, and from the cervical plexus. SECT. VI. Of the Lungs. The lungs fill the greater part of the cavity of the breast. They are of a soft and spongy tex- Y ture, 338 PART IV. OF THE THORAX. ture, and are divided into two lobes, which are separated from each other by the mediastinum, and are externally covered by a production of the pleura. Each of these is divided into two or three lesser lobes; and we commonly find three in the right side of the cavity, and two in the left. To discover the structure of the lungs, it is required to follow the ramifications of the bron- chi, which were described in the last section. These becoming gradually more and more mi- nute, at length terminate in the cellular spaces or vesicles, which make up the greatest part of the substance of the lungs, and readily communicate with each other. The lungs seem to possess but little sensibility. Their nerves, which are small, and few in num- ber, are derived from the intercostal and eighth pair. This last pair having reached the thorax, sends off a branch on each side of the trachea, called the recurrent, which reascends at the back of the trachea, to which it furnishes branches in its ascent, as well as to the œsophagus, but it is chiefly distributed to the larynx and its muscles. By dividing the recurrent and superior laryngeal nerves at their origin, an animal is deprived of its voice. There are two series of arteries which carry blood to the lungs: these are the arteriæ bron- chiales, and the pulmonary artery. The 339 PART IV. OF THE THORAX. The arteriæ bronchiales begin usually by two branches; one of which commonly arises from the right intercostal, and the other from the trunk of the aorta: but sometimes there are three of these arteries, and in some subjects only one. The use of these arteries is to serve for the nou- rishment of the lungs, and their ramifications are seen creeping every where on the branches of the bronchi. The blood is brought back from them by the bronchial vein into the vena azygos. The pulmonary artery and vein are not in- tended for the nourishment of the lungs; but the blood in its passage through them is destined to undergo some changes, or to acquire certain essential properties (from the action of the air), which it has lost in its circulation through the other parts of the body. The pulmo- nary artery receives the blood from the right ventricle of the heart, and dividing into two branches, accompanies the bronchi every where, by its ramifications through the lungs; and the blood is afterwards conveyed back by the pulmo- nary vein, which gradually forming a consider- able trunk, goes to empty itself into the left ventricle of the heart; so that the quantity of blood which enters into the lungs, is perhaps greater than that which is sent in the same pro- portion of time through all the other parts of the body. Y2 SECT. 340 PART IV. OF THE THORAX. SECT. VII. Of Respiration. RESPIRATION constitutes one of those func- tions which are properly termed vital, as being essential to life; for to live and to breathe are in fact synonymous terms. It consists in an alter- nate contraction and dilatation of the thorax, by first inspiring air into the lungs, and then expell- ing it from them in exspiration. It will perhaps be easy to distinguish and point out the several phenomena of respiration; but to explain their physical cause will be attended with difficulty: for it will naturally be enquir- ed, how the lungs, when emptied of the air, and contracted by exspiration, become again in- flated, they themselves being perfectly passive? How the ribs are elevated in opposition to their own natural situation? and why the diaphragm is contracted downwards, towards the abdomen? Were we to assert that the air, by forcing its way into the cavity of the lungs, dilated them, and consequently elevated, the ribs, and pressed down the diaphragm, we should speak erroneously. What induces the first inspiration, it is not easy to ascertain; but after an animal has once respir- ed, it would seem likely that the blood, after ex- spiration, finding its passage through the lungs obstructed, becomes a stimulus, which induces the intercostal muscles and the diaphragm to con- tract, and enlarge the cavity of the thorax, in consequence 341 PART IV. OF THE THORAX. consequence perhaps of a certain nervous influ- ence, which we will not here attempt to explain. The air then rushes into the lungs; every branch of the bronchial tubes, and all the cellular spaces into which they open, become fully dilated; and the pulmonary vessels being equally distend- ed, the blood flows through them with ease. But as the stimulus which first occasioned this dila- tation ceases to operate, the muscles gradually contract, the diaphragm rises upwards again, and diminishes the cavity of the chest; the ribs re- turn to their former state; and as the air passes out in exspiration, the lungs gradually collapse, and a resistance to the passage of the blood again takes place. But the heart continuing to receive and expel the blood, the pulmonary artery be- gins again to be distended, the stimulus is re- newed, and the same process is repeated, and continues to be repeated, in a regular succession, during life: for though the muscles of respiration, having a mixed motion, are (unlike the heart) in some measure dependent on the will, yet no human being, after having once respired, can live many moments without it. In an attempt to hold one's breath, the blood soon begins to distend the veins, which are, unable to empty their contents into the heart; and we are able only, during a very little time, to resist the sti- mulus to inspiration. In drowning, the circu- lation seems to be stopped upon this principle; and in hanging, the pressure made on the jugu- lar 342 PART IV. OF THE THORAX. lar veins, may co-operate with the stoppage of respiration in bringing on death. Till within these few years physiologists were entirely ignorant of the use of respiration. It was at length discovered in part by the illustrious Dr. Priestley. He found that the air exspired by animals was phlogisticated; and that the air was fitter for respiration, or for supporting ani- mal life, in proportion as it was freer from the phlogistic principle. It had long been observ- ed, that the blood in passing through the lungs acquired a more florid colour. He therefore suspected, that it was owing to its having im- parted phlogiston to the air: and he satisfied himself of the truth of this idea, by experi- ments, which showed, that the crassamentum of extravasated blood, phlogisticated air in propor- tion as it lost its dark colour. He farther found, that blood thus reddened had a strong attraction for phlogiston; insomuch that it was capable of taking it from phlogisticated air, thereby becom- ing of a darker colour, from hence it appeared that the blood, in its circulation through the ar- terial system, imbibes a considerable quantity of phlogiston, which is discharged from it to the air in the lungs. This discovery has since been prosecuted by two very ingenious physiologists, Dr. Crawford and Mr. Elliot. It had been shown by profe- sors Black and Irvine, that different bodies have different capacities for containing fire. For ex- ample, 343 PART IV. OF THE THORAX. ample, that oil and water, when equally hot to the sense and the thermometer, contain different proportions of that principle; and that unequal quantities of it are required, in order to raise those substances to like temperatures. The en- quiries of Dr. Crawford and Mr. Elliot tend to prove, that the capacities of bodies for contain- ing fire are diminished by the addition of phlo- giston, and increased by its separation: the ca- pacity of calx of antimony, for example, being greater than that of the antimony itself. Com- mon air contains a great quantity of fire; com- bustible bodies very little. In combustion, a double elective attraction takes place; the phlo- giston of the body being transferred to the air, the fire contained in the air to the combustible body. But as the capacity of the latter is not increased so much as that of the former is dimi- nished, only part of the extricated fire will be absorbed by the body. The remainder therefore will raise the temperature of the compound; and hence we may account for the heat attending combustion. As the use of respiration is to dephlogisticate the blood, it seems probable, that a like double elective attraction takes place in this process; the phlogiston of the blood being transferred to the air, and the fire contained in the air to the blood; but with this difference, that the capacities being equal, the whole of the extricated fire is absorbed by the latter. The blood in this state circulating through the body, imbibes 344 PART IV. OF THE THORAX. imbibes phlogiston, and of course gives out its fire; part only of which is absorbed by the parts furnishing the phlogiston; the remainder, as in combustion, becoming sensible; and is therefore the cause of the heat of the body, or what is called animal heat. In confirmation of this doctrine it may be observed, that the venous blood contains less fire than the arterial; combustible bodies less than incombustible ones; and that air contains less of this principle, according as it is rendered, by combination with phlogiston, less fit for res- piration (Z). In ascending very high mountains, respiration is found to become short and frequent, and some- times to be attended with a spitting of blood. These symptoms seem to be occasioned by the air being too rare and thin to dilate the lungs sufficiently; and the blood gradually accumulat- ing in the pulmonary vessels, sometimes bursts through their coats, and is brought up by cough- ing. This has likewise been accounted for in a different way, by supposing that the air contain- ed in the blood, not receiving an equal pressure from that of the atmosphere, expands, and at length ruptures the very minute branches of the pulmonary vessels; upon the same principle that fruits and animals put under the receiver of an air- (Z) See Crawford's Experiments and Observations on Animal Heat, and Elliot's Philosophical Observations. 345 PART IV. OF THE THORAX. air-pump, are seen to swell as the outer air be- comes exhausted. But Dr. Darwin of Litchfield has lately published some experiments, which seem to prove, that no air or elastic vapour does exist in the blood-vessels, as has been generally supposed: and he is induced to impute the spit- ting of blood, which has sometimes taken place in ascending high mountains, to accident, or to violent exertions; as it never happens to animals that are put into the exhausted receiver of an air- pump, where the diminution of pressure is ma- ny times greater than on the summit of the high- est mountains. SECT. VIII. Of the Voice. RESPIRATION has already been described as affording us many advantages; and next to that of life, its most important use seems to be that of forming the voice and speech. The ancients, and almost all the moderns, have considered the organ of speech as a kind of musical instrument, which may be compared to a flute, to an haut- boy, to an organ, &c. and they argue after the following manner. The trachea, which begins at the root of the tongue, and goes to terminate in the lungs, may be compared to the pipe of an organ the lungs di- lating like bellows during the time of inspiration; and as the air is driven out from them in exspi- ration, it finds its passage straitened by the carti- lages 346 PART IV. OF THE THORAX. lages of the larynx, against which it strikes. As these cartilages are more or less elastic, they occa- sion in their turn more or less vibration in the air, and thus produce the sound of the voice; the variation in the sound and tone of which de- pends on the state of the glottis, which, when straitened, produces an acute tone, and a grave one when dilated. The late M. Ferein communicated to the French Academy of Sciences a very ingenious theory on the formation of the voice. He con- sidered the organ of the voice as a firing, as well as a wind, instrument; so that what art has hi- therto been unable to construct, and what both the fathers Mersenne and Kircher so much wish- ed to see, M. Ferein imagined he had at length discovered in the human body. He observes, that there are at the edges of the glottis certain tendinous chords, placed horizontally across it, which are capable of considerable vibration, so as to produce sound, in the same manner as it is produced by the strings of a violin or a harpsi- chord: and he supposes that the air, as it passes out from the lungs, acts as a bow on these strings, while the efforts of the breast and lungs regulate its motion, and produce the variety of tones. So that according to this system the variation in the voice is not occasioned by the dilatation or contraction of the glottis, but by the distension or relaxation of these strings, the sound being more or less acute in proportion as they are more or less 347 PART IV. OF THL THORAX. less stretched out. Another writer on this sub- ject supposes, that the organ of voice is a double instrument, which produces in unison two sounds of a different nature; one by means of the air, and the other by means of the chords of the glottis. Neither of these systems, however, are universally adopted. They are both liable to insuperable difficulties; so that the manner in which the voice is formed has never yet been sa- tisfactorily ascertained: we may observe, how- ever, that the sound produced by the glottis is not articulated. To effect this, it is required to pass through the mouth, where it is differently modified by the action of the tongue, which is either pushed against the teeth, or upwards to- wards the palate; detaining it in its passage, or permitting it to flow freely, by contracting or di- lating the mouth. SECT. IX. Of Dejection. By dejection we mean the act of voiding the fæces at the anus; and an account of the man- ner in which this is conducted was reserved for this part of the work, because it seemed to re- quire a knowledge of respiration to be perfectly understood. The intestines were described as having a pe- ristaltic motion, by which the fæces were gra- dually advancing towards the anus. Now, when- ever the fæces are accumulated in the intestinum rectum 348 PART IV. OF THE THORAX. rectum in a sufficient quantity to become trouble- some, either by their weight or acrimony, they excite a certain uneasiness which induces us to go to stool.—To effect this, we begin by mak- ing a considerable inspiration; in consequence of which the diaphragm is carried downwards to- wards the lower belly; the abdominal muscles are at the same time contracted in obedience to the will; and the intestines being compressed on all sides, the resistance of the sphincter is over- come, and the fæces pass out at the anus; which is afterwards drawn up by its longitudinal fibres, which are called levatores ani, and then by means of its sphincter is again contracted: but it some- times happens, as in dysenteries, for instance, that the fæces are very liquid, and have consi- derable acrimony; and then the irritation they occasion is more frequent, so as to promote their discharge without any pressure from the dia- phragm or abdominal muscles; and sometimes involuntarily, as is the case when the sphincter becomes paralytic. SECT. X. Of the Pericardium, and of the Heart and its Auricles. THE two membranous bags of the pleura, which were described as forming the mediasti- num, recede one from the other, so as to afford a lodgement to a firm membranous sac, in which the heart is securely lodged; this sac, which is the 349 PART IV. OF THE THORAX. the pericardium, appears to be composed of two tunics, united to each other by cellular mem- brane.—The outer coat, which is thick, and in some places of tendinous complexion, is a pro- duction of the mediastinum; the inner coat, which is extremely thin, is reflected over the auricles and ventricles of the heart, in the same manner as the tunica conjunctiva, after lining the eye lids, is reflected over the eye. This bag adheres to the tendinous part of the diaphragm, and contains a coagulable lymph, the liquor pericardii, which serves to lubricate the heart and facilitate its motions; and seems to be secreted and absorbed in the same manner as it is in the other cavities of the body. The arteries of the pericardium are derived from the phrenic, and its veins pass into veins of the same name; its nerves are likewise branches of the phrenic. The size of the pericardium is adapted to that of the heart, being usually large enough to con- tain it loosely. As its cavity does not extend to the sternum, the lungs cover it in inspiration; and as it every where inverts the heart, it effectu- ally secures it from being injured by lymph, pus, or any other fluid, extravasated into the cavities of the thorax. The heart is a hollow muscle of a conical shape, situated transversely between the two laminæ of the mediastinum, at the lower part of the thorax; having its basis turned towards the right side, and its 350 PART IV. OF THE THORAX. its point or apex towards the left.—Its lower sur- face is somewhat flattened towards the diaphragrm Its basis, from which the great vessels originate, is covered with fat, and it has two hollow and fleshy-appendages, called auricles.—Round these several openings, the heart seems to be of a firm ligamentous texture, from which all its fibres seem to originate; and as they advance from thence towards the apex, the substance of the heart seems to become thinner. The heart includes two cavities or ventricles, which are separated from each other by a fleshy septum; one of these is called the right and the other the left, ventricle; though perhaps, with respect to their situation, it would be more pro- per to distinguish them into the anterior and pos- terior ventricles. The heart is exteriorly covered by a very fine membrane; and its structure is perfectly muscu- lar or fleshy, being composed of fibres which are described as passing in different directions; some as being extended longitudinally from the basis to the apex; others, as taking an oblique or spiral course; and a third sort as being placed in a transverse direction (A).—Within the two ventricles we observe several furrows; and there are (A) Authors differ about the course and distinctions of these fibres; and it seems right to observe, that the structure of the heart being more compact than that of other muscles, its fibres are not easily separated. 351 PART IV. OF THE THORAX. are likewise tendinous strings, which arise from fleshy columnæ in the two cavities, and are attach- ed to the valves of the auricles: That the use of these and the other valves of the heart may be understood, it must be observed, that four large vessels pass out from the basis of the heart, viz. two arteries and two veins; and that each of these vessels is furnished with a thin membran- ous production, which is attached all round to the borders of their several orifices, from whence hanging loosely down they appear to be divided into two or three distinct portions. But as their uses in the arteries and veins are different, so are they differently disposed. Those of the arteries are intended to give way to the passage of the blood into, them from the ventricles, but to op- pose its return: and, on the contrary, the valves of the veins are constructed so as to allow the blood only to pass into the heart. In consequence of these different uses, we find the valves of the pulmonary artery and of the aorta attached to the orifices of those vessels, so as to have their concave surfaces turned towards the artery; and their convex, surfaces, which mutually meet to- gether, being placed towards the ventricle, only permit the blood to pass one way, which is into the arteries. There are usually three of these valves belonging to the pulmonary artery, and as many to the aorta; and from their figure they are called valvulæ semilunares. The communi- cation between the two great veins and the ven- tricles 352 PART IV. OF THE THORAX. tricles is by means of the two appendages or au- ricles into which the blood is discharged; so that the other valves which may be said to belong to the veins, are placed in each ventricle, where the auricle opens into it. The valves in the right ventricle are usually three in number, and are named valvulæ tricuspides; but in the left ventricle we commonly observe only two, and these are the valvulæ mitrales. The membranes which form these valves in each cavity are at- tached so as to project somewhat forward; and both the tricuspides and the mitrales are connect- ed with the tendinous strings, which were de- scribed as arising from the fleshy columnæ. By the contraction of either ventricle, the blood is driven into the artery which communicates with that ventricle; and these tendinous strings being gradually relaxed as the sides of the cavity are brought nearer to each other, the valves natu- rally close the opening into the auricle, and the blood necessarily directs its course into the then only open passage, which is into the artery; but after this contraction, the heart becomes relaxed, the tendinous strings are again stretched out, and, drawing the valves of the auricle downwards, the blood is poured by the veins into the ventri- cle, from whence, by another contraction, it is again thrown into the artery, as will be describ- ed hereafter. The right ventricle is not quite so long, though somewhat larger, than the left; but the latter has more substance than the other: 3 and 353 PART IV. OF THE THORAX. and this seems to be, because it is intended to transmit the blood to the most distant parts of the body, whereas the right ventricle distributes it only to the lungs. The heart receives its nerves from the par va- gum and the intercostals. The arteries which serve for its nourishment are two in number, and arise from the aorta. They surround in some measure the basis of the heart, and from this course are called the coronary arteries. From these arteries the blood is returned by veins of the same name into the auricles, and even into the ventricles. The muscular bags called the auricles are situ- ated at the basis of the heart, at the sides of each other; and, corresponding with the two ventri- cles, are like those two cavities distinguished in- to right and left. These sacs, which are inte- riorly unequal, have externally a jagged appen- dix; which, from its having been compared to the extremity of an ear, has given them their name of auricles. SECT. XI. Angiology, or a Description of the Blood-vessels. The heart has been described as contracting itself, and throwing the blood from its two ventricles into the pulmonary artery and the aorta, and then as relaxing itself and receiv- ing a fresh supply from two large veins, which Z are 354 PART IV. OF THE THORAX. are the pulmonary vein and the vena cava. We will now point out the principal distributions of these vessels. The pulmonary artery arises from the right ventricle by a large trunk, which soon divides into two considerable branches, which pass to the right and left lobes of the lungs: each of these branches is afterwards divided and subdi- vided into an infinite number of branches and ramifications, which extend through the whole substance of the lungs; and from these branches the blood is returned by the veins, which, con- trary to the course of the arteries, begin by very minute canals, and gradually become larger, forming at length four large trunks called pul- monary veins, which terminate in the left auricle by one common opening, from whence the blood passes into the left ventricle. From this same ventricle arises the aorta or great artery, which at its beginning is nearly an inch in dia- meter : it soon sends off two branches, the coro- naries, which go to be distributed to the heart and its auricles. After this, at or about the third or fourth vertebra of the back, it makes a con- siderable curvature; from this curvature (B) arise three (B) Anatomists usually call the upper part of this curva- ture aorta ascendens; and the other part of the artery to its division at the iliacs, aorta descendens: but they differ about the place where this distinction is to be introduced; and it seems sufficiently to answer every purpose, to speak only of the aorta and its curvature. 355 PART IV. OF THE THORAX. three arteries; one of which soon divides into two branches. The first two are the left sub- clavian and the left carotid, and the third is a common trunk to the right subclavian and right carotid; though sometimes both the carotids arise distinctly from the aorta. These two carotids ascend within the subcla- vians, along the sides of the trachea; and when they have reached the larynx, divide into two principal branches, the internal and external ca- rotid. The first of these runs a little way back- wards in a bending direction; and having reach- ed the under part of the ear, passes through the canal into the os petrosum, and entering into the cavity of the cranium, is distributed to the brain and the membranes which invelope it, and like- wise to the eye. The external carotid divides into several branches, which are distributed to the larynx, pharynx, and other parts of the neck; and to the jaws, lips, tongue, eyes, temples; and all the external parts of the head. Each subclavian is likewise divided into a great number of branches. It sends off the ver- tebral artery, which passes through the openings we see at the bottom of the transverse processes of the vertebræ of the neck, and in its course sends off many ramifications to the neighbour- ing parts. Some of its branches are distributed to the spinal marrow, and after a considerable inflexion it enters into the cranium, and is dis- tributed to the brain. The subclavian likewise Z2 sends 356 PART IV. OF THE THORAX. sends off branches to the muscles of the neck and the scapula; and mediastinum, thymus, peri- cardiu, diaphragm, the breasts, and the mus- cles of the thorax, and even of the abdomen, derive branches from the subclavian, which are distinguished by different names, alluding to the parts to which they are distributed; and the mammary, the phrenic, the intercostal, &c. But notwithstanding the great number of branches which have been described as arising from the subclavian, it is still a considerable artery when it reaches the axilla, where it drops its former name, which alludes to its passage under the cla- vicle, and is called the axillary artery; from which a variety of branches are distributed to the muscles of the breast, scapula, and arm.—But its main trunk taking the name of brachialis, runs along on the inside of the arm near the os humeri, till it reaches the joint of the fore-arm, and then it divides into branches. This divi- sion however is different in different subjects; for in some it takes place higher up and in others lower down. When it happens to divide above the joint, it may be considered as a happy dis- position in case of an accident by bleeding; for supposing the artery to be unfortunately punc- tured by the lancet, and that the hæmorrhage could only be stopped by making a ligature on the vessel, one branch would remain unhurt, through which the blood would pass uninterupt- ed to the fore-arm and hand. One of the two branches 357 PART IV. OF THE THORAX. branches of the brachialis plunges down under the flexor muscles, and runs along the edge of the ulna; while the other is carried along the outer surface of the radius, and is easily felt at the wrist, where it is only covered by the com- mon integuments. Both these branches com- monly unite in the palm of the hand, and form an arterial arch from whence branches are de- tached to the fingers. The aorta, after having given off at its cur- vature the carotids and subclavians which convey blood to all the upper parts of the body, descends upon the bodies of the vertebræ a little to the left, as far as the os sacrum, where it drops the name of aorta, and divides into two considerable branches. In this course, from its curvature to its bifurca- tion, it sends off several arteries in the following order: 1. One or two little arteries, first demon- strated by Ruysch as going to the bronchi, and called arteriæ bronchinales Ruyschii. 2. The ar- teriæ œsophageæ. These are commonly three or four in number. They arise from the fore- part of the aorta, and are distributed chiefly to the œsophagus. 3. The inferior intercostal ar- teries, which are distributed between the ribs in the same manner as the arteries of the three or four superior ribs are, which are derived from the subclavian. These arteries send off branches to the medulla spinalis. 4. The diaphragmatic or inferior phrenic arteries, which go to the diaphragm, stomach, omentum, duodenum, pancreas 358 PART IV. OF THE THORAX. pancreas, spleen, liver, and gall-bladder. 5. The cœliac, which sends off the coronary-sto- machic, the splenic, and the hepatic artery. 6. The superior mesenteric artery and small intes- tines. 7. The emulgents, which go to the kid- neys. 8. The arteries, which are distributed to the glandulæ renales. 9. The spermatic. 10. The inferior mesenteric artery, which ra- mifies through the lower portions of the mesen- tery and the large intestines.—A branch of this artery which goes to the rectum is called the in- ternal hemorrhoidal. 11. The lumbar arteries, and a very small branch called the sacra, which are distributed to the muscles of the loins and abdomen, and to the os sacrum and medulla spinalis. The trunk of the aorta, when it has reached the last vertebra lumborum, or the os sacrum, drops the name of aorta, and separates into two fork- ed branches called the iliacs. Each of these soon divides into two branches; one of which is call- ed internal iliac, or hypogastric artery, and is dis- tributed upon the contents of the pelvis and up- on the muscles on its outer side. One branch, called pudenda communis, sends small ramifica- tions to the end of the rectum under the name of hemorrhoides externæ, and is afterwards dis- tributed upon the penis. The other branch, the external iliac, after having given off the cir- cumflex artery of the os ilium and the epigastric, which is distributed to the recti-muscles, passes out 359 PART IV. OF THE THORAX. out of the abdomen under Poupart's ligament, and takes the name of crural artery. It de- scends on the inner part of the thigh close to the os femoris, sending off branches to the mus- cles, and then sinking deeper in the hind part of the thigh, reaches the ham, where it takes the name of popliteal: after this it separates into two considerable branches: one of which is called the anterior tibial artery; the other divides into two branches, and these arteries all go to be dis- tributed to the leg and foot. The blood, which is thus distributed by the aorta to all parts of the body, is brought back by the veins, which are supposed to be continued from the ultimate branches of arteries; and uniting together as they approach the heart, at length from the large trunks, the vena cava ascendens, and vena cava descendens. All the veins which bring back the blood from the upper extremities, and from the head and breast, pass into the vena cava descendens; and those which return it from the lower parts of the body terminate in the vena cava ascendens; and these two cavas uniting together as they ap- proach the heart, open by one common orifice into the left auricle. It does not here seem to be necessary to fol- low the different divisions of the veins as we did those of the arteries; and it will be sufficient to remark, that in general every artery is accom- panied by its vein, and that both are distinguished by 360 PART IV. OF THE THORAX. by the same name. But, like many other ge- neral rules, this too has its exceptions (C). The veins for instance, which accompany the exter- nal and internal carotid, are not called the car- toid veins, but the external and internal jugular. —In the thorax, there is a vein distinguished by a proper name, and this is the azygos, or ve- na sine parti. This vein, which is a pretty con- siderable one, runs along by the right side of the vertebræ of the back, and is chiefly destin- ed to receive the blood from the intercostals on that side, and from the lower half of those on the left side, and to convey it into the vena cava descendens. In the abdomen we meet with a vein, which is still a more remarkable one, and this is the vena portæ, which performs the office both of an artery and a vein. It is formed by a re-union of all the veins which come from the stomach, intestines, omentum, pancreas, and spleen, so as to compose one great trunk, which goes to ramify through the liver; and after hav- ing deposited the bile, its ramifications unite and bring back into the vena cava, not only the blood which the vena portæ had carried into the liver, but likewise the blood from, the hepatic artery. Every artery has a vein which corresponds with it; but the trunks and branches of the veins are (C) In the extremities, some of the deep-seated veins, and all the superficial one, take a course different from that of the arteries. 361 PART IV. OF THE THORAX. are more numerous than those of the arteries. —The reasons for this disposition are perhaps more difficult to be explained; the blood in its course through the veins is much farther remov- ed from the source and cause of its motion, which are in the heart, than it was when in the arteries; so that its course is consequently less rapid, and enough of it could not possibly be brought back to the heart in the moment of its dilatation, to equal the quantity which is driven into the arteries from the two ventricles, at the time they contract; and the equilibrium which is so essential to the continuance of life and health would consequently be destroyed, if the capacity of the veins did not exceed that of the arteries, in the same proportion that the ra- pidity of the blood's motion through the arteries exceeds that of its return through the veins. A large artery ramifying through the body, and continued to the minute branches of veins, which gradually unite together to form a large trunk, may be compared to two trees united to each other at their tops; or rather as having their ramifications so disposed that the two trunks terminate in one common point; and if we farther suppose, that both these trunks and their branches are hollow, and that a fluid is inces- santly circulated through them, by entering into one of the trunks and returning through the other, we shall be enabled to conceive how the blood is 362 PART IV. OF THE THORAX. is circulated through the vessels of the human body. Every trunk of an artery, before it divides, is nearly cylindrical, or of equal diameter through its whole length, and so are all its branches when examined separately. But every trunk seems to contain less blood than the many branches do into which that trunk separates; and each of these branches probably contains less blood than the ramifications do into which it is subdivided: and it is the same with the veins; the volume of their several ramifications, when considered together, being found to exceed that of the great trunk which they form by their union. The return of the blood through the veins to the heart, is promoted by the action of the mus- cles, and the pulsation of the arteries. And this return is likewise greatly assisted by the valves which are to be met with in the veins, and which constitute one of the great distinctions between them and the arteries. These valves, which are supposed to be formed by the inner coat of the veins, permit the blood to flow from the extremities towards the heart, but oppose its return. They are most frequent in the smaller veins. As the column of blood increases, they seem to become less necessary; and therefore in the vena cava ascendens, we meet with only one valve, which is near its origin. The arteries are composed of several tunics. Some 363 PART IV. OF THE THORAX. Some writers enumerate five of these tunics; but perhaps we may more properly reckon only three, viz. the nervous, muscular, and cuticular coats. The veins are by some anatomists de- scribed as having the same number of coats as arteries; but as they do not seem to be irrita- ble, we cannot with propriety suppose them to have a muscular tunic. We are aware of Dr. Verschuir's * experiments to prove that the ju- gular and some other veins possess a certain de- gree of irritability; but it is certain, that his experiments, repeated by others, have produc- ed a different result; and even he himself allows, that sometimes he was unable to distinguish any such property in the veins. Both these series of vessels are nourished by still more minute arteries and veins, which are seen creeping over their coats, and ramifying through their whole substance, and are called vasa vasorum; they have likewise many minute branches of nerves. The arteries are much stronger than the veins, and they seem to require this force to be enabled to resist the impetus with which the blood circu- lates through them, and to impel it on towards the veins. When the heart contracts, it impels the blood into the arteries, and sensibly distends them; and these vessels again contract, as the heart be- comes relaxed to receive more blood from the auricles; * De Arteriarum et Venarum vi irritabili, 4to. 364 PART IV. OF THE THORAX. auricles; so that the cause of the contraction and dilatation of the arteries seems to be easy to be understood, being owing in part to their own contractile power, and in part to the action of the heart; but in the veins, the ef- fects of this impulse not being so sensibly felt, and the vessels themselves having little or no con- tractile power, the blood seems to flow in a con- stant and equal stream: and this, together with its passing gradually from a small channel into a larger one, seems to be the reason why the veins have no pulsatory motion, except the large ones near the heart; and in these it seems to be oc- casioned by the motion of the diaphragm, and by the regurgitation of the blood in the cavas. SECT. XII. Of the Action of the Heart, Auri- cles, and Arteries. The heart, at the time it contracts, drives the blood from its ventricles into the arteries; and the arteries being thus filled and distended, are naturally inclined to contract the moment the heart begins to dilate, and ceases to supply them with blood. These alternate motions of contraction and dilatation of the heart and arte- ries, are distinguished by the names systole and diastole. When the heart is in a state of con- traction or systole, the arteries are at that instant distended with blood, and in their diastole; and it 365 PART IV. OF THE THORAX. it is in this state we feel their pulsatory motion, which we call the pulse. When the heart dilates, and the arteries contract, the blood is impelled onwards into the veins, through which it is re- turned back to the heart. While the heart, how- ever, is in its systole, the blood cannot pass from the veins into the ventricles, but is detained in the auricles, which are two reservoirs formed for this use, till the diastole, or dilatation of the heart, takes place; and then the distended au- ricles contract, and drive the blood into the ven- tricles: so that the auricles have an alternate sys- tole and diastole as well as the heart. Although both the ventricles of the heart con- tract at the same time, yet the blood passes from one to the other. In the same moment, for in- stance, that the left ventricle drives the blood into the aorta, the right ventricle impels it into pulmonary artery, which is distributed through all the substance of the lungs. The blood is afterwards brought back into the left ventricle by the pulmonary vein, at the same time that the blood is returned by the cavas, into the right ventricle, from all the other parts of the body. This seems to be the mode of action of the heart and its vessels: but the cause of this action has, like all other intricate and interesting sub- jects, been differently explained. It seems to depend on the stimulus made on the different parts of the heart by the blood itself, which by its 366 PART IV. OF THE THORAX. its quantity and heat, or other properties (D), is perhaps capable of first exciting that motion, which is afterwards continued through life, in- dependent of the will, by a regular return of blood to the auricles, in a quantity proportion- ed to that which is thrown into the arteries. The heart possesses the vis insita, or principle of irritability, in a much greater degree than any other muscle of the body. The pulse is quicker in young than in old subjects, because the former are cæt. par. more irritable than the latter. Up- on the same principle we may explain, why the pulse is constantly quicker in weak than in ro- bust persons. SECT. XIII. Of the Circulation. After what has been observed of the struc- ture and action of the heart and its auricles, and likewise of the arteries and veins, there seem to be but very few arguments required to demon- strate the circulation of the blood, which has long since been established as a medical truth. This circulation may be defined to be a perpetual mo- tion of the blood, in consequence of the action of (D) Dr. Harvey long ago suggested, that the blood is pos- sessed of a living principle; and Mr. J. Hunter has lately en- deavoured to revive this doctrine; in support of which he has adduced many ingenious arguments. The subject is a curious one, and deserves to be prosecuted as an inquiry which cannot but be interesting to physiologists. 367 PART IV. OF THE THORAX. of the heart and arteries, which impel it through all the parts of the body, from whence it is brought back by the veins of the heart. A very satisfactory proof of this circulation, and a proof easy to be understood may be de- ducted from the different effects of pressure on an artery and a vein. If a ligature, for instance, is passed round an artery, the vessel swells con- siderably between the ligature and the heart; whereas, if we tie up a vein, it only becomes filled between the extremity and the ligature, and this is what we every day observe in bleed- ing. The ligature we pass round the arm on these occasions, compresses the superficial veins; and the return of the blood through them be- ing impeded, they become distended. When the ligature is too loose, the veins are not suffi- ciently compressed, and the blood continues its progress towards the heart; and, on the con- trary, when it is made too tight, the arteries themselves becomes compressed : and the flow of the blood through them being impeded, the veins cannot be distended. Another phænomenon, which effectually proves the circulation, is the loss of blood that every living animal sustains by opening only a single artery of a moderate size; for it continues to flow from the wounded vessel till the equili- brium is destroyed which is essential to life. This truth was not unknown to the ancients; and it seems, strange that it did not lead them to a knowledge 368 PART IV. OF THE THORAX. knowledge of the circulation, as it sufficiently proves, that all the other vessels must communi- cate with that which is opened. Galen, who lived more than 1500 years ago, drew this con- clusion from it; and if we farther observe, that he describes (after Erasistratus, who flourished about 450 years before him) the several valves of the heart, and determines their disposition and uses, it will appear wonderful, that a pe- riod of near 200 years should afterwards elapse before the true course of the blood was ascertain- ed. This discovery, for which we are indebt- ed to the immortal Harvey, has thrown new lights on physiology and the doctrine of dis- eases, and constitutes one of the most impor- tant periods of anatomical history. SECT. XIV. Of the Nature of the Blood. BLOOD, recently drawn from a vein into a bason, would seem to be an homogeneous fluid of a red colour (E); but when suffered to rest, it soon coagulates, and divides into two parts, which are distinguished by the names of crassa- mentum and serum. The crassamentum is the red coagulum, and the serum is the water in which it floats. Each of these may be again se- parated into two others; for the crassamentum, by (E) The blood, as it flows through the arteries, is ob- served to be more florid than it is in the veins; and this red- ness is acquired in its passage through the lungs. Vid. sect. vii. 369 PART IV. OF THE THORAX. by being repeatedly washed in warm water, gives out all its red globules, and what remains ap- pears to be composed of the coagulable lymph (F), which is a gelatinous substance, capable of being hardened by fire till it becomes perfectly horny: and if we expose the serum to a certain degree of heat, part of it will be found to coagulate like the white of an egg, and there will remain a clear and lympid water, resembling urine both in its appearance and smell. The serum and crassamentum differ in their proportion in different constitutions; in a strong person, the crassamentum is in a greater pro- portion to the serum than in a weak one;* and the same difference is found to take place in dis- eases (G). Aa SECT. XV. (F) It may not be improper to observe, that till of late the coagalable lymph has been confounded with the serum of the blood, which contains a substance that is likewise coagula- ble, though only when exposed to heat, or combined with certain chemical substances; whereas the other coagulates spontaneously when exposed to the air or to rest. * Hewson's Experim. Enq. Part. I. (G) When the blood separates into serum and crassamen- tum, if the latter be covered with a crust of a whitish or buff colour, it, has been usually considered as a certain proof of the blood's being in a state of too great viscidity. This appearance commonly taking place in inflammatory diseases, has long served to confirm the theory which ascribes the cause of inflammation to lentor and obstrudsions. But from the late Mr. Hewson's experiments it appears, that when the action of the arteries is increased, the blood, in- stead of being more viscid, is, on the conrtary, more fluid than 370 PART IV. OF THE THORAX. SECT. XV. Of Nutrition. THE variety of functions which we have de- scribed as being incessantly performed by the liv- ing body, and the continual circulation of the blood through it, must necessarily occasion a constant dissipation of the several parts which enter into its composition. In speaking of the insensible perspiration, we observed how much was incessantly passing off from the lungs and the surface of the skin. The discharge by urine is likewise every day considerable; and great part of the bile, saliva, &c. are excluded by stool. But the solid, as well as the fluid parts of the body, require a constant renewal of nu- tritious particles. They are exposed to the at- trition of the fluids which are circulated through them; and the contraction and relaxation they repeat so many thousand times in every day, would necessarily occasion a dissolution of the machine, if the renewal was not proportioned to the waste. It is easy to conceive how the chyle formed from the aliment is assimilated into the nature of the than in the ordinary state, previous to inflammation : and that in consequence of this, the coagulable lymph suffers the red globules, which are the heaviest part of the blood, to fall down to the bottom before it coagulates : so that the crassamentum is divided into two parts; one of which is found to consist of the coagulable lymph alone (in this case termed the buff;) and the other, partly of this and partly of the red globules. 371 PART IV. OF THE THORAX. the blood, and repairs the loss of the fluid parts of our body; but how the solids are renewed, has never yet been satisfactorily explained. The nutritious parts of the blood are probably de- posited by the arteries by exsudation through their pores into the tela cellulosa; and as the solid parts of the body are in the embryo only a kind of jelly, which gradually acquires the degree of consistence they are found to have when the body arrives at a more advanced age; and these same parts which consist of bones, cartilages, ligaments, muscles, &c. are some- times reduced again by diseases to a gelati- nous state; we may, with some degree of pro- bability, consider the coagulable lymph as the source of nutrition. If the supply of nourishment exceeds the de- gree of waste, the body increases; and this hap- pens in infancy and in youth: for at those peri- ods, but more particularly the former one, the fluids bear a large proportion to the solids; and the fibres being soft and yielding, are proporti- onably more capable of extension and increase. But when the supply of nutrition only equals the waste, we neither increase or decrease; and we find this to be the case when the body has attained its full growth or acme: for the solids having then acquired a certain degree of firm- ness and rigidity do not permit a farther in- crease of the body. But as we approach to old age, rigidity begins to be in excess, and the Aa2 fluids 372 PART IV. OF THE THORAX. fluids (H) bear a much less proportion to the so- lids than before. The dissipation of the body is greater than the supply of nourishment; many of the smaller vessels become gradually imper- vious (I); and the fibres losing their moisture and their elasticity, appear flaccid and wrinkled. The lilies and the roses disappear, because the fluids by which they were produced can no longer reach the extremities of the capillary vessels of the skin. As these changes take place, the nervous power being proportionably weakened, the irritability and sensibility of the body, which were formerly so remarkable, are greatly diminished; and in advanced life, the hearing, the eye-sight, and all the other senses, become gradually impaired. SECT. XVI. Of the Glands and Secretions. THE glands are commonly understood to be small, roundish, or oval bodies formed by the convolution of a great number of vessels, and destined to separate particular humours from the mass of blood. They are usually divided into two classes; but it (H) As the fluids become less in proportion to the solids, their acrimony is found to increase; and this may perhaps compensate for the want of fluidity in the blood, by dimi- nishing its cohesion. (I) In infancy, the arteries are numerous and large in respect to the veins, and the lymphatic glands are larger than at any other time of life; whereas, in old age, the ca- pacity of the venous system exceeds that of the arteries, and the lymphatic system almost disappears. 373 PART IV. OF THE THORAX. it seems more proper to distinguish three kinds of glands, viz. the mucous, conglobate, and con- glomerate. The mucous glands, or follicles, as they are most commonly called, are small cylindrical tubes continued from the ends of arteries. In some parts of the body, as in the tonsils, for example, several of these follicles may be seen folded to- gether in one common covering, and opening into one common sinus. These follicles are the vessels that secrete and pour out mucus in the mouth, œsophagus, stomach, intestines, and other parts of the body. The conglobate glands are peculiar to the lym- phatic system. Every lymphatic vein passes through a gland of this kind in its way to the thoracic duct. They are met with in different parts of the body, particularly in the axilla, groin, and mesentery, and are either solitary or in distinct clusters. The conglomerate glands are of much greater bulk than the conglobate, and seem to be an as- semblage of many smaller glands. Of this kind are the liver, kidneys, &c. Some of them, as the pancreas, parotids, &c. have a granulated ap- pearance. All these conglomerate glands are plentifully supplied with blood-vessels; but their nerves are in general very minute, and few in number. Each little granulated portion fur- nishes a small tube, which unites with other si- milar 374 PART IV. OF THE THORAX. milar ducts, to form the common excretory duct of the gland. The principal glands, and the humours they secrete, have been already described in different parts of this work; and there only remains for us to examine the general structure of the glands, and to explain the mechanism of secretion. On the first of these subjects two different systems have been formed; each of which has had, and still continues to have, its adherents. One of these systems was advanced by Malpighi, who supposed that an artery entering into a gland ra- mifies very minutely through its whole substance; and that its branches ultimately terminate in a vesicular cavity or follicle, from whence the se- creted fluid passes out through the excretory duct. This doctrine at first met with few opponents; but the celebrated Ruysch, who first attempted minute injections with wax, afterwards disputed the existence of these follicles, and asserted, that every gland appears to be a continued series of vessels, which after being repeatedly convoluted in their course through its substance, at length terminate in the excretory duct. Anatomists are still divided between these two systems : that of Malpighi, however; seems to be the best founded. The mode of secretion has been explained in a variety of ways, and they are all perfectly hy- pothetical. In such an inquiry it is natural to ask, how one gland constantly separates a parti- cular 375 PART IV. OF THE THORAX. cular humour, while another gland secretes one of a very different nature from the blood? The bile, for instance, is separated by the liver, and the urine by the kidneys. Are these secretions to be imputed to any particular dispositions in the fluids, or is their cause to be looked for in the solids? It has been supposed, that every gland con- tains within itself a fermenting principle, by which it is enabled to change the nature of the blood it receives, and to endue it with a particu- lar property. So that, according to this system, the blood, as it circulates through the kidneys, becomes mixed with the fermenting principle of those glands, and a part of it is converted into urine; and again, in the liver, in the salival and other glands, the bile, the saliva, and other juices, are generated from a similar cause. But it seems to be impossible for any liquor to be con- fined in a place exposed to the circulation, with- out being carried away by the torrent of blood, every part of which would be equally affected; and this system of fermentation has long been rejected as vague and chimerical. But as the cause of secretion continued to be looked for in the fluids, the former system was succeeded by another, in which recourse was had to the ana- logy of the humours. It was observed, that if paper is moistened with water, and oil and wa- ter are afterwards poured upon it, that the water only will be permitted to pass through it; but that, 376 PART IV. OF THE THORAX. that, on the other hand, if the paper has been previously soaked in oil instead of water, the oil only, and not the water, will be filtered through it. These observations led to a supposition, that every secretory organ is originally furnished with a humour analogous to that which it is after- wards destined to separate from the blood; and that in consequence of this disposition, the secre- tory vessels of the liver, for instance, will only admit the bilous particles of the blood, while all the other humours will be excluded. This sys- tem is an ingenious one, but the difficulties with which it abounds are unanswerable; for oil and water are immiscible; whereas the blood, as it is circulated through the body, appears to be an homogeneous fluid. Every oil will pass through a paper moistened only with one kind of oil; and wine, or spirits mixed with water, will easily be filtered through a paper previously soaked in water. Upon the same principle, all our humours, though differing in their other properties, yet agreeing in that of being perfectly miscible with each other, will all easily pass through the same filtre. But these are not all the objections to this system. The humours which are supposed to be placed in the secretory vessels for the determination of simi- lar particles of the blood, must be originally se- parated without any analogous fluid; and that which happens once, may as easily happen al- ways. Again, it sometimes happens from a vi- cious disposition, that humours are filtered through glands which are naturally not intended to 377 PART IV. OF THE THORAX. to afford them a passage; and when this once has happened, it ought, according to this system, to be expected always to do so : whereas this is not the case; and we are, after all, naturally led to seek for the cause of secretions in the solids. It does not seem right to ascribe it to any parti- cular figure of the excretory vessels; because the soft texture of those parts does not permit them to preserve any constant shape, and our fluids seem to be capable of accommodating themselves to every kind of figure, Some have imputed it to the difference of diameter in the orifices of the different secretory vessels. To this doctrine objections have likewise been raised; and it has been argued, that the vessels of the liver, for in- stance, would, upon this principle, afford a pas- sage not only to the bile, but to all the other hu- mours of less consistence with it. In reply to this objection, it has been supposed, that secon- dary vessels exist, which originate from the first, and permit all the humours thinner than the bile to pass through them. Each of these hypotheses is probably very re- mote from the truth. EXPLANATION OF PLATE XXVIII. This plate represents the Heart in situ, all the large Arteries and Veins, with some of the Muscles, &c. Muscles, &c.—SUPERIOR EXTREMITY.— a, Masseter. b, Complexus. C, Digastricus. d, Os 378 PART IV. OF THE THORAX. d, Os hyoides. e, Thyroid gland. f, Levator scapulæ. g, Cucullaris. h h, The clavicles cut. i, The deltoid muscle. k, Biceps flexor cubiti cut. l, Caraco-brachialis. m, Triceps exten- sor cubiti. n, The heads of the pronator teres, flexor carpi radiales, and flexor digitorum sub- limis, cut. o, The flexor carpi ulnaris, cut at its exteremity. p, Flexor digitorum profundus. q, Supinator radii longus, cut at its extremity. r, Ligamentum carpi transversale. s, Extensores carpi radiales. t, Latissimus dorsi. u, Anterior edge of the serratus anticus major, vv, The in- ferior part of the diaphragm. w w, Its anterior edge cut. xx, The kidneys. y, Transversus abdominis. z, Os ilium. INFERIOR EXTREMITY. a, Psoas magnus. b, Iliacus internus. c, The fleshy origin of the tensor vagina femoris. dd, The ossa pubis cut from each other. e, Musculus pectineus cut from its origin. f, Short head of the triceps ab- ductor femoris cut. g, The great head of the triceps. h, The long head cut. i, Vastus in- turnus. k, Vastus externus. l, Crureus. m, Gemellus. n, Soleus. o, Tibia. p, Peronæus longus. q, Peronæus brevis. r, Fibula. HEART and BLOOD-VESSELS.—A, The heart, with the coronary arteries and veins. B, The right auricle of the heart. C, The aorta ascen- dens. D, The left subclavian artery. E, The left carotid artery. F, The common trunk which sends off the right subclavian and right carotid arteries. ANATOMY PLATE XXVIII  379 PART IV. OF THE THORAX. arteries. G, The carotis externa. H, Arteria facialis, which sends off the coronary arteries of the lips. I. Arteria temporalis profunda. K, Aorta descendens. LL, The iliac arteries,— which send off MM, The femoral or crural ar- teries. N. B. The other arteries in this figure have the same distribution as the veins of the same name:—And generally, in the anatomi- cal plates, the description to be found on the one side, points out the same parts in the other. 1, The frontal vein. 2, The facial vein. 3, Vena temporalis profunda. 4, Vena occipitalis. 5, Vena jugularis externa. 6, Vena jugularis in- terna, covering the arteria carotis communis. 7, The vascular arch on the palm of the hand, which is formed by, 8, The radial artery and vein, and, 9, The ulnar artery and vein. 10 10, Cephalic vein. 11, Basilic vein, that on the right side cut. 12, Median vein. 13, The humeral vein, which, with the median, covers the humeral ar- tery. 14 14, The external thoracic or mam- mary arteries and veins. 15, The axillary vein, covering the artery. 1616, The subclavian veins, which, with (66) the jugulars, form, 17, The vena cava superior. 18, The cutaneous arch of veins on the fore part of the foot. 19, The vena tibialis antica, covering the artery. 20, The vena profunda femoris, covering the artery. 21, The upper part of the vena saphena major. 22, The femoral vein. 2323, The iliac veins. 24 24, Vena cava inferior. 2525, The renal veins covering the arteries. 2626, The dia- phragmatic veins. PART V. 380 Part V. OF THE BRAIN. PART V. OF THE BRAIN AND NERVES. SECT. I. Of the Brain and its Integuments. THE bones of the cranium were described in the osteological part of this work, as in- closing the brain, and defending it from external injury: but they are not its only protection; for when we make an horizontal section through these bones, we find this mass every where sur- rounded by two membranes (K), the dura and pia mater.—The first of these lines the interior surface of the cranium, to which it every where adheres strongly (L), but more particularly at the sutures, and at the many foramina through which vessels pass between it and the pericranium. The dura (K) The Greeks called these membranes meninges; but the Arabians, supposing them to be the source of all the other membranes of the body, afterwards gave them the names of dura and pia mater; by which they are now usu- ally distinguished. (L) In young subjects this adhesion is greater than in adults; but even then, in the healthy subject, it is no where easily separated, without breaking through some of the mi- nute vessels by means of which it is attached to the bone. 381 PART V. AND NERVES. dura mater (M) is perfectly smooth and inelastic, and its inner surface is constantly bedewed with a fine pellucid fluid, which every where separates it from the pia mater. The dura mater sends off several considerable processes, which divide the brain into separate portions, and prevent them from compressing each other. Of these processes there is one superior and longitudinal, called the falx, or falciform process, from its resemblance to a scythe. It arises from the spine of the os frontis, near the christa galli, and extending along in the direction of the sagittal suture, to beyond the lambdoidal suture, divides the brain into two hemispheres. A little below the lambdoidal su- ture, it divides into two broad wings or expan- sions called the transverse or lateral processes, which prevents the lobes of the cerebrum from pressing on the cerebellum. Besides these there is a fourth, which is situated under the trans- verse processes, and being continued to the spine of the occiput, divides the cerebellum into two lobes. The (M) This membrane is commonly described as consisting of two laminæ ; of which the external one is supposed to perform the office of periosteum internum to the cranium, while the internal one forms the folds and processes of the dura mater. In the natural state, however, no such separa- tion is apparent; like other membranes, we may indeed di- vide it, not into two only, but many laminæ; but this divi- sion is artificial, and depends on the dexterity of the ana- tomist. 382 PART V. OF THE BRAIN. The blood, after being distributed through the cavity of the cranium by means of the ar- teries, is returned, as in, the other parts of the body, by veins which all pass on to, certain chan- nels, situated behind these several processes. These canals or sinuses communicate with each other, and empty themselves into the internal jugular veins, which convey the blood into the vena cava. They are in fact triangular veins, running through the substance of the dura mater, and, like the processes, are distinguished into longitudinal and lateral; and where these three meet, and where the fourth process passes off, we observe a fourth sinus, which is called torcular; Herophilus, who first described it, having sup- posed that the blood at the union of these two veins, is, as it were, in a press. Besides these four canals, which were known to the ancients, modern anatomists enumerate many others, by giving the appellation of sinuses to other veins of the dura mater, which for the most part empty themselves into some of those we have just now described. There are the in- ferior longitudinal sinus, the superior and inferior petrous sinuses, the cavernous sinuses, the circu- lar sinus, and the anterior and posterior occipital sinuses. These sinuses or veins, by being conveyed through a thick dense membrane, firmly suspend- ed, as the dura mater is, within the cranium, are less liable to rupture; at the same time they 3 are 383 PART. V. AND NERVES. are well supported, and by running every where along the inner surface of the bones, they are prevented from pressing on the substance of the brain. To prevent too great a dilatation of them, we find filaments (called chordæ Willisii, from their having been first noticed by Willis) stretch- ed across their cavities; and the oblique manner in which the veins from the brain run through the substance of the brain into these channels, serves the purpose of a valve, which prevents the blood from turning back into the smaller and weaker vessels of the brain. The pia mater is a much softer and finer mem- brane than the dura mater; being exceedingly delicate, transparent, and vascular. It invests every part of the brain, and sends off an infinite number of elongations, which insinuate them- selves between the convolutions, and even into the substance of the brain. This membrane is composed of two laminæ; of which the exterior one is named tunica arachnoidea, from its thin- ness, which is equal to that of a spider's web. These two laminæ are intimately adherent to each other at the upper part of the brain, but are easily separable at the basis of the brain, and through the whole length of the medulla spinalis. The external layer, or tunica arachnoidea, appears to be spread uniformly over the surface of the brain, but without entering into its furrows as the in- ner layer does; the latter being found to insinu- ate itself between the convolutions, and even into the 384 PART V. OF THE BRAIN. the interior cavities of the brain. The blood- vessels of the brain are distributed through it in their way to that organ, and are therefore divided into very minute ramifications, before they pene- trate the substance of the brain. There are several parts included under the ge- neral denomination of brain. One of these, which is of the softest consistence, and fills the greatest part of the cavity of the cranium, is the cerebrum, or brain properly so called. Another portion, which is seated in the inferior and pos- terior part of the head, is the cerebellum; and a third, which derives its origin from both these, is the medulla oblongata. The cerebrum is a medullary mass of a mode- rate consistence, filling up exactly all the upper part of the cavity of the cranium, and divided into two hemispheres by the falx of the dura mater. Each of these hemispheres is usually dif- tinguished into an interior, a middle, and a pos- terior lobe. The first of these is lodged on the orbital processes of the os frontis; the middle lobes lie on the middle fossæ of the basis of the cranium, and the posterior lobes are placed on the transverse septum of the os occipitis, imme- diately over the cerebellum, from which they are separated by the lateral processes of the dura ma- ter. These two portions afford no distinguishing mark of separation; and on this account Haller, and many other modern anatomists, omit the distinction 385 PART V. AND NERVES. distinction of middle lobe, and speak only of the anterior and posterior lobes of the brain. The cerebrum appears to be composed of two distinct substances. Of these, the exterior one, which is of a greyish or ash-colour, is called the cortex, and is somewhat softer than the other, which is very white, and is called medulla or sub- stantia alba. After having removed the falx, and separated the two hemispheres from each other, we per- ceive a white convex body, the corpus callosum, which is a portion of the medullary substance, uniting the two hemispheres to each other, and not invested by the cortex. By making an ho- rizontal incision in the brain, on a level with this corpus callosum, we discover two oblong cavi- ties, named the anterior or lateral ventricles, one in each hemisphere. These two ventricles, which communicate with each other by a hole immediately under the plexus choroides, are se- parated laterally by a very fine medullary parti- tion, called septum lucidum, from its thinness and transparency. The lower edge of this septum is fixed to the fornix, which is a kind of medullary arch (as its name implies) situated under the cor- pus callosum, and nearly of a triangular shape. Anteriorly the fornix sends off two medullary chords, called its anterior crura; which seem to be united to each other by a portion of medulla- ry substance, named commissura anterior cerebri. These crura diverging from one another, are lost Bb at 386 PART V. OF THE BRAIN. at the other side of the lower and fore-part of the third ventricle. Posteriorly the fornix is formed into two other crura, which unite with two me- dullary protuberances called pedes hippocampi, and sometimes cornua ammonis, that extend along the back-part of the lateral ventricles. The concave edge of the pedes hippocampi is co- vered by a medullary lamina, called corpus sim- briatum. Neither the edges of the fornix, nor its poste- rior crura, can be well distinguished, till we have removed the plexus choroides. This is a pro- duction of the pia mater, which is spread over the lateral ventricles. Its loose edges are collect- ed, so as to appear like a vascular band on each side. When we have removed this plexus, we discover several other protuberances included in the lateral ventricles. These are the corpora striata, the thalami nervorum opticorum, the tubercula quadrugemina, and the pineal gland. The corpora striata are two curved oblong emi- nences, that extend along the anterior part of the lateral ventricles. They derive their name from their striated appearance, which is owing to an intermixture of the cortical and medullary sub- stances of the brain. The thalami nervorum op- ticorum, are so called, because the optic nerves arise chiefly from them, and they are likewise composed both of the cortex and medulla. They are separated from the corpora striata only by a kind 387 PART V. AND NERVES. kind of medullary chord, the geminum centrum semi-circulare. The thalami are nearly of an oval shape, and are situated at the bottom of the upper cavity of the lateral ventricles. They are closely united, and at their convex part seem to become one body. Anteriorly, in the space between the thalami, we observe an orifice by which the lateral ven- tricles communicate, and another leads down from this, under the different appellations of foramen commune anterius, vulva iter ad infundi- bulum, but more properly iter ad tertium ventri- culum; and the separation of the thalami from each other posteriorly, forms another opening or interstice called anus. This has been supposed to communicate with the third ventricle; but it does not, the bottom of it being shut up by the pia mater. The back part of the anus is formed by a kind of medullary band, which connects the thalami to each other, and is called commissura posterior cerebri. Behind the thalami and commissura posterior, we observe a small, soft, greyish, and oval body, about the size of a pea. This is the glandula pi- nealis; it is described by Galen under the name of conarion, and has been rendered famous by Descartes, who supposed it to be the seat of the soul. Galen seems formerly to have entertained the same opinion. Some modern writers have, with as little reason, imagined that the soul is placed in the corpus callosum. Bb2 The 388 PART V. OF THE BRAIN. The pineal gland rests upon four remarkable eminences, disposed in pairs, and seated imme- diately below it. These tubercles, which by the ancients were called testes and nates, have, since the time of Winslow, been more commonly nam- ed tubercula quadrugemina. Under the thalami we observe another cavity, the third ventricle, which terminates anteriorly in a small medullary canal, the infundibulum, that leads to the glandula pituitaria. It has been doubted, whether the infundibulum is really hol- low; but some late experiments on this part of the brain * by Professor Murray of Upsal, dearly prove it to be a medullary canal, surrounded by both laminæ of the pia mater. After freezing the brain, this channel was found filled with ice; and de Haen tells † us, he found it dilated, and filled with a calcareous matter (N). The soft spongy body in which the infundibu- lum terminates, was by the ancients supposed to be of a glandular structure, and destined to filter the serosity of the brain. Spigelius pretended to have discovered its excretory duct, but it seems certain that no such duct exists. It is of an oblong * Diso. de Infundibulo Cerebri. † Ratio Med. tom. vi. p. 271. (N) The under part of it, however, appears to be imper- vious; at least no injection that can be depended on has been made to pass from it into the glandula pitoitaria without lace- ration of parts. 389 PART V. AND NERVES. oblong shape, composed, as it were, of two lobes. In ruminant animals it is much larger than in man. From the posterior part of the third ven- tricle, we see a small groove or channel, de- scending obliquely backwards. This channel, which is called the aqueduct of Sylvius, though it was known to the ancients, opens into another cavity of the brain, placed between the cerebel- lum and medulla oblongata, and called the fourth ventricle. The cerebellum, which is divided into two lobes, is commonly supposed to be of a firmer texture than the cerebrum; but the truth is, that in the greater number of subjects, there ap- pears to be no sensible difference in the con- sistence of these two parts. It has more of the cortical than of the medullary substance in its composition. The furrow that divides the two lobes of the cerebellum leads anteriorly to a process, compos- ed of medullary and cortical substances, covered by the pia mater; and which, from its being di- vided into numerous furrows, resembling the rings of the earth-worm, is named processus ver- miformis. This process forms a kind of ring in its course between the lobes. The surface of the cerebellum does not af- ford those circumvolutions which appear in the cerebrum; but instead of these, we observe a great number of minute furrows, running pa- rallel 390 PART V. OF THE BRAIN. rallel to each other, and nearly in a transverse di- rection. The pia mater insinuates itself into these furrows. When we cut into the substance of the cere- bellum, from above downwards, we find the medullary part running in a kind of ramifying course, and exhibiting an appearance that has gotten the name of arbor vitæ. These ramifi- cations unite to form a medullary trunk; the the middle, anterior, and most considerable part of which forms two processes, the crura cerebel- li, which unite with the crura cerebri, to form the medulla oblongata. The last furnishes two other processes, which lose themselves under the nates, and thus unite the lobes of the cerebellum to the posterior part of the cerebrum. Under the nates we observe a transverse medullary line, or linea alba, running from one of these processes to the other; and between them we find a very thin medullary lamina, covered with the pia ma- ter, which the generality of anatomists have (though seemingly without reason) considered as a valve formed for closing the communica- tion between the fourth ventricle and the aquæ- ductus Sylvii. Vieussens named it valvula ma- jor cerebri. The medulla oblongata is situated in the mid- dle, lower, and posterior part of the cranium, and may be considered as a production or con- tinuation of the whole medullary substance of the cerebrum and cerebellum, being formed by the 391 PART V. AND NERVES. the of two considerable medullary processes of the cerebrum, called crura cerebri, with two other smaller ones from the cerebellum, which were just now spoken of under the name or crura cerebelli. The crura cerebri arise from the middle and lower part of each hemisphere. They are sepa- rated from each other at their origin, but are united below, where they terminate in a middle protuberance, the pons Varolii, so called, because Varolius compared it to a bridge. This name, however, can convey no idea of its real appear- ance. It is, in fact, nothing more than a me- dullary protuberance, nearly of a semi-spherical shape, which unites the crura cerebri to those of the cerebellum. Between the crura cerebri, and near the ante- rior edge of the pons Varolii, are two tubercles, composed externally of medullary, and internal- ly of cineritious substance, to which Eustachius first gave the name of eminentiæ mamillares. Along the middle of the posterior surface of the medulla oblongata, where it forms the ante- rior part of the fourth ventricle, we observe a kind of furrow which runs downwards and ter- minates in a point. About an inch above the lower extremity of this fissure, several medullary filaments are to be seen running towards it on each side in an oblique direction, so as to give it the appearance of a writing-pen; hence it is called calamus scriptorius. From 392 PART V. OF THE BRAIN. From the posterior part of the pons Varolii, the medulla oblongata descends obliquely back- wards ; at its fore-part, immediately behind the pons Varolii, we observe two pair of eminences, which were described by Eustachius, but receiv- ed no particular appellation till the time of Vi- eussens, who gave them the names of corpora olivaria and corpora pyramidalia. The former are the outermost, being placed one on each side. They are nearly of an oval shape, and are com- posed of medulla, with streaks of cortical sub- stance. Between these are the corpora pyra- midalia, each of which terminates in a point. In the human subject these four eminences are sometimes not easily distinguished. The medulla spinalis, or spinal marrow, which is the name given to the medullary chord that is extended down the vertebral canal, from the great foramen of the occipital bone to the bot- tom of the last lumbar vertebra, is a continua- tion of the medulla oblongata. Like the other parts of the brain, it is invested by the dura and pia mater. The first of these, in its passage out of the cranium, adheres to the foramen of the os occipitis. Its connection with the ligamentary substance that lines the cavity of the spine, is on- ly by means of cellular membrane; but between the several vertebræ, where the nerves pass out of the spine, it sends off prolongations, which adhere strongly to the vertebral ligaments. Here, as in the cranium, the dura mater has its sinuses or 393 PART V. AND NERVES. or large veins. These are two in number, and are seen running on each side of the medullary column, from the foramen magnum of the os occipitis to the lower part of the os sacrum. They communicate together by ramifying branches at each vertebra, and terminate in the vertebral, intercostal, and sacral veins. The pia mater is connected with the dura ma- ter by means of a thin transparent substance, which from its indentations between the spinal nerves has obtained the name of ligamentum den- ticulatum. It is somewhat firmer than the tu- nica arachnoidea, but in other respects resem- bles that membrane. Its use is to support the spi- nal marrow, that it may not affect the medulla oblongata by its weight. The spinal marrow itself is externally of a white colour; but upon cutting into it we find its middle-part composed of a darker coloured mass, resembling the cortex of the brain. When the marrow has reached the first lumbar verte- bra, it becomes extremely narrow, and at length terminates in an oblong protuberance; from the extremity of which the pia mater sends off a prolongation or ligament, resembling a nerve, that perforates the dura mater, and is fixed to the os coccygis. The medulla spinalis gives rise to 30 or 31 pair of nerves, but they are not all of the same size, nor do they all run into in the same direc- tion. The upper ones are thinner than the rest, and 394 PART V. OF THE BRAIN .. and are placed almost transversely: as we descend we find them running more and more obliquely downwards, till at length their course is almost perpendicular, so that the lowermost nerves ex- hibit an appearance that is called cauda equina, from its resemblance to a horse's tail. The arteries that ramify through the different parts of the brain, are derived from the inter- nal carotid and from the vertebral arteries. The medulla spinalis is supplied by the anterior and posterior spinal arteries, and likewise receives branches, from the cervical, the inferior and su- perior intercostal, the lumbar, and the sacral ar- teries. SECT. II. Of the Nerves. THE nerves are medullary chords, differing from each other in size, colour, and consistence, and deriving their origin from the medulla ob- longata and medulla spinalis. There are 39, and sometimes 40, pair of these nerves; nine (O) of which originate from the medulla oblongata, and 30 or 31 from the medulla spinalis. They ap- pear to be perfectly inelastic, and likewise to pos- sess (O) It has been usual to describe the ten pair of nerves as arising from the medulla oblongata; but as the tenth pair arise in the same manner as the other spinal nerves, Santorini, Heister, Haller, and others, seem very properly to have classed them among the nerves of the spine. 395 PART V. AND NERVES. sess no irritability. If we irritate muscular fi- bres, they immediately contract; but nothing of this sort happens if we irritate a nerve. They carry with them a covering from the pia mater; but derive no tunic from the dura mater, as hath been generally, though erroneously, supposed, ever since the time of Galen (P), the outer cover- ing of the nerves being in fact nothing more than the cellular membrane. This covering is very thick where the nerve is exposed to the action of muscles; but where it runs through a bony canal, or is secure from pressure, the cel- lular tunic is extremely thin, or altogether want- ing. We have instances of this in the portio mollis of the auditory nerve, and in the nerves of the heart. By elevating, carefully and gently, the brain from the basis of the cranium, we find the first nine pair arising in the following order: 1. The nervi olfactorii, distributed through the pituita- ry membrane, which constitutes the organ of smell. 2. The optici, which go to the eyes, where they receive the impressions of visible objects. 3. The oculorum motores, so called because they are distributed to the muscles of the eye. 4. The (P) Baron Haller and Professor Zinn seem to have been the first who demonstrated, that the dura mater is reflected upon and adheres to the periosteum at the edges of the fora- mina that afford a passage to the nerves out of the cranium, and vertebral canal, or is soon lost in the cellular substance, 396 PART V. OF THE BRAIN. The pathetici, distributed to the superior oblique muscles of the eyes, the motion of which is expressive of certain passions of the soul. 5. The nerves of this pair soon divide into three prin- cipal branches, and each of these has a different name. Its upper division its he ophthalmicus, which is distributed to various parts of the eyes, eye-lids, forehead, nose, and integu- ments of the face. The second is called the maxillaris superior, and the third maxillaris in- ferior; both of which names allude to their dis- tribution. 6. The abductores; each of these nerves is distributed to the abductor muscle of the eye, so called, because it helps to draw the globe of the eye, from the nose. 7. The auditorii (Q), which are distributed through the organs of hearing. 8. The par vagum, which derives its name from the great number of parts to which it gives branches both in the tho- rax and abdomen. 9. The linguales, or hypo- glossi, which are distributed to the tongue, and appear (Q) This pair, soon after its entrance into the meatus audi- torius internus, separates into two branches. One of these is of a very soft and pulpy consistence, it is called the portio mollis of the seventh pair, and is spread over the inner part of the ear. The other passes out through the aqueduct of Fallopius in a firm chord, which is distinguished as the por- tio dura, and is distributed to the external ear and other parts of the neck and face. 397 PART V. AND NERVES. appear to contribute both to the organ of taste and to the motions of the tongue (R). It has already been observed, that the spinal marrow sends off 30 or 31 pair of nerves; these are chiefly distributed to the exterior parts of the trunk and to the extremities. They are com- monly distinguished into the cervical, dorsal, lumbar, and sacral nerves. The cervical, which pass out from between the several vertebræ of the neck, are eight (S) in number; the dorsal, twelve; the lumbar five; and the sacral, five or six; the number of the latter depending on the number of holes in the os sacrum. Each spinal nerve at its origin is composed of two fasciculi of medullary fibres. One of these fasciculi arises from the anterior, and the other from the pos- terior surface of the medulla. These fasciculi are (R) Heister has summed up the uses of these nine pair of nerves in the two following Latin verses : "Olfaciens, cernens, oculosque novens, patiensque, "Gastans, abducens, audiensque, vagansque, loquensque." (S) Besides these, there is another pair called accessorii, which arises from the medulla spinalis at its beginning; and ascending the great foramen of the os occipitis into the cranium, passes out again close to the eighth pair, with which, however, it does not unite; and it is afterwards distributed chiefly to the muscles of the neck, back, and scapula. In this course it sends off filaments to different parts, and likewise communicates with several other nerves. Physiologists are at a loss how to account for the singular origin and course of these nervi accessorii. The indents considered them as a branches of the eighth pair, dis- tributed to muscles of the scapula: Willis likewise considered them as appendages to that pair, and on that account named them accessorii. They are sometimes called the spinal pair; but as this latter name is applicable to all the nerves of the spine indiscriminately, it seems better to adapt that given by Willis. 398 PART V. OF THE BRAIN. are separated by the ligamentum denticulatum; after which we find them contiguous to one ano- ther. They then perforate the dura mater, and unite to form a considerable knot or ganglion. Each of these ganglions sends off two branches; one anterior, and the other posterior. The an- terior branches communicate with each other at their coming out of the spine, and likewise send off one, and sometimes more branches, to assist in the formation of the intercostal nerve. The knots or ganglions of the nerves just now spoken of, are not only to be met with at their exit from the spine, but likewise in various parts of the body. They occur in nerves of the medulla oblongata, as well as in those of the spine. They are not the effects of dis- ease, but are to be met with in the same parts of the same nerves, both in the fœtus and adult. They are commonly of an oblong shape, and of a greyish colour, somewhat inclined to red, which is perhaps owing to their being extreme- ly vascular. Internally we are able to distinguish something like an intermixture of the nervous filaments. Some writers have considered them as so ma- ny little brains; Lancisi fancied he had disco- vered muscular fibres in them, but they are cer- tainly not of an irritable nature. A late writer, Dr. Johnstone*, imagines they are intended to deprive us of the power of the will over cer- tain * Essay on the Use of the Ganglions of the Nerves. 399 PART V. AND NERVES. tain parts, as the heart, for instance: but if this hypothesis were well founded, we should meet with them only in the nerves leading to involun- tary muscles; whereas it is certain, that the vo- luntary muscles receive their nerves through gan- glions. Doctor Monro, from observing the ac- curate intermixture of the minute nerves which compose them, considers them as new sources of nervous energy †. The nerves, like the blood-vessels, in their course through the body, communicate with each other; and each of these communications constitutes what is called a plexus, from whence branches are again detached to different parts of the body. Some of these are constant and con- siderable enongh to be distinguished by particu- lar names, as the semilunar plexus; the pulmo- nary plexus; the hepatic, the cardiac, &c. It would be foreign to the purpose of this work, to follow the nerves through all their dis- tributions; but it may be remembered, that in describing the different viscera, mention was made of the nerves distributed to them. There is one pair, however, called the intercostal or great sympathetic nerve, which seems to require particular notice, because it has an almost uni- versal connection and correspondence with all the other nerves of the body. Authors are not perfectly agreed about the origin of the intercos- tal; † Observations on the Nervous system. 400 PART V. OF THE BRAIN. tal; but it may perhaps not improperly be de- scribed, as beginning from filaments of the fifth and sixth pair; it then passes out of the cranium, through the bony canal of the carotid, from whence it descends laterally close to the bodies of the vertebræ, and receives branches from al- most all the vertebral nerves; forming almost as many ganglions in its course through the thorax and abdomen. It sends off an infinite number of branches to the viscera in those cavities, and forms several plexuses with the branches of the eight pair or par vagum. That the nerves are destined to convey the principles of motion and sensibility to the brain from all parts of the system, there can be no doubt; but how these effects are produced, no one has ever yet been able to to determine. The inquiry has been a constant source of hypothe- sis in all ages, and has produced some ingenious ideas, and many erroneous positions, but with- out having hitherto afforded much satisfactory information. Some physiologists have considered a trunk of nerves as a solid chord, capable of being divided into an infinite number of filaments, by means of which the impressions of feeling are convey- ed to the sensorium commune. Others have supposed it to be a canal, which afterwards se- parates into more minute channels; or, perhaps, as being an assemblage of many very small and distinct tubes, connected to each other, and thus 3 forming 401 PART V. AND NERVES. forming a cylindrical chord. They who con- tend for their being solid bodies, are of opinion that feeling is occasioned by vibration; so that, for instance, according to this system, by prick- ing the finger, a vibration would be occasioned in the nerve, distributed through its substance; and the effects of this vibration, when extended to the sensorium, would be an excital of pain. But the inelasticity, the softness, the connection, and the situation of the nerves, are so many proofs that vibration has no share in the cause of feeling. Others have supposed, that in the brain and spinal marrow, a very subtile fluid is secreted, and from thence conveyed through the imper- ceptible tubes, which they consider as existing in the nerves. They have farther supposed, that this very subtile fluid, to which they have given the name of animal spirits, is secreted in the cortical substance of the brain and spinal marrow, from whence it passes through the me- dullary substance. This, like the other system, is founded altogether on hypothesis; but it seems to be an hypothesis derived from much more probable principles, and there are many ingenious arguments to be brought in its sup- port. Cc EXPLA- 402 PART V. OF THE BRAIN. EXPLANATION OF PLATE XXIX. FIG. 1. Represents the inferior part of the brain;—the Anterior part of the whole Spine, including the Medulla Spinalis;—with the ori- gin and large portions of all the NERVES. A A, The anterior lobes of the cerebrum. B B, The lateral lobes of the cerebrum. CC, The two lobes of the cerebellum. D, Tuber annulare. E, The passage from the third ven- tricle to the infundibulum. F, The medulla ob- longata, which sends off the medulla spinalis through the spine. G G, That part of the os occipitis which is placed above (HH) the trans- verse processes of the first cervical vertebra. II, &c. The seven cervical vertebræ, with their interme- diate cartilages. KK, &c. The twelve dorsal vertebræ, with their intermediate cartilages. LL, &c. The five lumbar vertebræ, with their in- termediate cartilages. M, The os sacrum. N, The os coccygis. NERVES.—11, The first pair of nerves, named olfactory, which go to the nose. 22, The second pair, named optic, which goes to form the tunica retina of the eye. 33, The third, named motor oculi; it supplies most of the mus- cles of the eye-ball. 44, The fourth pair, named pathetic,—which is wholly spent upon the musculus trochlearis of the eye. 55, The fifth 403 PART V. AND NERVES. fifth pair divides into three branches.—The first, named ophthalmic, goes to the orbit, supplies the lachrymal gland, and sends branches out to the forehead and nose.—The second, named supe- rior maxillary, supplies the teeth of the upper jaw, and some of the muscles of the lips.—The third named inferior maxillary, is spent upon the muscles and teeth of the lower jaw, tongue, and muscles of the lips. 66, The sixth pair, which, after sending off the beginning of the intercos- tal or great sympathetic, is spent upon the ab- ductor oculi. 77, The seventh pair, named au- ditory, divides into two branches.—The largest, named portio mollis, is spent upon the internal ear.—The smallest, portio dura, joins to the fifth pair within the internal ear by a reflected branch from the second of the fifth; and with- in the tympanum, by a branch from the third of the fifth, named chorda tympani.—Vid. fig. 3. near B. 88, &c. The eighth pair, named par vagum,—which accompanies the intercostal, and is spent upon the tongue, larynx, pharynx, lungs, and abdominal viscera. 99, The ninth pair, which are spent upon the tongue. 1010, &c. The intercostal, or great sympathetic, which is seen from the sixth pair to the bottom of the pelvis on each side of the spine, and joining with all the nerves of the spine;—in its progress supplying the heart, and, with the par vagum, the contents of the abdomen and pelvis. 11, 11, The accessorius, which is spent upon Cc2 the 404 PART V. OF THE BRAIN. the sternocleido-mastoidæus and trapezius mus- cles. 12 12, The first cervical nerves;—1313, The second cervical nerves;—both spent up- on the muscles that lie on the neck, and te- guments of the neck and head. 1414, The third cervical nerves, which, after sending off (15 15, &c.) the phrenic nerves to the diaphragm, supply the muscles and teguments that lie on the side of the neck and top of the shoulder. 16 16, The brachial plexus, formed by the fourth, fifth, sixth, seventh cervicals, and first dorsal nerves. —which supply the muscles and teguments of the superior extremity. 1717, The twelve dorsal, or proper intercostal nerves, which are spent upon the intercostal muscles and some of the large muscles which lie upon the thorax. 1818, The five lumbar pairs of nerves, which supply the lumbar and abdominal muscles, and some of the teguments and muscles of the infe- rior extremity. 1919, The sacro-sciatic, or posterior crural nerve, formed by the two infe- rior lumbar, and three superior of the os sacrum. This large nerve supplies the greatest part of the muscles and teguments of the inferior extremity. 20, The stomachic plexus, formed by the eighth pair. 2121, Branches of the solar or cæliac plexus, formed by the eighth pair and intercos- tals, which supply the stomach and chylopoietic viscera. 2222, Branches of the superior and inferior mesenteric plexuses, formed by the eighth pair and intercostals, which supply the chylo- poietic viscera, with part of the organs of urine and ANATOMY PLATE XXIX  405 PART V. AND NERVES. and generation. 2323, Nerves which accom- pany the sphermatic chord. 2424, The hypo- gastric plexus, which supplies the organs of u- rine and generation within the pelvis. FIG. 2, 3, 4, 5. Shows different Views of the Inferior part of the Brain, cut perpendicular- ly through the Middle,—with the origin and large Portions of the Nerves which pass out through the bones of the Cranium,—and the three first Cervical. A, The anterior lobe. B, The lateral lobe of the cerebrum. C, One of the lobes of the ce- rebellum. D, Tuber annulare. E, Corpus py- ramidale, in the middle of the medulla oblon- gata. F, The corpus olivare, in the side of the medulla oblongata. G, The medulla oblongata. H, The medulla spinalis. Nerves.—1 2 3 4 5 6 7 8 and 9, Pairs of nerves. 1010, Nervus accessorius, which comes from —11, 12, and 13, The three first cervical nerves. PART VI. 406 PART VI. OF THE SENSE. PART VI. O THE SENSES, AND THEIR ORGANS. IN treating of the senses, we mean to confine ourselves to the external ones of touch, taste, smelling, hearing, and vision. The word sense, when applied to these five, seems to imply not only the sensation excited in the mind by certain impressions made on the body, but likewise the organ destined to receive and transmit these im- pressions to the sensorium. Each of these or- gans being of a peculiar structure, is suscepti- ble only of particular impressions, which will be pointed out as we proceed to describe each of them separately. SECT. I. Of Touch. THE sense of touch may be defined to be the faculty of distinguishing certain properties of bo- dies by the feel. In a general acceptation, this definition might perhaps not improperly be ex- tended to every part of the body possessed of sensibility (T), but it is commonly confined to the (T) In the course of this article, mention has often been made of the sensibility or insensibility of different parts of the body; it will therefore, perhaps, not be amiss to observe 407 PART VI. AND THEIR ORGANS. the nervous papillæ of the cutis, or true skin which, with its appendages, and their several uses, have been already described. The exterior properties of bodies, men as their solidity, moisture, inequality, smoothness, dryness, or fluidity, and likewise their degree of heat, seem all to be capable of making different impressions on the papillæ, and consequently of exciting different ideas in the sensorium com- mune. But the organ of touch, like all the other senses, is not equally delicate in every part of the body, or in every subject ; being in some much more exquisite than it is in others. SECT. in this place, that many parts which were formerly suppos- ed to possess the most exquisite sense. are now known to have but little or no feeling, at least in a sound state: for in an inflamed state, even the bones, the most insensible parts of any, become susceptible of the most painful sensations. This curious discovery is due to the late Baron Haller. His experiments prove, that the bones, cartilages, ligaments, tendons, epidermis and membranes (as the pleura, pericar- dium, dura and pia mater, periosteum, &c.) may in a healthy state be considered as insensible. As sensibility depends on the brain and nerves, of course different parts will possess a greater or less degree of feeling in proportion as they are supplied with a greater or smaller number or nerves. Upon the principle it is, that the skin, muscles, stomach, intestines, urinary bladder, ureters, uterus, vagina, penis, tongue, and retina are extremely sensible while the lungs and glands have only an obscure degree of feeling. 408 PART VI. OF THE SENSES. SECT. II. Of the Taste. The sense of taste is seated chiefly in the tongue; the situation and figure of which are sufficiently known. On the upper surface of this organ we may observe a great number of papillæ, which, on account of their difference in size and shape, are commonly divided into three classes. The largest are situated towards the basis of the tongue. Their number commonly varies from seven to nine, and they seem to be mucous fol- licles. Those of the second class are somewhat smaller, and of a cylindrical shape. They are most numerous about the middle of the tongue. Those of the third class are very minute, and of a conical shape. They are very numerous on the apex and edges of the tongue, and have been supposed to be formed by the extremities of its nerves. We observe a line, the linea linguæ mediana, running along the middle of the tongue, and di- viding it as it were into portions. Towards the basis of the tongue, we meet with a little cavity, named by Morgagni foramen cæcum, which seems to be nothing more than a common termi- nation of some of the excretory ducts of mu- cous glands situated within the substance of the tongue. We 409 PART VI. AND THEIR ORGANS. We have already observed, that this organ is every where covered by the cuticle, which, by forming a reduplication, called the frænum, at its under part, serves to prevent the too great motion of the tongue, and to fix in it its situation. But, besides this attachment, the tongue is con- nected by means of its muscles and membranous ligaments, to the lower jaw, the os hyoides, and the styloid processes. The principal arteries of the tongue are the linguales, which arise from the external carotid. Its veins empty themselves into the external ju- gulars. Its nerves arise from the fifth, eighth, and ninth, pair. The variety of tastes seems to be occasioned by the different impressions made on the papil- læ by the food. The different state of the pa- pillæ with respect to their moisture, their figure, or their covering, seems to produce a considera- ble difference in the taste, not only in diffe- rent people, but in the same subject, in sickness and in health. The great use of the taste seems to be to enable us to distinguish wholesome and salutary food from that which is unhealthy ; and we observe that many quadrupeds, by hav- ing their papillæ (U) very large and long, have the (U) Malpighi's description of the papillæ, which has been copied by many anatomical writers, seems to have been taken chiefly from the tongues of sleep. 410 PART VI. OF THE SENSES. the faculty of distinguishing flavours with infi- nite accuracy. SECT III. Of Smelling. THE sense of smelling, like the sense of taste, seems intended to direct us to a proper choice of aliment, and is chiefly seated in the nose, which is distinguished into its external and internal parts. The situation and figure of the former of these do not seem to require a definition. It is compos- ed of bones and cartilages, covered by muscu- lar fibres and by the common teguments. The bones make up the upper portion, and the car- tilages the lower one. The septum narium, like the nose, is likewise in part bony, and in part cartilaginous. These bones and their con- nexions were described in the osteology. The internal part of the nose, besides the ossa spongiosa, has six cavities or sinuses, the max- illary, the frontal, and the sphenoid, which were all described with the bones of the head. They all open into the nostrils; and the nose likewise communicates with the mouth, larynx, and pharynx, posteriorly behind the velum palati. All these several parts, which are included in the internal division of the nose, viz. the inner surface of the nostrils, the lamellæ of the ossa spongiosa, and the sinuses, are lined by a very vascular membrane, which though not unknown to the ancients, was first well described by Schneider 411 PART VI. AND THEIR ORGANS. Schneider *, and is therefore now commonly named membrana pituitaria Schneideri. This membrane is truly the organ of smelling; but its real structure does not yet seem to be perfect- ly understood. It appears to be a continuation of the cuticle, which lines the inner surface of the mouth, in some parts of the nose it is smooth and firm, and in others it is loose and spongy. It is constantly moistened by a mu- cous secretion; the finer parts of which are carried off by the air we breathe, and the re- mainder, by being retained in the sinuses, ac- quires considerable consistence. The manner in which this mucus is secreted has not yet been sa- tisfactorily ascertained; but it seems to be by means of mucous follicles. Its arteries are branches of the internal max- illary and internal carotid. Its veins empty themselves into the internal jugulars. The first pair of nerves, the olfactory, are spread over every part of it, and it likewise receives branches from the fifth pair. After what has been said of the pituitary mem- brane, it will not be difficult to conceive how the air we draw in at the nostrils, being im- pregnated with the effluvia of bodies, excites in us that kind of sensation we call smelling. As these effluvia, from their being exceedingly light and volatile, cannot be capable in a small quantity De Catarrho, lib. iii. 412 PART VI. OF THE SENSES. quantity of making any great impression on the extremities of the olfactory nerves, it was ne- cessary to give considerable extent to the pituitary membrane, that by this means a greater num- ber of odoriferous particles might be admitted at the same time. When we wish to take in much of the effluvia of any thing, we naturally close the mouth, that all the air we inspire may pass through the nostrils; and at the same time, by means of the muscles of the nose, the nos- trils are dilated, and a greater quantity of air is drawn into them. In many quadrupeds, the sense of smelling is much more extensive and delicate than it is in the human subject; and in the human subject it seems to be more perfect the less it is vitiated by a variety of smells. It is not always in the same state of perfection, being naturally affect- ed by every change of the pituitary membrane, and of the lymph with which that membrane is moistened. SECT. IV. Of Hearing. BEFORE we undertake to explain the manner in which we are enabled to receive the impressi- ons of sound, it will be necessary to describe the ear, which is the organ of hearing. It is com- monly distinguished into external and internal. The former of these divisions includes all that we are able to discover without dissection, and the 413 PART VI. AND THEIR ORGANS the meatus auditorius, as far as the typanum; and the latter, all the other parts of the ear. The external ear is a cartilaginous funnel co- vered by the common integuments, and attach- ed by means of its ligaments and muscles, to the temporal bone. Although capable only of a very obscure motion, it is found to have seve- ral muscles. Different parts of it are distinguish- ed by several names; all its cartilaginous part is called ala or wing, to distinguish it from the soft and pendent part below, called the lobe. Its outer circle or border is called helix, and the se- micircle within this, antihelix. The moveable cartilage placed immediately before the meatus auditorius, which it may be made to close ex- actly, is named tragus; and an eminence op- posite to this at the extremity of the antihelix, is called antitragus. The concha is a consider- able cavity formed by the extremities of the he- lix and antihelix. The meatus auditorius, which at its opening is cartilaginous, is lined with a very thin membrane, which is a conti- nuation of the cuticle from the surface of the ear. In this canal we find a yellow wax, which is secreted by a number of minute glands or folli- cles, each of which has an excretory duct. This secretion; which is at first of an oily consistence, defends the membrane of the typanum from the injuries of the air; and by its bitterness, pre- vents minute insects from entering into the ear. But 414 PART VI. OF THE SENSES. But when from neglect or disease it accumulates in too great a quantity, it sometimes occasions deafness. The inner extremity of the meatus is closed by a very thin transparent membrane, the membrana tympani, which is set in a bony cir- cle like the head of a drum. In the last century Rivinus, professor at Leipsic, fancied he had discovered a hole in this membrane, surrounded by a sphincter, and affording a passage to the air, between the external and internal ear. Cow- per, Heister, and some other anatomists, have admitted this supposed foramen, which certain- ly does not exist. Whenever there is any open- ing in the membrana tympani, it may be consi- dered as accidental. Under the membrana tym- pani runs a branch of the fifth pair of the nerves, called chorda tympani; and beyond this mem- brane is the cavity of the tympanum, which is about seven or eight lines wide, and half so ma- ny in depth; it is semispherical, and every where lined by a very fine membrane. There are four openings to be observed in this cavity. It communicates with the mouth by means of the Eustachian tube. This canal, which is in part bony and in part cartilaginous, begins by a very narrow opening at the anterior and almost superior, part of the tympanum, increasing in size as it advances towards the palate of the mouth, where it terminates by an oval opening. This tube is every where lined by the same mem- brane that covers the inside of the mouth. The real 415 PART VI. AND THEIR ORGANS. real use of this canal does not seem to have been hitherto satisfactorily ascertained; but sound would seem to be conveyed through it to the membrana tympani, deaf persons being often observed to listen attentively with their mouths open. Opposite to this is a minute passage, which leads to the sinuosities of the mastoid pro- cess; and the two other openings, which are in the internal process of the os petrosum, are the fenestra ovalis, and fenestra rotunda, both of which are covered by a very fine membrane. There are three distinct bones in the cavity of the tympanum; and these are the malleus, incus, and stapes. Bedsides these there is a fourth, which is the os orbiculare, considered by some anatomists as a process of the stapes, which is necessarily broken off by the violence we are obliged to use in getting at these bones; but when accurately considered, it seems to be a distinct bone. The malleus is supposed to resemble a ham- mer, being larger at one extremity, which is its head, than it is at the other, which is its han- dle. The latter is attached to the membrana tympani, and the head of the bone is articulat- ed with the incus. The incus, as it is called from its shape, though it seems to have less resemblance to an anvil than to one of the dentes molares with its roots wide- ly separated from each other, is distinguished in- to its body and its legs. One of its legs is plac- 3 ed 416 PART VI. OF THE SENSES. ed at the entry of the canal which leads to the mastoid process; and the other, which is some- what longer, is articulated with the stapes, or rather with the os orbiculare, which is placed between them. The third bone is very properly named stapes, being perfectly shaped like a stirrup. Its basis is fixed into the fenestra ovalis, and its upper part is articulated with the os orbiculare. What is called the fenestra rotunda, though perhaps improperly, as it is more oval than round, is observed a little above the other, in an emi- nence formed by the os petrosum, and is closed by a continuation of the membrane that lines the inner surface of the tympanum. The stapes and malleus are each of them furnished with a little muscle, the stapedeus and tensor tym- pani. The first of these, which is the smallest in the body, arises from a little cavern in the pos- terior and upper part of the cavity of the tym- panum; and its tendon, after passing through a hole in the same cavern, is inserted at the back part of the head of the stapes. This mus- cle, by drawing the stapes obliquely upwards, as- sists in stretching the membrana tympani. The tensor tympani (X,) or internus malei, as it is called by some writers, arises from the carti- (X) Some anatomists describe three muscles of the mal- leus: but only this one seems to deserve the name of muscle; what are called the externus and obliquus mollei, seeming to be ligaments rather than muscles. 417 PART VI. AND THEIR ORGANS. cartilaginous extremity of the Eustachian tube, and is inserted into the back part of the handle of the malleus, which it serves to pull inwards, and of course helps to stretch the membrana tym- pani. The labyrinth is the only part of the ear which remains to be described. It is situated in the os petrosum, and is separated from the tympanum by a partition which is every where bony, except at the two fenestræ. It is composed of three parts; and these are the vestibulum, the semicir- cular canals, and the cochlea. The vestibulum is an irregular cavity, much smaller than the tympanum, situated nearly in the centre of the os petrosum, between the tym- panum, the cochlea, and the semicircular canals. It is open on the side of the tympanum by means of the fenestra ovalis, and communicates with the upper portion of the cochlea by an oblong foramen, which is under the fenestra ovalis, from which it is separated only by a very thin parti- tion. Each of the three semicircular canals forms about half a circle of nearly a line in diameter, and running each in a different direction, they are distinguished into vertical, oblique, and hori- zontal. These three canals open by both their extremities into the vestibulum; but the vertical and the oblique being united together at one of their extremities, there are only five orifices to be seen in the vestibulum. Dd The 418 PART VI. OF THE SENSES. The cochlea is a canal which takes a spiral course, not unlike the shell of a snail. From its basis to its apex it makes two turns and a half; and is divided into two canals by a very thin la- mina or septum, which is in part bony and in part membranous, in such a manner that these two canals only communicate with each other at the point. One of them opens into the vestibu- lum, and the other is covered by the membrane that closes the fenestra rotunda. The bony la- mella which separates the two canals is exceed- ingly thin, and fills about two thirds of the dia- meter of the canal. The rest of the septum is composed of a most delicate membrane, which lines the whole inner surface of the cochlea, and seems to form this division in the same man- ner as the two membranous bags of the pleura, by being applied to each other, form the medias- tinum. Every part of the labyrinth is furnished with a very delicate periosteum, and filled with a wa- tery fluid, secreted as in other cavities. This fluid transmits to the nerves the vibrations it re- ceives from the membrane closing the fenestra ro- tunda, and from the basis of the stapes, where it rests on the fenestrum ovale. When this fluid is collected in too great quantity, or is compressed by the stapes, it is supposed to escape through two minute canals or aqueducts, lately described by 419 PART VI. AND THEIR ORGANS. by Dr Cotunni *, an ingenious physician at Na- ples. One of these aqueducts opens into the bottom of the vestibulum, and the other into the cochlea, near the fenestra rotunda. They both pass through the os petrosum, and com- municate with the cavity of the cranium where the fluid that passes through them is absorbed; and they are lined by a membrane which is sup- posed to be a production of the dura mater. The arteries of the external ear come from the temporal and other branches of the external carotid, and its veins pass into the jugular. The internal ear receives branches of arteries from the basilary and carotids, and its veins empty them- selves into the sinuses of the dura mater, and into the internal jugular. The portio mollis of the seventh pair is distributed through the cochlea, the vestibu- lum, and the semi-circular canals; and the portio dura sends off a branch to the tympa- num, and other branches to the external ear and parts near it. The sense of hearing, in producing which all the parts we have described assist, is occasioned by a certain modulation of the air collected by the funnel-like shape of the external ear, and con- veyed through the meatus auditorius to the mem- brana tympani. That sound is propagated by means of the air, is very easily proved by ring- Dd2 ing * De aquæductibus Auris Humanæ Internæ, 8ve, 1760. 420 PART VI. OF THE SENSES. ing a bell under the receiver of an air-pump; the sound it affords being found to diminish gra- dually as the air becomes exhausted, till at length it ceases to be heard-at all. Sound moves through the air with infinite velocity; but the degree of its motion seems to depend on the state of the air, as it constantly moves faster in a dense and dry, than it does in a moist and rarefied air. See Acoustics, n° 20. That the air vibrating on the membrana tym- pani communicates its vibration to the different parts of the labyrinth, and by means of the fluid contained in this cavity affects the auditory nerve so as to produce sound, seems to be very probable : but the situation, the minuteness, and the variety of the parts which compose the ear, do not permit much to be advanced with certain- ty concerning their mode of action. Some of these parts seem to constitute the im- mediate organ of hearing, and these are all the parts of the vestibulum: but there are others which seem intended for the perfection of this sense, without being absolutely essential to it. It has happened, for instance, that the membrana tympani, and the little bones of the ear, have been destroyed by disease, without depriving the patient of the sense of hearing (Y). Sound (Y) This observation has led to a supposition, that a per- foration of this membrane may in some cases of deafness be useful; and Mr. Cheselden relates, that, some years ago, a 421 Part VI. AND THEIR ORGANS. Sound is more or less loud in proportion to the strength of the vibration; and the variety of sounds seems to depend on the difference of this vibration; for the more quick and frequent it is, the more acute will be the sound, and vice versa. Before we conclude this article, it will be right to explain certain phenomena, which will be found to have a relation to the organ of hearing. Every body has, in consequence of particular sounds, occasionally felt that disagreeable sensa- tion which is usually called setting the teeth on edge: and the cause of this sensation may be traced to the communication which the portio dura of the auditory nerve has with the branches of the fifth pair that are distributed to the teeth, being probably occasioned by the violent tremor produced in the membrana tympani by these ve- ry acute sounds. Upon the same principle we may explain the strong idea of sound which a person has who holds a vibrating string between his teeth. The humming which is sometimes perceived in the ear, without any exterior cause, may be occasioned either by an increased action of the arteries in the ears, or by convulsive contractions of malefactor was pardoned on condition that he should submit to this operation; but the public clamour raised against it was so great, that it was thought right not to perform it. 422 PART VI. OF THE SENSES. of the muscles of the malleus and stapes, affecting the auditory nerve in such a manner as to produce the idea of found. An ingenious philosophical writer * has lately discovered, that there are sounds liable to be excited in the ear by irritation, and without any assistance from the vibrations of the air. SECT. V. Of Vision. The eyes, which constitute the organ of vi- sion, are situated in two bony cavities named orbits, where they are surrounded by several parts, which are either intended to protect them from external injury, or to assist in their motion. The globe of the eye is immediately covered by two eye-lids or palpebræ, which are compos- ed of muscular fibres covered by the common in- teguments, and lined by a very fine and smooth membrane, which is from thence extended over part of the globe of the eye, and is called tunica conjunctiva. Each eye-lid is cartilaginous at its edge; and this border which is called tarsus, is furnished with a row of hairs named cilia or eye- lashes. The * Elliot's Philosophical Observations on the Senses of Vi- sion and Hearing, 8vo. 423 PART VI. AND THEIR ORGANS. The cilia serve to protect the eye from insects and minute bodies floating in the air, and like- wise to moderate the action of the rays of light in their passage to the retina. At the roots of these hairs there are sebaceous follicles, first no- ticed by Meibomius, which discharge a glutinous liniment. Sometimes the fluid they secrete has too much viscidity, and the eye-lids become glued to each other. The upper border of the orbit is covered by the eye-brows or supercilia, which by means of their two muscles are capable of being brought towards each other, or of being carried upwards. They have been considered as serving to protect the eyes, but they are probably intended more for ornament than utility (Z). The orbits, in which the eyes are placed, are furnished with a good deal of fat, which affords a soft bed on which the eye performs its several motions. The inner angle of each orbit, or that part of it which is near the nose, is called canthus major, or the great angle; and the outer angle, which is on the opposite side of the eye, is the canthus minor, or little angle. The little reddish body which we observe in the great angle of the eye-lids, and which is call- ed caruncula lachrymalis, is supposed to be of a glandular structure, and, like the follicles of the eye- (Z) It is observable, that the eye-brows are peculiar to the human species. 424 PART VI. OF THE SENSES. eye-lids, to secrete an oily humour. But its structure and use do not seem to have been hi- therto accurately determined. The surface of the eye is constantly moistened by a very fine limpid fluid called the tears, which is chiefly, and per- haps wholly, derived from a large gland of the conglomerate kind, situated in a small depression of the os frontis near the outer angle of the eye. Its excretory ducts pierce the tunica conjunctiva just above the cartilaginous borders of the upper eye-lids. When the tears were supposed to be secreted by the caruncule, this gland was called glandula innominata; but now that its structure and uses are ascertained, it very properly has the name of glandula lachrymalis. The tears pour- ed out by the ducts of this gland are, in a natu- ral and healthy state, incessantly spread over the surface of the eye, to keep it clear and transpa- rent, by means of the eye-lids, and as constantly pass out at the opposite corner of the eye or in- ner angle, through two minute orifices, the punc- ta lachrymalia (A); being determined into these little openings by a duplication of the tunica con- jundiva, shaped like a crescent, the two points of which answer to the puncta This reduplica- tion is named membrana, or valvula semilunaris. Each (A) It sometimes happens, that this very pellucid fluid, which moistens the eye, being poured out through the ex- cretory ducts of the lachrymal gland faster than it can be carried off through the puncta, trickles down the cheek, and is then strictly and properly called tears. 425 PART VI. AND THEIR ORGANS. Each of these puncta is the beginning of a small excretory tube, through which the tears pass into a little pouch or reservoir, the sacculus lachryma- lis, which lies in an excavation formed partly by the nasal process of the os maxillare superius, and partly by the os unguis. The lower part of this sac forms a duct called the ductus ad nares, which is continued through a bony channel, and opens into the nose, through which the tears are occasionally discharged (B). The motions of the eye are performed by six muscles: four of which are straight and two oblique. The straight muscles are distinguished by the names of elevator, depressor, adductor, and abductor, from their several uses in elevating and depressing the eye, drawing it towards the nose, or carrying it from the nose towards the temple. All these four muscles arise from the bottom of the orbit, and are inserted by flat tendons into the globe of the eye. The oblique muscles are intended for the more compound motions of the eye. The first of these muscles, the obliquus superior, does not, like the other four muscles we have described, arise from the bottom of the orbit, but from the edge of the foramen that transmits the optic nerve, which separates the origin (B) When the ductus ad nares becomes obstructed in con- sequence of disease, the tears are no longer able to pass into the nostrils; the sacculus lachrymalis becomes distended; and inflammation, and sometimes ulceration, taking place, constitute the disease called fistula lachrymalis. 426 PART VI. OF THE SENSES. origin of this muscle from that of the others. From this beginning it passes in a straight line towards a very small cartilaginous ring, the situ- ation of which is marked in the skeleton by a little hollow in the internal orbitar process of the os frontis. The tendon of the muscle, after passing through this ring, is inserted into the up- per part of the globe of the eye, which it serves to draw forwards, at the same time turning the pupil downwards. The obliquus inferior arises from the edge of the orbit, under the opening of the ductus lach- rymalis; and is inserted somewhat posteriorly into the outer side of the globe, serving to draw the eye forwards and turn the pupil upwards. When either of these two muscles acts separately, the eye is moved on its axis; but when they act together, it is compressed both above and below. The eye itself, which is now to be described, with its tunics, humours, and component parts, is nearly of a spherical figure. Of its tunics, the conjunctiva has been already described as a par- tial covering, reflected from the inner surface of the eye-lids over the anterior portion of the eye. What has been named albuginea cannot pro- perly be considered as a coat of the eye, being in fact nothing more than the tendons of the straight muscles spread over some parts of the sclerotica. The immediate tunics of the eye, which are to be demonstrated when its partial coverings, and 427 PART VI. AND THEIR ORGANS. and all the other parts with which it is surround- ed, are removed, are the sclerotica, cornea, cho- roides, and retina. The sclerotica, which is the exterior coat, is every where white and opaque, and is joined at its anterior edge to another, which has more convexity than any other part of the globe, and being exceedingly transparent is called cornea (C). These two parts are perfectly different in their structure; so that some anatomists suppose them to be as distinct from each other as the glass of a watch is from the case into which it is fixed. The sclerotica is of a compact fibrous structure; the cornea, on the other hand, is composed of a great number of lamina united by cellular mem- brane. By macerating them in boiling water, they do not separate from each other, as some writers have asserted; but the cornea soon soft- ens, and becomes of a glutinous consistence. The ancients supposed the sclerotica to be a continuation of the dura mater. Morgagni and some other modern writers are of the same opi- nion; but this point is disputed by Winslow, Haller, Zin, and others. The truth seems to be, that the sclerotica, though not a production of the dura mater, adheres intimately to that membrane. The (C) Some writers, who have given the name of cornea to all this outer coat, have named what is here and most com- monly called sclerotica, cornea opaca; and its anterior and transparent portion, cornea lucida. 428 PART VI. OF THE SENSES. The choroides is so called because it is furnish- ed with a great number of vessels. It has like- wise been named uvea, on account of its resem- blance to a grape. Many modern anatomical writers have considered it as a production of the pia mater. This was likewise the opinion of the ancients; but the strength and thickness of the choroides, when compared with the delicate structure of the pia mater, are sufficient proofs of their being two distinct membranes. The choroides has of late generally been de- scribed as consisting of two laminæ; the inner- most of which has been named after Ruysch, who first described it. It is certain, however, that Ruysch's distinction is ill founded, at least with respect to the human eye, in which we are una- ble to demonstrate any such structure, although the tunica choroides of sheep and some other quadrupeds may easily be separated into two layers. The choroides adheres intimately to the scle- rotica round the edge of the cornea; and at the place of this union, we may observe a little whitish areola, named ligamentum ciliare, though it is not of a ligamentous nature. They who suppose the choroides to be com- posed of two laminæ, describe the external one as terminating in the ligamentum ciliare, and the internal one as extending farther to form the iris, which is the circle we are able to distinguish through the cornea: but this part is of a very different 429 PART VI. AND THEIR ORGANS. different structure from the choroides; so that some late writers have perhaps not improperly considered the iris as a distinct membrane. It derives its name from the variety of its colours, and is perforated in the middle. This perfora- tion, which is called the pupil or sight of the eye, is closed in the fœtus by a very thin vascular membrane. This membrana pupillaris common- ly disappears about the seventh month. On the under side of the iris we observe many minute fibres, called ciliary processes, which pass in radii or parallel lines from the circumference to the centre. The contraction and dilatation of the pupil are supposed to depend on the ac- tion of these processes. Some have considered them as muscular, but they are not of an irritable nature; others have supposed them to be fila- ments of nerves: but their real structure has never yet been clearly ascertained. Besides these ciliary processes, anatomists usu- ally speak of the circular fibres of the iris, but no such seem to exist. The posterior surface of the iris, the ciliary processes, and part of the tunica choroides, are covered by a black mucus for the purposes of ac- curate and distinct vision; but the manner in which it has been secreted has not been deter- mined. Immediately under the tunica choroides we find the third and inner coat, called the retina, which 430 PART VI. OF THE SENSES. which seems to be merely an expansion of the pulpy substance of the optic nerve, extending to the border of the crystaline humour. The greatest part of the globe of the eye, within these several tunics, is filled by a very transparent and gelatinous humour of consider- able consistence, which, from its supposed resem- blance to fused glass, is called the vitreous humour. It is invested by a very fine and delicate mem- brane, called tunica vitrea, and sometimes arach- noides.—It is supposed to be composed of two laminæ ; one of which dips into its substance, and by dividing the humour into cells adds to its firmness. The fore-part of the vitreous hu- mour is a little hollowed, to receive a very white and transparent substance of a firm texture, and of a lenticular and somewhat convex shape, named the crystaline humour. It is included in a capsula, which seems to be formed by a separa- tion of the two laminæ of the tunica vitrea. The fore-part of the eye is filled by a very thin and transparent fluid, named the aqueous humour, which occupies all the space between the crystal- line and the prominent cornea.—The part of the choroides which is called the iris, and which comes forward to form the pupil, appears to be suspended as it were in this humour, and has oc- casioned this portion of the eye to be distin- guished into two parts. One of these, which is the little space between the anterior surface of the crystalline and the iris, is called the posterior chamber; 431 PART VI. AND THEIR ORGANS. chamber; and the other, which is the space between the iris and the cornea, is called the anterior chamber of the eye (D). Both these spaces are completely filled with the aqueous humour. (E). The eye receives its arteries from the internal carotid through the foramina optica; and its veins pass through the foramina lacera, and empty themselves into the lateral sinuses. Some of the ramifications of these vessels appear on the inner surface of the iris, where they are seen to make very minute convolutions, which are sufficiently remarkable to be distinguished by the name of circulus arteriosus, though per- haps improperly, as they are chiefly branches of veins. The (D) We are aware that some anatomists, particularly Lieu- taud, are of opinion, that the iris is every where in close contact with the crystalline, and that it is of course right to speak only of one chamber of the eye; but as this does not appear to be the case, the situation of the iris and the two chambers of the eye are here described in the usual way. (E) When the crystalline becomes opaque, so as to pre- vent the passing of the rays of light to the retina, it con- stitutes what is called a contract; and the operation of couching consists in removing the diseased crystalline from its bed in the vitreous humour. In this operation the cor- nea is perforateed, and the aqueous humour escapes out of the eye, but it is constantly renewed again in a very short time. The manner, however, in which it is secreted, has not yet been determined. 432 PART VI. OF THE SENSES. The optic nerve passes in at the posterior part of the eye, in a considerable trunk, to be ex- panded for the purposes of vision, of which it is now universally supposed to be the immediate seat. But Messrs. Mariotte and Mery contend- ed, that the choroides is the seat of this sense; and the ancients supposed the crystalline to be so. Besides the optic, the eye receives branches from the third, fourth, fifth, and sixth pair of nerves. The humours of the eye, together with the cornea, are calculated to refract and converge the rays of light in such a manner as to form at the bottom of the eye a distinct image of the object we look at; and the point where these rays meet is called the focus of the eye. On the the retina, as in the camera obscura, the object is painted in an inverted position; and it is on- ly by habit that we are enabled to judge of its true situation, and likewise of its distance and magnitude. To a young gentleman who was born blind, and who was couched by Mr. Che- selden, every object (as he expressed himself) seemed to touch his eyes as what he felt did his skin; and he thought no objects so agreeable as those which were smooth and regular, although for some time he could form no judgment of their shape, or guess what it was in any of them that was pleasing to him. In order to paint objects distinctly on the re- tina, the cornea is required to have such a de- 3 gree 433 PART VI. AND THEIR ORIGIN. gree of convexity, that the rays of light may be collected at a certain point, so as to terminate exactly on the retina—If the cornea is too pro- minent, the rays, by diverging too soon, will be united before they reach the retina, as is the case with near-sighted people or myopes; and on the contrary, if it is not sufficiently convex, the rays will not be perfectly united when they reach the back part of the eye; and this hap- pens to long-sighted people or presbi, being found constantly to take place as we approach to old age, when the eye gradually flattens (F). These defects are to be supplied by means of glasses. He who has too prominent an eye, will find his vision improved by means of a concave glass; and upon the same principles, a convex glass will be found useful to a person whose eye is na- turally too flat. (F) Upon this principle, they who in their youth are near-sighted may expert to see better as they advance in life, as their eyes gradually become more flat. Ee EXPLA- 434 PART VI. OF THE SENSES. EXPLANATION OF PLATE XXX. FIGURE 1. Shows the Lachrymal Canals, after the Common Teguments and Bones have been cut away. a, The lachrymal gland, b, The two puncta lachrymalia, from which the two lychrymal ca- nals proceed to c, The lachrymal sac. d, The large lachrymal duct. e, Its opening into the nose. f, The caruncula lachrymalis. g, The eye-ball. FIG. 2. An interior View of the Coats and Hu- mours of the Eye. aaaa, The tunica sclerotica cut in four an- gles, and turned back, b b b b, The tunica cho- roides adhering to the inside of the sclerotica, and the ciliary vessels are seen passing over—cc, The retina which covers the vitreous humour. dd, The ciliary processes, which were conti- nued from the choroid coat, ee, The iris. f, The pupil. FIG. 3. Shows the Optic Nerves, and Muscles of the Eye. aa, The two optic nerves before they meet. b, The two optic nerves conjoined. c, The right 435 PART VI. AND THEIR ORGANS. right optic nerve. d, Musculus attolens palpe- bræ superioris. e, Attollens oculi. f, Abduc- tor. gg, Obliquus superior, or trochlearis. h, Adductor. i, The eye-ball. FIG. 4. Shows the Eye-balls with its Muscles, a, The optic nerve. b, Musculus trochlearis, c, Part of the os frontis, to which the trochlea or pully is fixed, through which,—d, The ten- dons of the trochlearis passes, e, Attollens oc- culi. f, Adductor oculi. g, Abductor oculi. h, Obliquus inferior, i, Part of the superior maxillary bone to which it is fixed. k, The eye-ball. FIG. 5. Represents the Nerves and Muscles of the Right Eye, after part of the Bones of the orbit have been cut away. A, The eye-ball. B, The lachrymal gland. C, Musculus abductor oculi. D, Attolens. E, Levator palpebræ superioris. F, Depressor oculi. G. Adductor. H, Obliquus superior, with its pul- ly. I, Its insertion into the sclerotic coat. K, Part of the obliquus inferior. L, The anterior part of the os frontis cut. M, The crista galli of the ethmoid bone. N, The posterior part of the sphenoid bone. O, Transverse spinous pro- cess of the sphenoid bone. P, The carotid ar- tery, denuded where it passes through the bones. Q, The 436 PART VI. OF THE SENSES. Q, The carotid artery within the cranium. R, The ocular artery. Nerves—aa, The optic nerve. b, The third pair.—c, Its joining with a branch of the first branch of the fifth pair, to form l,—The len- ticular ganglion, which sends off the ciliary nerves, d, ee, The fourth pair. f, The trunk of the fifth pair. g, The first branch of the fifth pair, named ophthalmic. h, The frontal branch of it. i, Its ciliary branches, along with which the nasal twig is sent to the nose. k, Its branch to the lachrymal gland. l, The lenticular gan- glion. m, The second branch of the fifth pair, named superior maxillary. n, The third branch of the fifth pair, named inferior maxillary, o, The sixth pair of nerves,—which sends off p, The beginning of the great sympathetic. q, The remainder of the sixth pair, spent on c, The ab- ductor oculi. FIG. 6. Represents the head of a youth, where the upper part of the cranium is sawed off,—to show the upper part of the brain, covered by the pia mater, the vessels of which are minutely filled with wax. AA, The cut edges of the upper part of the cranium. B, The two tables and intermediate diploë. BB, The two hemispheres of the ce- rebrum. CC, The incisure made by the falx. D, Part of the tentorium cerebello super ex- pansum. 437 PART VI. AND THEIR ORGANS. pansum. E, Part of the falx, which is fixed to the crista galli. FIG. 7. Represents the parts of the external Ear, with the Parotid Gland and its Ducts. a a, The helix, b, The antihelix. c, The antitragus. d, The tragus. e, The lobe of the ear. f, The cavitas innominata. g, The scapha. h, The concha. ii, The parotid gland. k, A lymphatic gland, which is often found before the tragus. l. The duct of the parotic gland. m, Its opening into the mouth. FIG. 8. A view of the posterior part of the ex- ternal ear, meatus auditorius, tympanum, with its small bones, and Eustachian tube of the right side. a, The back part of the meatus, with the small ceruminous glands. b, The incus. c, Malleus. d, The chorda tympani. e, Mem- brana tympani. f, Eustachian tube. g, Its mouth from the fauces. FIG. 9. Represents the anterior part of the right external ear, the cavity of the tympanum— its small bones, cochlea, and semicircular ca- nals. a, The malleus. b, Incus with its long leg, resting upon the stapes. c, Membrana tympani. I d e, The 438 PART VI. OF THE SENSES, &c. d, e, The Eustachian tube, covered by part of — f f, The musculus circumflexus palati. 1 2 3, The three semicircular canals. 4, The vestible. 5, The cochlea. 6, The portio mollis of the seventh pair of nerves. FIG. 10. Shews the muscles which compose the fleshy substance of the tongue. a a, The tip of the tongue, with some of the papillæ minimæ. b, The root of the tongue. c, Part of the membrane of the tongue, which covered the epiglottis. dd, Part of the mus- culus hyo-glossus. e, The lingualis. f, Genio- glossus. gg, Part of the stylo-glossus. THE END. DIRECTIONS FOR PLACING THE PLATES. Plate 19 to face page 138 20 144 21 } 158 22 } 23 } 236 24 } 25 320 26 } 326 27 } 28 378 29 404 30 438 ANATOMY Plate XXX  Just Published, Sold by THOMAS DOBSON at the Stone-House, No. 41. South Second Street. The EDINBURGH NEW DISPENSATORY: CONTAINING, I. The Elements of Pharmaceutical Chemistry. II. The Materia Medica; or, An Account of the Natural History, Qualities, Operations and Uses, of the different Substances employed in Medicine. III. 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