NASAL CHARACTERISTICS, HUMAN AND COMPARATIVE. BY Mb. CHARLES H. WARD. REPRINTED FROM THE TRANSACTIONS OF THE AMERICAN LARYNGOLOGICAL ASSOCIATION, Seventeenth Annual Meeting, 1895. NASAL CHARACTERISTICS, HUMAN AND COMPARATIVE* By Mr. CHARLES H. WARD. MR. CHAIRMAN AND GENTLEMEN : When your president called on us last Thursday to arrange for a visit from the members of the American Laryngological Association, T regretted that fate had not set the date of your meeting some months hence. Not that you are any the less welcome to-day, but that we are unprepared to entertain you in your chosen line of investigation. As an establishment we devote our energies to the making of scientific collections-collections of rocks, of fossils, of meteorites, of shells, of corals, of skins, and skeletons-in fact, of almost every- thing within the broad domains of natural history. Many are the special collections which have been gradually built up within these walls. That of comparative odontology, purchased by the New York Odontological Society, and of sponges, which pretty nearly fills the next hall, are examples. The making of a collection which shall illustrate exhaustively and well a given subject is a great educator. Probably no single person is so much benefited by a collection as the maker. Starting with but a rudimentary or even superficial knowledge of the subject treated, the museum-builder works up from the simple to the complex, from generalities to specialization. He becomes an enthusiast, a monomaniac, a devourer of mono- graphs and costly subscription books-in fact, to an unappreciative public, a crank. * Read before the American Laryngological Association at its seventeenth annual congress on the occasion of their visit to Ward's Natural Science Estab- lishment. Meeting held in the Hall of Invertebrates. 2 NASAL CHARACTERISTICS, But at about the stage when the embryo odontologist begins to speak of this skull as exhibiting a monophyodont homodont denti- tion, or of that as " a splendid case of pyorrhoea alveolaris" the col- lection is completed, perhaps sold, and our enthusiast returns to earth again, to become absorbed in turn by other and possibly remote subjects. Superficial in a sense he must perhaps ever remain, but far from ignorant. Those weeks or months of rapt study, amid a profusion of material, a library of authorities, and frequently with occasional advice and assistance from the purchaser-always an en- thusiast and a master (for, gentlemen, it is the rule that those require most illustrations who best understand their subjects)-this short, high-pressure, practical course has etched certain truths deep into his cortex, never to be entirely effaced. It has been my good fortune as a member of the working force of this institution to contribute to the series of special collections, particularly in the department of human anatomy. Later, after Professor Ward has shown you his sponge collec- tion, which competent authorities have adjudged the finest in any museum, American or foreign, I shall be pleased to exhibit a series illustrating human odontology. Some time last fall the speaker made a little suite of sectioned skulls, showing the auxiliary fossae of the nose, for Dr. George M. Lefferts, of New York, a duplicate of these here exhibited, the idea being to limit the attention of the student to one point on each specimen. The maxillary, frontal, ethmoidal, and sphenoidal sinuses are in turn revealed in four of these preparations, the fifth being an antero-posterior section of the entire skull a little to one side of the median line. (See Fig. 1.) Quite recently we agreed to make for the same gentleman a series of fifty skulls to illustrate variations of nasal septum, spurs, angles, dislocations and fractures of vomer, etc. We expect also to include a limited number of ethnological peculiarities of the anterior nares, examples of diseased turbinals, etc. As this order was re- ceived but a fortnight ago the collection is as yet hardly commenced. Hence my expressed wish that your meeting had chanced a little later. At present, we will call it the nucleus of a collection, and the labeling merely provisional. As appears from the titles of papers in your programme, not alone the larynx, but the nares, the sinuses, and adjacent territory, are embraced in your repertoire. A few re- marks on this region, from both an anthropological and a compara- tive standpoint, may not, therefore, prove amiss. HUMAN AND COMPARATIVE. 3 We love to reread even a well-known tale in an illustrated edition. Permit me, then, to review a few anatomical facts for the sake of the illustrations. We distinguish as the nose the partially bony, partially fleshy Fig. 1.-Sectioned skulls. and cartilaginous anterior prolongation of the nasal chamber. It has, in the human species, a twofold function: (1) The lodging of the first paired organs of special sense, the olfactory apparatus; and (2) as the external orifices (also paired) of the respiratory system. It also subserves, as shown by Charles Bell, by Darwin, and in detail by Dr. Harrison Allen, as a means of expression, certain flex- ures of its muscles following certain emotions of the mind. To Lavater it stood for much more : it was the outwaid and visible sign of a temperament, a character. We instinctively asso- ciate the Roman nose with command; the Grecian with a fine aesthetic sense ; the extremely thin and sharp nose with penetration, 4 NASAL CHARACTERISTICS, and the like. Besides all this it is a most marked hereditary fea- ture. Cowper's humble hero, who " Would not, with a peremptory tone, Assert the nose upon his face his own," was perhaps wise. It might have been his grandmother's. Developed, doubtless, originally as a primitive food inspector, a toxicological expert ever on the alert, the position of the nose above and in advance of the mouth is well considered. As we glance through the zoological series, the nose or its homo- logue assumes strange and bizarre forms and functions. Absent altogether in many lowly forms, to all of whom respira- tion is a necessity, the presence of an olfactory organ is either ex- tremely problematic, or suspected but unproved, in numerous cases, as, for example, the jellyfish. That this sense is one of the earliest to appear, however, can not be doubted. The olfactory sense in insects and molluscs is, when demonstrably present, frequently far removed from the respiratory system, the former breathing through a system of pulmonary tracheae that opens on the exterior by numerous spiracles, the latter, either by gills, as in the oyster, or by a single contractile spiracle, as in the shell-less land snail. Passing to the vertebrates, in the skull-less amphioxus or lancelet, specimens of which we have here, we find the respiration still carried on by means of gills, while on the left side of the anterior end of the body-the head end-an involution of the integument, a little ciliated pit, reaches inward toward a slight prominence on the spinal cord. Like the ear or the mouth, an involution of the skin marks the first appearance in vertebrates of the olfactory apparatus. Unlike the ears, however, it begins as a single organ, as in the lancelet, the hagfish and the lamprey, in which latter the olfactory lobes exceed in size the cerebral hemispheres. Classifically, these single-nostril forms are differentiated as Monorrhina from all higher vertebrates, which, having paired nos- trils, are termed Amphirrhina. In the Teleostei, or bony fishes (this perch, for example), respira- tion is still by means of gills, while the nostrils or olfactory orifices, now paired, appear just in front of either eye. The nasal sacs, how- ever, do not communicate with the mouth. In the Dipnoi or lung-fish a curious modification occurs. Liv- ing in shallow streams which dry up during the hot season, it has HUMAN AND COMPARATIVE. 5 developed accessory lungs, which enable it to breathe air during its annual imprisonment in the dried mud of the river bed-a truly amphibian life. Its nasal sacs, as in the higher vertebrates, communicate with the mouth cavity. The frog, beginning life with gills, exchanges them later for lungs. Its nares, originally used as strictly olfactory orifices, now take part in respiration, while, owing to absence of both ribs and diaphragm, the frog can inhale only by a bellows-like action of the mouth cavity, gulping his air as required. He can not breathe with his mouth open. We note, en passant, that voice here appears; the musical notes that pulsate so persistently through these long spring evenings is a batrachian serenade-a love song-and strictly masculine. Voice, then, is primarily a sexual allurement. The vague emo- tional disturbances produced in the most prosaic mind by the wailing notes of the violin or the rhythmic drum-beat seem beyond reason in their intensity, stimulating as they do brain centres that were before reason in development. It is among reptiles that we first find true nostrils-combined respiratory and olfactory orifices. These are always paired, although in the ophidians associated with but one functional lung. The nasal bones, constant in other vertebrates, are absent in most chelonians. The larynx, while more perfect than in batrachians, is still far simpler than in birds. A certain historical serpent excepted, the class of reptiles may ,be described as voiceless. The crocodiles, highest of reptiles, have valves with which to close their nostrils when wrestling with their prey in deep water. Old males among the gavials, or Gangetic crocodiles (see cut e, Fig. 3), have also a con- siderable enlargement of the nasal sacs at the tip of the snout, which enables them to remain below the surface much longer than can the females. Furthermore, the crocodile can hold a victim steadily under the surface in his open mouth, breathing betimes through the nostrils which crown the tip of his extraordinarily elongated snout -another valvular arrangement cutting off all communication, for the time being, between mouth and respiratory passages; while still other valves, or cutaneous lids, seal his ears at will. Respiration reaches its highest degree of perfection among birds : in them the blood is more rapidly and perfectly oxygenated than in any other form of animal. The bodily temperature is consequently 6 NASAL CHARACTERISTICS, higher. Not only are the lungs capacious, but a system of subsid- iary air-sacs, nine in number, traverse the body-cavity, encroaching on the abdominal viscera; while, connecting also with the lungs, the hollow bones are stored with heated air, in some instances even as far as the phalanges, making a sort of living balloon. Nor is this arrangement confined to the birds of flight: this femur of the African ostrich, which gives attachment to muscles formidable in their strength, is but a shell; how different,, for example, this smaller bone-a humerus-from an animal which lives on the sea bottom, and whose specific gravity must therefore be great. (I refer to the manatee.) John Hunter ligated the trachea of a bird, which continued to breathe through an artificial opening in the humerus. What possibilities of operative procedure does not this suggest, were only human subjects able to breathe in the same way ! The true larynx in birds, although present, is not the organ of voice. It is in the syrinx, or so-called false larynx, a flexible enlarge- ment of the lower end of the trachea, just above the branching bron- chi, that the musical notes are made. The stork, the eagle, the ostrich, lacking the syrinx, are mute, or utter only discordant cries. Tn the crane and swan a most unusual modification appears. We have here the body of a swan in which you will observe that the trachea, instead of entering the thorax directly, descends to the ster- num, which it enters, and, describing a long horizontal loop in the body of this bone, re emerges and, rising, enters the lungs. Hence the booming trombone note so distinctive of these birds. As fore- shadowed in the frog, the songs of birds are in part sexual cries, the male almost invariably excelling the female as a vocalist. b c Fig. 2. The nares, invariably two, may present as mere slits, as in the pigeon, or be produced as horny tubes, such as characterize the albatross. They may open close to the head on a very long-billed HUMAN AND COMPARATIVE. 7 bird, as this pelican, or appear at the tip of the bill, as in that rare form, the'wingless apteryx of New Zealand-degenerate scion of those gigantic forms, the Dinornis and Note that, like the crocodile, the pelican can breathe with its mouth (bill) sub- merged, but, living above the water, its nostrils are brought close to the head, instead of at the tip of the elongated snout, as in the former. The relative position of the submerged prey to the ani- mal's air supply remains the same, while the animals themselves are reversed. The scent of birds is undoubtedly keen; how far this is supple- mented or supplanted in birds of prey by their extraordinary powers of vision is mainly conjectural, yet the frightful rapidity with which buzzards congregate from all quarters at the death of an animal sug- gests some signal system of videttes as perfect as unknown. d Fig. 3. f 9 Among mammals, the development of the cerebral hemispheres is usually accompanied by a corresponding diminution in size of the olfactory lobes. The monotremes, the lowest of mammalia, com- prise two genera-the Echidna, or spiny ant-eater, a land animal, and the duck-billed platypus, or Ornithorhynchus, an aquatic form. So closely allied yet so dissimilar in appearance, the first is distin- guished by a very large, the latter by a very small, cribriform plate. In ornithorhynchus but one olfactory nerve on either side pierces the cribriform plate; in echidna the olfactory nerves are very numer- ous, and are accommodated on broad ethmo-turbinals. That the degenerate olfactory apparatus of the platypus is the 8 NASAL CHARACTERISTICS, result of his aquatic habits appears probable on a survey of the more strictly aquatic cetacea. Among marsupials the olfactory lobes remain prominent, the nasal chambers in some kangaroos, remarkable for acuteness of smell, being much wider than the anterior portion of the mouth. (See Fig. 2, a.) This development of nares reaches its extreme limit in the wombat (Fig. 2, 6), in which the nares are absolutely wider than the brain cavity. Another peculiarity of the kangaroo deserves notice. Born in an immature state, the young kangaroo is transferred to the marsupium of the female, there to be nourished by milk ejected k i Fig. 4. m h n from the teat of the mother, the young one being as yet too helpless to suckle. Nature prevents the strangling of the baby kangaroo by building upward a prolongation of the larynx, which is embraced by the soft palate, respiration continuing unimpeded the while the lac- teal fluid trickles in an unbroken stream around the craterlike larynx into the oesophagus. Of edentates, most of which have highly developed olfactory or- gans, we will glance at the great ant eater (Myrmecophaga jubatd), whose skull (Fig. 3, d) is interesting from its enormous nasal bones, measuring in this specimen 17'2 centimetres in length, and forming nearly one thirty-ninth by weight of the entire skull. Among rodents, rabbits and hares are proverbially keen of scent, here obviously specialized as a protective measure. In the porcupine the nasals are the largest bones of the skull (Fig. 4, A). Most rodents have the entire nose, save at the middle of the sep- tum, covered with hair. HUMAN AND COMPARATIVE. 9 In moles, the external nose, produced into a sort of proboscis and worked by powerful muscles, is a most efficient excavator. The star-nosed mole (Condylura cristata) develops a curious, many-rayed, double star at the extremity of his snout, a sort of third hand, whose tac- tile impressions guide him in his subterranean rambles. Certain of the bats have leaf- like prolongations of the in- tegument rising from the nose. Singular conditions are ob- served among the cetacea. In all, the vocal cords are unde- veloped, while, as in the baby kangaroo, the elongated larynx meets and is embraced by the soft palate, not here, however, as a temporary arrangement, to disappear with weaning, but a permanent provision against the encroachment of water into the trachea while feeding. The mysticeti alone of cetaceans possess olfactory nerves; we can state positively that the sperm whale knows not the odor of the ambergris which he produces -a perfume, by the way, elaborated as faecal excreta. In the sperm whale also, in common with others of the odonto- ceti-as the epiodon, hyperoodon, etc.-the skull is remarkable for a lack of bilateral symmetry, particularly in the region of the anterior nares, where the right aperture becomes very much the larger, the nostril continuing through the intervening soft parts to the exterior as a single tube, with but a single sigmoid vent. To the lookout in the crow's-nest of a whaler the spouting whale is thus instantly classified by the single or double spout long ere the animal's outlines permit of recognition. Fig. 5 is a reproduction of a photograph of the dorsal surface of the skull of hyperoddon, the tip of the pointer resting on the ante- rior nares. A fine example of Nature's wondrous adaptitude is shown Fig. 5. 10 NASAL CHARACTERISTICS, in these skulls-one (Fig. 11, o) that of the Greenland seal (Phoca groenlandica), the other (Fig. 11, p) the tropical seal (Monachus tropicalis). In the former the inferior turbinals are expanded so as to present an immense extent of radiating surface, by which the icy northern air is heated before reaching the lungs. But the tropic seal, breathing warm air, needs no such provision. Perhaps the most specialized nose among mammals is that of the elephant, in Fig. C. which the greatly elongated trunk serves not only for breathing and smell, but for prehension, for drinking-by drawing up water which is discharged into the mouth-as a trumpet, and as a club. Here also the inferior turbinals are rudimentary alike in existing tropic and in extinct northern species, the respired air being sufficiently heated in its passage through the long trunk. Crowned by a horn, or even two, in the rhinoceros, it becomes a weapon of defense, while in the sensitive nostrils of the horse are volumes of expression de- nied to his artiodactyl neighbor, the ox. Keen in the hunted ungulate, through the gradual extermination of the less gifted in this respect, the olfactory sense subserves an offensive design in the carnivora. Like the nose itself, the memory of certain scents is hereditary ; HUMAN AND COMPARATIVE. 11 newborn kittens show unmistakable aversion to the odor of a puppy as conveyed by the naked hand. The odors of animals appear to be mainly of either a sexual, an individual, or a protective character ; the former, usually more marked in the male, may be permanent, as in the dog, or confined to the rutting season, as with goats. So implicitly do dogs depend on this sense that excision of the olfactories in young puppies has resulted in a total inability to distinguish sex in after years. Individual odors are so minutely analyzed that a hound dis- criminates not only between species, kindred, sex, harm- less and harmful invaders, but distinguishes the he- reditary enemy, and, in trail- ing his leather-shod mas- ter across a populous city, must encounter myriads of tracks so similar that the mind pauses in wonder at the deli- cacy of this sense, beside which our boasted instru- ments of precision are rude indeed. Protective odors may be emitted only at the moment of provocation, as in Mephitis, or may confer a permanent immunity from attack, as in certain in- sects. As with voice, we find that scent also stimulates the reproductive functions. With the male as the active agent in emitting both sound and characteristic odor, the role of the female is confined to receiving the ensuing auditory and olfactory impressions. She there- fore should be found more receptive, more sensitive, more impres- sionable to sounds and odors in general. Does not this hold true even of our own species ? Is not the use of perfumes a womanly habit ? A significant fact in this connection is the use of musk. A constituent of practically every perfume manufactured, it adds a peculiar zest to what were otherwise mere cloying sweetness. The demi-monde alone, however, as a class, are habitually redolent of its unadulterated emanations. Now, this odor, the base of every arti- ficial olfactory delight, the chosen incense of the votaries of Venus, Fig. 7. 12 NASAL CHARACTERISTICS, is a typical sexual stimulant, an emanation elaborated primarily as a love charm by certain specialized glands of the musk-deer. Among primates-man's own class-no such radical variations of the respiratory apparatus would seem warranted as have been noted among the lower mammals. Let us examine. Geoffroy Saint- Hilaire long ago separated the monkeys of the Old World from those of the New by nasal peculiarities-the former as catarrhine, the latter as platyrrhine. In our illustration (Fig. 6) the golden howler of Brazil is contrasted with the rhesius of India. Fig. 8. Extreme variations of the vocal apparatus are found here close to man, of which the great hyoidean drum of the howler (Fig. 7), and the immense laryngeal pouches of the male gorilla, reaching down to and under the armpits, are examples. The nose itself is wide in South American monkeys, thinner in Asiatic ones, flat in the anthropoid apes, proboscislike in Nasalis, and widely varied in man. Projecting far beyond the face in most Caucasians, it may be so sunken between prominent cheek bones, as in certain Eskimos de- scribed by King, that a ruler will bridge from one cheek to the other without touching the nose. Savages and barbarians seem superior to the civilized races in keenness of scent. Humboldt's assertion that the Peruvians have three distinct words by which to designate the odor of the European, the Indian, and the negro re- spectively, reveals a delicacy of discrimination (or a pungency of odor) beyond our powers. Krafft-Ebing's account of the Swiss libertine who confessed to rousing the passions of his victims by the odor of his perspiration proves that even man has traces of a sexual odor. Compared with vertebrates generally, however, the sense of smell in man is poor, even vestigial, and is accompanied by a corre- HUMAN AND COMPARATIVE. 13 spending atrophy of the olfactory lobes. Here is a series of mod- els-casts of the brain cavities of various vertebrates-in which the proportion between olfactory lobes and cerebral hemispheres is well shown. Beginning with the ornithorhynchus, we note the sudden enlarge- ment of the olfactory region in the echidna, its absence in the por- poise, its marked prominence in the pig. Fig. 9. Compare the immense extent of this region in the extinct Dinoce- ras-that primitive ungulate whose spinal cord exceeded its brain in size-with the same area in our modern horse. Note its great de- velopment in the tiger, the expanse of the cerebral hemispheres with corresponding diminution of these lobes in the gorilla, and, lastly, their vestigial character in man. Fig. 10. A certain relation between the nasal chamber and the nasal bones is found, the latter forming its roof. As has been pointed out by an honored member of your society, Dr. Harrison Allen, a very long proboscis may, nevertheless, be associated with very short 14 NASAL CHARACTERISTICS, nasals, as in the elephant (Fig. 3, g), and very long nasals with a short proboscis, as in the pig (Fig. 3,/). Familiar to students of human anatomy as paired bones forming a portion of the border of the anterior nares, they may develop as a single bone, as in varanian lizards, or become a single bone by early ankylosis, as in the orang (Fig. 4, «), and frequently in adult life in negroes and other abo- rigines. Present in all vertebrates (with the exception of chelonia, already noted), yet few of the facial bones offer greater variations in location, in relative size, and in shape. In the gavial the nasals are far removed from the nares; the same is true in chameleons. In porpoises, according to Mivart, they occur as " small, rounded masses, each lying in a concavity on the frontal bone, and not at all rooting the nasal passages." 0 p Fig. 11. Even in a given species wide variations occur. Here are skulls of the lion (Fig. 4, n) and of the tiger (Fig. 4, m). Holden, in a note on the comparative osteology of the nasals {Human Osteology, seventh London edition, p. 106), says: "The only difference be- tween the skulls of a tiger and a lion is that in the lion the upper ends of the nasal bones and the nasal processes of the superior maxillary bones are on the same level, whereas in the tiger the nasal bones run up considerably beyond the nasal processes." This is well shown in the skulls before you. But were this the only distinction, the osteologist would often be in a quandary, as more than once in known specimens of the two skulls have we found the limits of the nasals absolutely identical. I had intended to touch briefly upon the weight of the nasal bones as found in man and in mammals generally, but time has not HUMAN AND COMPARATIVE. 15 permitted the assemblage of certain proposed data. In twelve European skulls examined, the weight of the right nasal varied from thirty centigrammes in an adult woman to one hundred and thirteen cen- tigrammes in another adult woman. In seven cases the right nasal was the heavier; in one, a female of twenty years, they were equal; in four the left were the larger. Greatest variation, thir- ty-five centigrammes(right) and twenty centigrammes (left) respectively in a skull estimated at nineteen to twenty years. Tabulating these weights in two series, one representing adults (over twenty-one years), the other series all under twenty-one years, we find, as would be expected, a preponderance of size in the older bones. Thus, of five adult skulls, the aver- age weight of the right nasal is centigrammes; of the left, 68| centi- grammes. In a very an- cient cliff-dweller skull from New Mexico the na- sals weighed respectively, right, seventy-four centi- grammes ; left, seventy centigrammes. Of the seven younger skulls (none of them under fifteen years) the average weight of the right nasal is centigrammes ; of the left, centigrammes. In other 5 4 Fig. 12. 3 2 1 16 NASAL CHARACTERISTICS, words, while the average weight of the two nasals is centi- grammes in the young series, in the adults it is 1431, an additional 1 2 Fig. 13. growth of more than sixty-eight per cent, by weight occurring after puberty. Of the adults, both the heaviest and the lightest nasals found were those of women. The proportional weight of both nasals to the entire skull as found for seven specimens was- Young Series. Adult Series. 11111 1 1 7021' 702|' 1091' 640' 328 975' 386 Truly a wide divergence. Note, however, that in both series those having largest nasals in proportion to skulls are male. These figures will, of course, be modified by future results. Mankind has been classified according to color, geographical distribution, ancestry, language, hair, shape of skull and of but- tocks, protrusion of jaw, height, political complexion, and religious faith. Professor Flower has introduced a dental index to the scientific world, which seems much more exact in its results than many of the indexes heretofore in vogue. To Dr. Paul Broca, the distin- guished French anthropologist, do we owe the use of the proportion between the maximum width and height of the skeletal nose as a classific concept. His method is as follows: The greatest width of the anterior nares, multiplied by a hundred, is divided by the dis- tance from the anterior spine to the centre of the naso-frontal suture. The resultant nasal index varies in Broca's tables from 35*71 (Rus- sian) to 72*22 (Bushman, Africa). Three natural groups are demon- HUMAN AND COMPARATIVE. 17 strated : The leptorrhini (thin-nosed), index 42 to 48 ; mesorrhini (medium-nosed), index 48 to 53 ; and the platyrrhini (broad-nosed), index 53 to 58 and above. The white race, with the Eskimos, comprise the first group; the Mongols and aboriginal Americans the second ; and the black races the third. Some of these forms are here shown. This fragment (No. 1, Fig. 12) is from a European woman, and presents the extraordinary nasal index of 30'9. Broca's lowest recorded index is the Russian, 35'71. Here is another leptorrhine skull, that of a mulatto girl, eighteen years of age (No. 3, Fig. 12). " But the negro is platyr- rhine," you protest. Granted. Yet, as a rule, the father of a mulatto is a white man and leptorrhine. Granting this and the further pos- tulate that girls usually resemble the father, and vice versa, and we find her apparently contradictory nose accounted for. In this male mulatto (No. 4, Fig. 12) from the same locality, the entire skull is markedly negroid, while the nasal index reaches 50, throwing it into the mesorrhine division. Of pla- tyrrhine skulls we have but one or two; this Micronesian, a native of New Ireland (No. 5, Fig. 12), is a good example, while this one (No. 1, Fig. 13) shows, in addition to the typical nose, a sabre cut cleanly dividing the nasals. (No. 2, Fig. 13, shows an abscess of both antrum and nares.) So far we have made only post-mortem measurements. The vivo-nasal index accomplishes the same end for the living, the great- est width between the alae now creeping in. We consequently obtain much higher indices and different limits are given to each class. Still another index, the antero-posterior, is determined thus: width : projection anteriorly :: X : 10. The anterior projection of the nose is generally in an inverse ratio to the width. Topinard enumerates the following essential points of ethnic variation of the nose: Fig. 14. Maximum height ) x , " breadth rjansverse 1Ddex- " projection ) antero Posteri°r index. 18 NASAL CHARACTERISTICS, Lobule l distinct (pinched, trilobed varieties) non-distinct. Ake ' extending beyond the nares. , near together. divergent. elliptical. Shape round, special. Nostrils - Their plane - looking downward, forward, outward, backward. Direction antero posterior. of their oblique. axis transverse. r Its angle of inclination. rectilinear. Bridge - Direction - bent or dinted, convex (aquiline), concave (retrousse), like a roof. Shape rounded off. broad and flat. Let us examine a single point, the direction of the axis of the nostril. Elliptical and nearly parallel from before backward in the 1 2 3 Fig. 15. 4 HUMAN AND COMPARATIVE. 19 purest white types, a gradual thickening of the posterior border of the nasal septum occurs as we descend the scale, and, as pointed out by Topinard, the nostrils, from being placed anteriorly, become transverse. This is well shown in this mask of the head of a Tasmanian (Fig. 14), a copy, by the way, of that in the Government Museum of Hobart Town, a death-mask of the last representative of this inter- esting race, who died in 1876. The nostrils, being in relation with the inferior border of the anterior nares, may reasonably be supposed to share with this region any peculiar ethnic features. To a considerable extent this is true. The skulls of all colored races are distinguished from those of whites by having this inferior border rounded; the nasal spine is also much reduced in size, smooth and blunt, both of which are marked simian characteristics, the nasal spine in particular being found only in man. I have here skulls of adult mulatto (1) and white (2) women, also skulls of negro (3) and of white child (4) at birth, in all of which these points are well shown (see Fig. 15). My own observations on the deflection of the nasal septum, not yet tabulated, extend over about three hundred specimens. No. No. No. No. No. No. No. No. No. zzz 6 zzz SEX SEX SEX SEX SEX SEX SEX SEX SEX RACE RACE RACE RACE RACE RACE RACE RACE RACE Section of Nasal Septum Chart, (reduced.) No. 23211 No. 73 No. 78 No. 1945 No. 1530 No. 1000 No. 2036 No. 623 No. 1895 r A L "x i SEX SEX SEX sex sex sex sex g sex $ MC£ Aleut. RACE Kanaka RACE Kanaka RACE Maori RACE Apache RACE Comanche RACE Cheyenne RACE Choctaw RACE Peruvian Same chart filled in. From skulls in Army Medical Museum. , R=Right side Nos. are those of specimens. Left side sex indicated in the usual way. / (Afars)-male, female. Fig. 16. The following diagrams were sent last Friday and Saturday to Washington and to New York. It seemed a saving of time to out- line the anterior nares ; then, in making the record, the septum is 20 NASAL CHARACTERISTICS. simply drawn in as it appears when facing the skull. Keeping in mind that for this reason the inflections are all reversed, this chart, filled in in two or three hours (through the courtesy of Mr. Frederic A. Lucas, curator of comparative anatomy of the United States National Museum), represents the condition of pretty nearly all the septa in the Army Medical Museum at Washington. A section only of the blank and of the filled forms is shown in Fig. 16. On the original chart about fifty septa were figures. Printed forms, filled in this way, express the condition of affairs fairly well, besides giving number of specimen, sex, and nationality. And now, gentlemen, let me thank you for your kind attention.